2019 in paleomalacology

This list 2019 in paleomalacology is a list of new taxa of ammonites and other fossil cephalopods, as well as fossil gastropods, bivalves and other molluscs that are scheduled to be described during the year 2019, as well as other significant discoveries and events related to molluscan paleontology that are scheduled to occur in the year 2019.

List of years in paleomalacology
  • 2009
  • 2010
  • 2011
  • 2012
  • 2013
  • 2014
  • 2015
  • 2016
  • 2017
  • 2018
  • 2019
  • 2020
  • 2021
  • 2022
  • 2023
  • 2024
  • 2025
  • 2026
  • 2027
  • 2028
  • 2029
In paleontology
2016
2017
2018
2019
2020
2021
2022

General research

Ammonites

Research

A Puzosia (Bhimaites) species juvenile shell in Burmese amber
  • Dimorphic Indonesian ammonite Macrocephalites keeuwensis is reported from the core of Jara Dome (Kutch district, India) by Jain (2019).[6]
  • Description of new fossils of members of the genus Lytohoplites from the Neuquén Basin (Argentina), taxonomic revision of the Lytohoplites species occurring in this basin, and a study on the ontogeny, probable sexual dimorphism and spectrum of intraspecific variability of members of this genus, is published by Vennari & Aguirre-Urreta (2019).[7]
  • A study on the intraspecific variation through the ontogeny of the Late Cretaceous ammonite Scaphites whitfieldi from the Western Interior of the United States is published by Klein & Landman (2019).[8]
  • A lower jaw of Spathites puercoensis is described from the Turonian Carlile Member of the Mancos Shale (New Mexico, United States) by Landman et al. (2019).[9]
  • A study on periodic variations in carbon and oxygen stable isotope profiles from Campanian and Maastrichtian ammonites belonging to the genus Baculites, and on their implications for inferring the life histories of these ammonites, is published by Ellis & Tobin (2019).[10]
  • A study on the hydrostatic properties of the orthoconic morphotype of Baculites compressus and on their implications for the knowledge of the mode of life of this animal is published by Peterman et al. (2019).[11]
  • A study on the ecology and growth of baculitid and scaphitid ammonites from the Maastrichtian Owl Creek Formation (Mississippi, United States), evaluating whether these taxa may have migrated, is published by Ferguson et al. (2019).[12]
  • A study on a sutural anomaly of ammonite specimens known as sutural pseudoinversion is published by Rogov (2019).[13]
  • A study on the systematics of Late Cretaceous ammonites from the southeastern San Juan Basin (Sandoval County, New Mexico) and on the stratigraphic framework of their fossils is published by Sealey & Lucas (2019).[14]
  • A study on the ontogenetic trajectories of septal spacing between succeeding chambers in Hypophylloceras subramosum and Phyllopachyceras ezoense is published online by Iwasaki, Iwasaki & Wani (2019).[15]
  • A study on the intra- and interspecific variation and ontogenetic trajectories of the kossmaticeratid species Maorites densicostatus and Maorites seymourianus is published by Morón-Alfonso (2019).[16]
  • A pathological ammonite specimen characterized by a pronounced left-right asymmetry in both ornamentation and suture lines is described from the Toarcian of southern France by Jattiot et al. (2019).[17]
  • A study on the morphology and function of hook‐like structures found in the body chambers of six specimens of Rhaeboceras halli is published online by Kruta et al. (2019).[18]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Albanesiceras[19]

Gen. et sp. nov

Valid

Galácz

Middle Jurassic (Bathonian)

 Italy

A member of the family Perisphinctidae. The type species is A. damianoi.

Albanites americanus[20]

Sp. nov

Valid

Brayard et al.

Early Triassic

 United States
( Idaho)

Anagaudryceras calabozoi[21]

Sp. nov

Valid

Raffi & Olivero in Raffi, Olivero & Milanese

Late Cretaceous (Campanian)

Cerro Toro Formation
Rabot Formation
Santa Marta Formation

Antarctica
(James Ross Island)
 Chile

Anagaudryceras sanctuarium[21]

Sp. nov

Valid

Raffi & Olivero in Raffi, Olivero & Milanese

Late Cretaceous (Campanian/Maastrichtian boundary)

Snow Hill Island Formation

Antarctica
(James Ross Island)

Anagaudryceras subcompressum[21]

Sp. nov

Valid

Raffi & Olivero in Raffi, Olivero & Milanese

Late Cretaceous (Maastrichtian)

Snow Hill Island Formation

Antarctica
(James Ross Island)

Araxoceltites[22]

Gen. et 3 sp. nov

Valid

Korn, Ghaderi & Tabrizi

Permian (Changhsingian)

 Iran

Genus includes new species A. sanestapanus, A. laterocostatus and A. cristatus.

Araxoceras insolens[23]

Sp. nov

Valid

Korn & Ghaderi

Permian (Wuchiapingian)

 Iran

Araxoceras truncatum[23]

Sp. nov

Valid

Korn & Ghaderi

Permian (Wuchiapingian)

 Iran

Beckeroceras[24]

Gen. et comb. nov

Valid

Naglik, De Baets & Klug

Devonian (Emsian)

Dzhaus Beds

 Uzbekistan

A member of Agoniatitida belonging to the family Mimosphinctidae. The type species is "Erbenoceras" khanakasuensis Yatskov (1990).

Beulia[25]

Gen. et sp. nov

Valid

Korn & Price

Devonian (Famennian)

 Germany

Genus includes new species B. wunderlichi.

Caribouceras[20]

Gen. et sp. nov

Valid

Brayard et al.

Early Triassic

 United States
( Idaho)

Genus includes new species C. slugense.

Carpathiceras[26]

Gen. et comb. et sp. nov

Valid

Meister & Schlögl

Early Jurassic (Sinemurian)

 Slovakia

A member of Asteroceratinae. The type species is "Coroniceras (Paracoroniceras)" bordoti Guérin-Franiatte (1966); genus also includes new species C. misiki.

Carthaginites multituberculatus[27]

Sp. nov

Valid

Kennedy in Gale, Kennedy & Walaszczyk

Late Cretaceous (Cenomanian)

Cauvery Basin

 India

A member of the family Turrilitidae.

Chieseiceras dolomiticum[28]

Sp. nov

Valid

Brack & Rieber

Middle Triassic (Late Anisian)

 Italy

Chtelnicaceras[26]

Gen. nov

Valid

Meister & Schlögl

Early Jurassic (Sinemurian)

 Slovakia

An ammonite with an intermediate morphology between Arietitidae and Oxynoticeratidae.

Docidoceras higginsi[29]

Sp. nov

Valid

Chandler

Middle Jurassic (Bajocian)

 United Kingdom

Dzhaprakoceras brodheadense[30]

Sp. nov

Valid

Work & Mason

Carboniferous (Tournaisian)

Borden Formation

 United States
( Kentucky)

Eogunnarites? elegans[27]

Sp. nov

Valid

Kennedy in Gale, Kennedy & Walaszczyk

Late Cretaceous (Cenomanian)

Cauvery Basin

 India

A member of the family Kossmaticeratidae.

Euomphaloceras varicostatum[27]

Sp. nov

Valid

Kennedy in Gale, Kennedy & Walaszczyk

Late Cretaceous (Cenomanian)

Cauvery Basin

 India

Falciclymenia cabrierensis[25]

Sp. nov

Valid

Korn & Price

Late Devonian

 France

Finiclymenia czarnockii[31]

Sp. nov

Valid

Korn & Price

Devonian (Famennian)

Central Europe

A gonioclymeniid ammonite.

Finiclymenia langei[31]

Sp. nov

Valid

Korn & Price

Devonian (Famennian)

Central Europe

A gonioclymeniid ammonite.

Follotites bighillensis[30]

Sp. nov

Valid

Work & Mason

Carboniferous (Tournaisian)

Borden Formation

 United States
( Kentucky)

Gaudryceras submurdochi[21]

Sp. nov

Valid

Raffi & Olivero in Raffi, Olivero & Milanese

Late Cretaceous (Campanian)

Rabot Formation
Santa Marta Formation

Antarctica
(James Ross Island)

Glaphyrites taimyricus[32]

Sp. nov

Valid

Konovalova & Borissenkov in Borissenkov, Konovalova & Kutygin

Carboniferous

Makarovskaya Formation

 Russia
( Krasnoyarsk Krai)

Glaucoceras[33]

Gen. et comb. nov

Valid

Matamales-Andreu et al.

Early Cretaceous (Hauterivian)

 Spain
  Switzerland

A new genus for "Baculites" renevieri Ooster (1860).

Hoevelia[25]

Gen. et sp. nov

Valid

Korn & Price

Devonian (Famennian)

 Germany

Genus includes new species H. schindewolfi.

Homoeoplanulites todaginensis[34]

Sp. nov

Valid

Poulton & Hall

Middle Jurassic (Callovian)

 Canada
( British Columbia)

Hoploscaphites macer[35]

Sp. nov

Valid

Landman et al.

Late Cretaceous (Maastrichtian)

Bearpaw Shale
Pierre Shale

 United States
( Colorado
 Montana
 Wyoming)

Ivoites meshchankinae[24]

Sp. nov

Valid

Naglik, De Baets & Klug

Devonian (Emsian)

 Czech Republic
 Uzbekistan

A member of Agoniatitida belonging to the family Mimosphinctidae.

Kamerunoceras multinodosum[27]

Sp. nov

Valid

Kennedy in Gale, Kennedy & Walaszczyk

Late Cretaceous (Turonian)

Karai Formation

 India

Kayutoceras pogrebitskyi[32]

Sp. nov

Valid

Borissenkov & Kutygin in Borissenkov, Konovalova & Kutygin

Carboniferous

Olen’ynskaya Formation

 Russia
( Krasnoyarsk Krai)

Kirsoceras arteseptatum[36]

Sp. nov

Valid

Bockwinkel, Korn & Herd

Devonian (Famennian)

 Germany

Kirsoceras gelamenum[36]

Sp. nov

Valid

Bockwinkel, Korn & Herd

Devonian (Famennian)

 Germany

Kitabobactrites[24]

Gen. et sp. nov

Valid

Naglik, De Baets & Klug

Devonian (Emsian)

 Czech Republic
 Uzbekistan

A member of Bactritida belonging to the family Bactritidae. The type species is K. salimovae.

Kumetaceras[19]

Gen. et sp. nov

Valid

Galácz

Middle Jurassic (Bathonian)

 Italy

A possible member of the family Spiroceratidae. The type species is K. enigmaticum.

Kunnamiceras[27]

Gen. et comb. nov

Valid

Kennedy in Gale, Kennedy & Walaszczyk

Late Cretaceous (Cenomanian)

Cauvery Basin

 India
 Madagascar

A member of the subfamily Acanthoceratinae. The type species is "Ammonites" tropicus Stoliczka (1865); genus also includes "Protacanthoceras" imperatoris Wright & Kennedy (1980).

Latiwitchellia atacamensis[37]

Sp. nov

Valid

Dietze & von Hillebrandt

Middle Triassic (Bajocian)

 Chile

Lissoceras maerteni[38]

Sp. nov

Valid

Pavia & Fernandez-Lopez

Middle Jurassic (Bajocian)

 France

A member of Haploceratoidea belonging to the family Lissoceratidae.

Lissoceras maizetense[38]

Sp. nov

Valid

Pavia & Fernandez-Lopez

Middle Jurassic (Bajocian)

 France

A member of Haploceratoidea belonging to the family Lissoceratidae.

Lissoceras ovale[38]

Sp. nov

Valid

Pavia & Fernandez-Lopez

Middle Jurassic (Bajocian)

El Pedregal Formation

 France
 Germany
 Hungary
 Spain

A member of Haploceratoidea belonging to the family Lissoceratidae.

Lissoceras sturanii[38]

Sp. nov

Valid

Pavia & Fernandez-Lopez

Middle Jurassic (Bajocian)

 France
 Italy
 Russia
( Dagestan)
 Turkmenistan

A member of Haploceratoidea belonging to the family Lissoceratidae.

Lissoceras submediterraneum[38]

Sp. nov

Valid

Pavia & Fernandez-Lopez

Middle Jurassic (Bajocian)

El Pedregal Formation

 France
 Spain

A member of Haploceratoidea belonging to the family Lissoceratidae.

Maeneceras tabasense[39]

Sp. nov

Valid

Korn et al.

Probably late early Famennian

 Iran

A member of the family Sporadoceratidae.

Medioclymenia media[31]

Sp. nov

Valid

Korn & Price

Devonian (Famennian)

Central Europe

A gonioclymeniid ammonite.

Megatirolites[40]

Gen. et comb. nov

Valid

Balini et al.

Early Triassic (Olenekian)

Sefid-Kuh Limestone
Tartaly Formation

 Iran
 Kazakhstan

A member of Ceratitida belonging to the superfamily Dinaritoidea and to the family Tirolitidae. The type species is "Tirolites" rossicus Kiparisova (1947); genus also includes "Tirolites" armatus Shevyrev (1968).

Mesodistichites[41]

Gen. et comb. et sp. nov

Valid

Krystyn et al.

Late Triassic (Norian)

Nayband Formation

 Iran

A member of Ceratitida belonging to the family Distichitidae. The type species is "Distichites" tozeri Seyed-Emami (1975); genus also includes new species M. evolutus.

Metabactrites rakhmonovi[24]

Sp. nov

Valid

Naglik, De Baets & Klug

Devonian (Emsian)

 Uzbekistan

A member of Agoniatitida belonging to the family Mimosphinctidae.

Nelomites[36]

Nom. nov

Valid

Bockwinkel, Korn & Herd

Devonian (Famennian)

 Kazakhstan

A replacement name for Melonites Bogoslovskii (1971).

Noridiscites nodosus[41]

Sp. nov

Valid

Krystyn et al.

Late Triassic (Norian)

Nayband Formation

 Iran

A member of Ceratitida belonging to the family Noridiscitidae.

Oraniceras calvum[42]

Sp. nov

Valid

Gulyaev

Middle Jurassic (Bathonian)

 Russia

A member of the family Parkinsoniidae.

Oraniceras issae[42]

Sp. nov

Valid

Gulyaev

Middle Jurassic (Bathonian)

 Russia

A member of the family Parkinsoniidae.

Pachydiscus wrighti[43]

Sp. nov

Valid

Kennedy

Late Cretaceous

 United Kingdom

Paroecotraustes minor[19]

Sp. nov

Valid

Galácz

Middle Jurassic (Bathonian)

 Italy

A member of the family Oppeliidae.

Phlycticeras tenue[19]

Sp. nov

Valid

Galácz

Middle Jurassic (Bathonian)

 Italy

A member of the family Strigoceratidae.

Praechebbites[44]

Gen. et sp. et comb. nov

Valid

Becker et al.

Devonian (Emsian)

 Morocco
 China?
 France?

A member of the family Mimosphinctidae belonging to the new subfamily Chebbitinae. The type species is P. debaetsi; genus might also include "Anetoceras (Erbenoceras)" mattei Feist (1970) and "Teicherticeras" ilanense Shen (1975).

Protacanthoceras parva[27]

Sp. nov

Valid

Kennedy in Gale, Kennedy & Walaszczyk

Late Cretaceous (Cenomanian)

Cauvery Basin

 India

A member of the subfamily Acanthoceratinae.

Pseudocrassiceras[45]

Gen. et comb et sp. nov

Valid

Rulleau & Jattiot

Early Jurassic (Toarcian)

 France
 Spain

A member of the family Hildoceratidae. The type species is "Ammonites" bayani Dumortier (1874); genus also includes "Ammonites" frantzi Reynès (1868), "Hildoceras" comense var. praegruneri Monestier (1931) (raised to the rank of a separate species Pseudocrassiceras praegruneri) and a new species P. arbaulti.

Pseudosonninia[46]

Gen. et sp. nov

Valid

Parent et al.

Middle Jurassic (Callovian)

Los Molles Formation

 Argentina

A member of the family Oppeliidae. The type species is P. chacaymelehuensis.

Puzosia (Bhimaites) falx[27]

Sp. nov

Valid

Kennedy in Gale, Kennedy & Walaszczyk

Late Cretaceous (Cenomanian)

Cauvery Basin

 India

Schlueterella stinnesbecki[47]

Sp. nov

Valid

Ifrim

Late Cretaceous (Turonian-Coniacian)

 Mexico

A member of the family Diplomoceratidae.

Semilissoceras[38]

Gen. et 3 sp. et comb. nov

Valid

Pavia & Fernandez-Lopez

Middle Jurassic (Bajocian)

 Algeria
 France
 Portugal
 Spain
 United Kingdom

A member of Haploceratoidea belonging to the family Lissoceratidae. The type species is S. ellipticum; genus also includes new species S. turgidulum and S. costellatum, as well as "Oppelia" depereti Flamand (1911) and "Lissoceras" semicostulatum Buckman (1923).

Sicilisphinctes[19]

Gen. et sp. nov

Valid

Galácz

Middle Jurassic (Bathonian)

 Italy

A member of the family Perisphinctidae. The type species is S. incertus.

Siemiradzkia mangoldi[48]

Sp. nov

Valid

Metodiev

Middle Jurassic (Bathonian)

Bov Formation

 Bulgaria  France

A member of the family Perisphinctidae.

Sikeliceras[19]

Gen. et sp. nov

Valid

Galácz

Middle Jurassic (Bathonian)

 Italy

A possible member of the family Spiroceratidae. The type species is S. costatum.

Sokurella elshankae[42]

Sp. nov

Valid

Gulyaev

Middle Jurassic (Bathonian)

 Russia

A member of the family Parkinsoniidae.

Sokurella pletniovkae[42]

Sp. nov

Valid

Gulyaev

Middle Jurassic (Bathonian)

 Russia

A member of the family Parkinsoniidae.

Stephanoceras (Skirroceras) englandi[49]

Sp. nov

Valid

Chandler

Middle Jurassic (Lower Bajocian)

Inferior Oolite Formation

 United Kingdom

Stephanoceras (Riccardiceras) eoeteosum[49]

Sp. nov

Valid

Chandler

Middle Jurassic (Middle or Upper Aalenian)

Inferior Oolite Formation

 United Kingdom

Uptonia schlatteri[50]

Sp. nov

Valid

Fauré & Téodori

Early Jurassic (Pliensbachian)

 France

Uzbekisphinctes[24]

Gen. et comb. nov

Valid

Naglik, De Baets & Klug

Early Devonian

Daleje Shale
Dzhaus Beds
Middle Kaub Formation

 China
 Czech Republic
 Germany
 Uzbekistan

A member of Agoniatitida belonging to the family Mimosphinctidae. The type species is "Teicherticeras" rudicostatum Bogoslovsky (1980); genus also includes "Convoluticeras" discordans Erben (1965) and "Teicherticeras" primigenitum Erben (1965).

Vedioceras fusiforme[23]

Sp. nov

Valid

Korn & Ghaderi

Permian (Wuchiapingian)

 Iran

A member of the family Araxoceratidae.

Watinoceras elegans[27]

Sp. nov

Valid

Kennedy in Gale, Kennedy & Walaszczyk

Late Cretaceous (Cenomanian)

Cauvery Basin

 India

Zelandites pujatoi[21]

Sp. nov

Valid

Raffi, Olivero & Milanese

Late Cretaceous (Campanian)

Rabot Formation
Santa Marta Formation

Antarctica
(James Ross Island)

Other cephalopods

Research

  • A study on hydrostatic properties of early Paleozoic nautiloid and endoceratoid cephalopods, and on its implications for the knowledge of ecology of these cephalopods, is published by Peterman, Barton & Yacobucci (2019).[51]
  • A study on the distribution and diversity of cephalopods during the Cambrian–Ordovician transition is published by Fang et al. (2019).[52]
  • A study on cephalopod occurrences in South China and adjacent areas during the Middle to Late Ordovician, aiming to define biogeographic provinces, is published by Fang et al. (2019).[53]
  • A study on changes of geographic range size and rates of speciation and extinction in nautiloid and ammonoid cephalopods living in the North American Midcontinent Sea from the Carboniferous (Pennsylvanian)-early Permian is published by Kolis & Lieberman (2019).[54]
  • Ascocerid fossils are described from the Hirnantian?–Llandovery strata of the Eusebio Ayala and Vargas Peña formations (southeastern Paraguay) by Cichowolski et al. (2019), representing the first ascocerid record from high paleolatitudes of Gondwana.[55]
  • A juvenile specimen of Dolorthoceras, representing the first Devonian cephalopod specimen preserving nacreous structures and the first ectocochleate cephalopod preserving fibrous structures within its conch, is described from the Frasnian Domanic Formation (Russia) by Doguzhaeva (2019).[56]
  • A study on the response of cephalopod assemblages from the area of the present-day European Russia to environmental changes during the early Aptian Oceanic Anoxic Event is published by Rogov et al. (2019).[57]
  • A study on the paleobiology of cephalopods from the Albian Mahajanga Basin of Madagascar will be published by Hoffmann et al. (2019).[58]
  • A proposal for revision of the classification of nautiloid cephalopods presented in Treatise Part K is presented by King & Evans (2019), who name a new subclass Tarphyceratia, new order Rioceratida and new family Bactroceratidae.[59]
  • A study on embryonic and early juvenile development of shell in Peismoceras is published by Manda & Turek (2019).[60]
  • A study on changes of the morphological diversity of Late Permian coiled nautiloids is published online by Korn et al. (2019).[61]
  • A study on the paleobiology of Angulithes mermeti, based on new material from the upper Cenomanian of Egypt, and on its implications for the knowledge of the evolution of Cenomanian lineage of Angulithes, is published by Wilmsen & Nagm (2019).[62]
  • A study on the anatomy of Gordoniconus beargulchensis and on its implications for the knowledge of evolution of coleoids is published by Klug et al. (2019).[63]
  • A study on a putative Sinemurian octocoral Mesosceptron from Montagna del Casale (Sicily, Italy) is published by Pignatti et al. (2019), who reinterpret the fossils of this taxon as incompletely preserved coleoid rostra, and consider Mesosceptron to be a subjective junior synonym of the genus Atractites.[64]
  • A study on four specimens of Clarkeiteuthis conocauda from the Toarcian Posidonia Shale (Germany) preserved with small bony fish in their arm crowns, on the feeding behaviour of members of this species, and on the differences in the mode of life of C. conocauda, Passaloteuthis bisulcata and Hibolithes semisulcatus, is published by Jenny et al. (2019).[65]
  • A study on belemnite body-size dynamics across the PliensbachianToarcian boundary in the Peniche section (Lusitanian Basin, Portugal), evaluating its implications for the knowledge of the impact of the Pliensbachian–Toarcian boundary warming event on belemnite body size, is published by Rita et al. (2019).[66]
  • New belemnite fossils are described from the BerriasianHauterivian of Arctic Canada and North-East Greenland by Mutterlose et al. (2019), who also evaluate the implications of these fossils for the knowledge of spatial distribution patterns of belemnites at northern high latitudes.[67]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Andamanoceras[68]

Gen. et sp. nov

Valid

Niko, Sone & Leman

Ordovician

Lower Setul Limestone

 Malaysia

A member of Pseudorthocerida. Genus includes new species A. densiseptum.

Basiloceras[69]

Gen. et sp. nov

Valid

Pohle, Klug & Haas

Devonian (Eifelian and Givetian)

 Morocco

A member of Oncocerida belonging to the family Acleistoceratidae. Genus includes new species B. goliath and B. david.

Beloitoceras thorslundi[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Oncoceratidae.

Cabaneroceras[71]

Gen. et sp. nov

Valid

Kröger & Gutiérrez-Marco

Ordovician (Darriwilian)

 Spain

A member of Intejocerida. Genus includes new species C. aznari.

Cameroceras motsognir[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

Cimomia kurkurensis[72]

Sp. nov

Valid

Hewaidy, Qot & Moneer

Early Paleocene

Kurkur Formation

 Egypt

A member of the family Hercoglossidae.

Clothoceras[70]

Gen. et sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Proteoceratidae. Genus includes new species C. thornquisti.

Danoceras skalbergensis[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

Deltoidonautilus hassani[72]

Sp. nov

Valid

Hewaidy, Qot & Moneer

Late Paleocene

Tarawan Formation

 Egypt

A member of the family Hercoglossidae.

Dicoelites aprilis[73]

Sp. nov

Valid

Dzyuba in Dzyuba, Mitta & Sherstyukov

Middle Jurassic (Bajocian)

Djangura Formation

 Russia
( Karachay-Cherkessia)

A belemnite.

Dicoelites octobris[73]

Sp. nov

Valid

Dzyuba in Dzyuba, Mitta & Sherstyukov

Middle Jurassic (Bajocian)

Djangura Formation

 Russia
( Karachay-Cherkessia)

A belemnite.

Discoceras amtjaernense[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

Discoceras nilssoni[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

Endoceras naekki[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

Engeseriteuthis[74]

Gen. et sp. nov

Valid

Fuchs et al.

Late Jurassic (late Kimmeridgian to ?early Tithonian)

Geisental Formation

 Germany

A member of Muensterelloidea. Genus includes new species E. arcuatus.

Eromangateuthis[75]

Gen. et comb. nov

Valid

Fuchs

Early Cretaceous (Albian)

Allaru Formation

 Australia

A member of Octobrachia belonging to the family Plesioteuthidae; a new genus for "Boreopeltis" soniae Wade (1993).

Etchesia[76]

Gen. et sp. nov

Valid

Fuchs

Late Jurassic (Tithonian)

Kimmeridge Clay Formation

 United Kingdom

A member of Coleoidea belonging to the family Muensterellidae. Genus includes new species E. martilli.

Furudaloceras[70]

Gen. et sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Cyrtogomphoceratidae. Genus includes new species F. tomtei.

Gujiaonautilus[77]

Gen. et sp. nov

Valid

Miao et al.

Permian (Changhsingian)

Dalong Formation

 China

A nautiloid. Genus includes new species G. longliensis.

Isbergoceras[70]

Gen. et 2 sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Graciloceratidae. The type species is I. niger; genus also includes I. consobrinum.

Isorthoceras nikwis[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Proteoceratidae.

Isorthoceras sylphide[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Proteoceratidae.

Isorthoceras urdr[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Proteoceratidae.

Kullsbergoceras[70]

Gen. et sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Cyrtogomphoceratidae. Genus includes new species K. nissei.

Lambeoceras (Lambeoceras) balashovi[78]

Sp. nov

Valid

Barskov & Marek

Ordovician (Katian)

 Russia

Lambeoceras (Lambeoceras) hanae[78]

Sp. nov

Valid

Barskov & Marek

Ordovician (Katian)

 Russia

Lambeoceras (Lambeoceras) sibiricum[78]

Sp. nov

Valid

Barskov & Marek

Ordovician (Katian)

 Russia

Langgunites[68]

Gen. et sp. nov

Valid

Niko, Sone & Leman

Ordovician

Lower Setul Limestone

 Malaysia

A member of Pseudorthocerida. Genus includes new species L. mucronulatus.

Malayorthoceras[68]

Gen. et sp. nov

Valid

Niko, Sone & Leman

Ordovician

Lower Setul Limestone

 Malaysia

A member of Orthocerida. Genus includes new species M. gracilentum.

Muensterella jillae[74]

Sp. nov

Valid

Fuchs et al.

Late Cretaceous (Turonian)

South Bosque Formation

 United States
( Texas)

Muensterella spinosa[74]

Sp. nov

Valid

Fuchs et al.

Late Jurassic (Tithonian)

Mörnsheim Formation

 Germany

Muensterellina[74]

Gen. et sp. nov

Valid

Fuchs et al.

Middle Jurassic (Callovian)

Oxford Clay Formation

 United Kingdom

A member of Muensterelloidea. Genus includes new species M. johnjagti.

Neotainoceras zhaoi[77]

Sp. nov

Valid

Miao et al.

Permian (Changhsingian)

Dalong Formation

 China

A nautiloid.

Nodopleuroceras gujiaoense[77]

Sp. nov

Valid

Miao et al.

Permian (Changhsingian)

Dalong Formation

 China

A nautiloid.

Ordogeisonoceras uppsalaensis[70]

Sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Geisonoceratidae.

Sepia vandervoorti[79]

Sp. nov

Valid

Košťák, Jagt & Schlögl

Miocene (late Langhian–earliest Serravallian)

Köselerli Formation

 Turkey

A species of Sepia.

Shanthaiceras[68]

Gen. et sp. nov

Valid

Niko, Sone & Leman

Ordovician

Lower Setul Limestone

 Malaysia

A member of Pseudorthocerida. Genus includes new species S. amplum.

Sibumasuoceras kilimense[68]

Sp. nov

Valid

Niko, Sone & Leman

Ordovician

Lower Setul Limestone

 Malaysia

A member of Pseudorthocerida.

Sibumasuoceras scrivenori[68]

Sp. nov

Valid

Niko, Sone & Leman

Ordovician

Lower Setul Limestone

 Malaysia

A member of Pseudorthocerida.

Tofangoceras kedahense[68]

Sp. nov

Valid

Niko, Sone & Leman

Ordovician

Lower Setul Limestone

 Malaysia

A member of Orthocerida.

Tofangoceras rayense[68]

Sp. nov

Valid

Niko, Sone & Leman

Ordovician

Lower Setul Limestone

 Malaysia

A member of Orthocerida.

Tofangoceras staiti[68]

Sp. nov

Valid

Niko, Sone & Leman

Ordovician

Lower Setul Limestone

 Malaysia

A member of Orthocerida.

Tyrionella[74]

Gen. et sp. nov

Valid

Fuchs et al.

Late Jurassic (Tithonian)

Altmühltal Formation

 Germany

A member of Muensterelloidea. Genus includes new species T. fauseri.

Valkyrioceras[70]

Gen. et sp. nov

Valid

Kröger & Aubrechtová

Late Ordovician

Kullsberg Limestone Formation

 Sweden

A member of the family Valcouroceratidae. Genus includes new species V. dalecarlia.

Gastropods

Research

  • A study on the diversity of gastropod larval conch fossil assemblages of ages ranging from the Ordovician to Carboniferous is published online by Dzik (2019).[80]
  • A study on the recovery of gastropods in the aftermath of the Cretaceous–Paleogene extinction event, based on data from fossils from the former Ankerpoort‐Curfs quarry (Geulhem, the Netherlands), is published online by Vellekoop et al. (2019).[81]
  • A probable cyclophoroidean gastropod, possibly a member of the family Cyclophoridae, is described from the Cretaceous amber from Myanmar by Xing et al. (2019), constituting the first confirmed and oldest record of soft-bodied preservation of a snail in Cretaceous amber.[82]
  • A study on the nature of the size dimorphism in the Miocene gastropod Persististrombus deperditus, and on the causes of its evolution, is published by Halder & Paira (2019).[83]
  • New specimens of Spinucella reimersi are described from the Miocene Gram Clay of Denmark by Schnetler (2019), who also provides an emended description of this species.[84]
  • A revision of extant and Neogene members of the genus Antistreptus from the southwestern Atlantic Ocean is published by Pastorino & Griffin (2019).[85]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Acirsa torensis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Acirsa.

Acmaea selandica[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Acmaea.

Actinella fecundaerrata[87]

Nom. nov

Valid

Teixeira et al.

Quaternary

 Madeira

A species of Actinella.

Agaronia almaghribensis[88]

Sp. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

A species of Agaronia.

Ageria laxa[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Ageria skeldervigensis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Allmonia[89]

Gen. et comb. nov

Valid

Harzhauser & Landau

Oligocene (Rupelian) to Miocene (Tortonian)

 Algeria
 Austria
 Belgium
 Bosnia and Herzegovina
 Bulgaria
 Czech Republic
 France
 Germany
 Greece
 Hungary
 India
 Iran
 Italy
 Netherlands
 Pakistan
 Portugal
 Slovakia
 Slovenia
  Switzerland
 Turkey

A member of the family Turritellidae. The type species is "Turritella" cathedralis Brongniart (1823); genus also includes several European Cenozoic species previously placed in Protoma.

Alvania amphitrite[90]

Sp. nov

Valid

Thivaiou, Harzhauser & Koskeridou

Miocene (Aquitanian)

Pentalofos Formation

 Greece

A species of Alvania.

Amaea alicae[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Amaea stevniensis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Amaurellina ravni[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Amaurotoma? multispirata[91]

Sp. nov

Valid

Ketwetsuriya, Cook & Nützel

Permian (Wordian)

Ratburi Group

 Thailand

A member of the family Trochonematidae.

Ananias belaensis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Eotomariidae.

Ananias bashkiriensis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Eotomariidae.

Anematina indiscreta[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Elasmonematidae.

Aneudaronia[93]

Gen. et sp. nov

Valid

Guzhov

Jurassic

 Russia

A member of the family Stuoraxidae. Genus includes new species A. elegans.

Anomphalus glaucus[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

Apachella rugosa[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Eotomariidae.

Aphera chattica[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Aphera.

Aquitanobursa[95]

Gen. et comb. nov

Valid

Sanders, Merle & Puillandre

Oligocene (Rupelian) to early Pliocene

 Dominican Republic
 France
 Italy
 Pakistan
 Venezuela
 United States?

A member of the family Bursidae. The type species is "Ranella" grateloupi d’Orbigny (1852); genus also includes "Bursa" amphitrites Maury (1917), "Apollon" inaequicrenatus Cossmann & Peyrot (1924), "Ranella" morrisi d’Archiac & Haime (1853), "Ranella" tuberosa Grateloup (1833) and possibly also "Bursa (Bufonariella)" chipolana Schmelz (1997).

Archaeocyclotus[96]

Gen. et sp. nov

Valid

Asato & Hirano in Hirano et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Cyclophoridae. The type species is A. plicatula.

Archaeocyclotus plicatula

Archimediella carpathica[89]

Sp. nov

Valid

Harzhauser & Landau

Middle Miocene

 Austria
 Bosnia and Herzegovina
 Hungary
 Poland
 Romania
 Slovakia

A member of the family Turritellidae.

Arene jakobseni[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Arene.

Argentisioliella[97]

Gen. et sp. nov

Valid

Miquel

Early Pliocene

Irene Formation

 Argentina

A member of the family Cochliopidae. Genus includes new species A. pardignasi.

Arribazona seleukensis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Orthonematidae.

Ascheria salina[98]

Sp. nov

Valid

Kiel et al.

Early Oligocene

Talara Basin

 Peru

A member of the family Hokkaidoconchidae.

Ataphrus knoesensis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Athleta benoisti[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Athleta.

Athleta confusus[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Athleta.

Athleta maculosus[94]

Sp. nov

Valid

Lozouet

Miocene

 France

A species of Athleta.

Athleta memoirae[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Athleta.

Athleta peyreirensis[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Athleta.

Attiliosa gallica[99]

Sp. nov

Valid

Landau et al.

Miocene (Tortonian)

 France

A species of Attiliosa.

Attiliosa pouweri[99]

Sp. nov

Valid

Landau et al.

Miocene (Tortonian)

 France

A species of Attiliosa.

Auritoma[100]

Gen. et sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Genus includes new species A. lenticula.

Bandelzyga[92]

Gen. et sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudozygopleuridae. Genus includes new species B. elegans.

Bathrotomaria jakobseni[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Bayania falconneti[101]

Sp. nov

Valid

Pacaud

Paleogene

 France

Baylea longispira[92]

Sp. nov

Valid

Mazaev

Permian

 China
 Russia

A member of the family Lophospiridae.

Bellazona acuta[102]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

Bellerophon shakhtauensis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

Birgella burchi[103]

Sp. nov

Valid

Naranjo-García & Aguillón in Vega et al.

Late Cretaceous (late Campanian)

Cerro del Pueblo Formation

 Mexico

A member of the family Hydrobiidae.

Bleytonella saalensis[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of the family Pickworthiidae.

Bobellis[105]

Gen. et sp. nov

Valid

Peel

Early Silurian

Samuelsen Høj Formation

 Greenland

A member of Euomphaloidea belonging to the family Pycnomphalidae. The type species is B. oliveri.

Bocourtia (Kuschelenia?) bonariensis[97]

Sp. nov

Valid

Miquel

Early Pliocene

Irene Formation

 Argentina

A member of the family Bulimulidae.

Bolicharewia[92]

Gen. et sp. et comb. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Anomphalidae. Genus includes new species B. edita, as well as "Anomphalus" straparoliformis Mazaev (1997).

Bonellitia cryptica[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Bonellitia.

Bonellitia peyroti[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Bonellitia.

Boreomica costaspiralis[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Rissooidea belonging to the family Palaeorissoinidae.

Busambrella[100]

Gen. et sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Genus includes new species B. fasciata.

Buvignieria convexa[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Rissooidea belonging to the family Palaeorissoinidae.

Buvignieria racitana[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Rissooidea belonging to the family Palaeorissoinidae.

Callispira taosiformis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Sinuspiridae.

Calyptraea baldii[106]

Sp. nov

Valid

Vicián, Kovács & Stein

Late Oligocene – early Miocene

 Hungary

A species of Calyptraea.

Ceratia rodvigensis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Ceratia.

Cerion uva gouldi[107]

Subsp. nov

Valid

Harasewych

Pleistocene

 Aruba

A subspecies of Cerion uva.

Cerithiella selandica[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Cerithiella.

Cerithiopsis restemlongis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Cerithiopsis.

Chartronella belaensis[108]

Sp. nov

Valid

Szabó & Jaitly

Middle Jurassic (Callovian)

Chari Formation

 India

A member of the family Paraturbinidae.

Chelotia pervicina[109]

Sp. nov

Valid

Berezovsky & Pacaud

Middle Eocene

 Ukraine

A member of the family Pleurotomariidae.

Chlorozyga asiatica[91]

Sp. nov

Valid

Ketwetsuriya, Cook & Nützel

Permian (Wordian)

Ratburi Group

 Thailand

A member of the family Imoglobidae.

Chriskingia[110]

Gen. et comb. nov

Valid

Janssen & Wilson

Miocene

 Dominican Republic
 Trinidad and Tobago

A member of the family Atlantidae; a new genus for "Atlanta" rotundata Gabb (1873).

Clavatula danuvii[106]

Sp. nov

Valid

Vicián, Kovács & Stein

Late Oligocene – early Miocene

 Hungary

A species of Clavatula.

Clavilithes tamdakhtensis[88]

Nom. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

A species of Clavilithes.

Claviscala minor[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Cleopatra adami[111]

Nom. nov

Valid

Neiber & Glaubrecht

Early Pleistocene

Kaiso Series

 Uganda

A species of Cleopatra; a replacement name for Viviparus cylindricus Adam (1957).

Clithon (Pictoneritina) coeuvrensis[112]

Sp. nov

Valid

Vrinat

Eocene (Ypresian)

 France

A species of Clithon.

Clithon (Pictoneritina) occultatus[112]

Sp. nov

Valid

Vrinat

Eocene (Ypresian)

 France

A species of Clithon.

Clithon (Pictoneritina) tigrinus[112]

Sp. nov

Valid

Vrinat

Eocene (Ypresian)

 France

A species of Clithon.

Cochlis odovychenorum[113]

Sp. nov

Valid

Pedriali, Sosso & Dell'Angelo

Miocene

 Ukraine

A member of the family Naticidae.

Cochlis ukrainensis[113]

Sp. nov

Valid

Pedriali, Sosso & Dell'Angelo

Miocene

 Ukraine

A member of the family Naticidae.

Coninoda strekwera[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of the family Eustomatidae.

Coniscala umbilica[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Contortia grateloupi[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Cancellariidae.

Contortia intermedia[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Cancellariidae.

Contortia zitae[106]

Sp. nov

Valid

Vicián, Kovács & Stein

Late Oligocene – early Miocene

 Hungary

A member of the family Cancellariidae.

Coptostylus harzhauseri[101]

Nom. nov

Valid

Pacaud

Paleogene

 France

Cosmocerithium veliger[114]

Sp. nov

Valid

Guzhov

Late Jurassic

 Russia

A member of the family Cerithiopsidae.

Cossmannina alfischeri[115]

Sp. nov

Valid

Foster et al.

Early Triassic

 China

Cranopsis cretacea[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Cranopsis.

Cristalloella (Wonwalica) permica[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Tofanellidae.

Cryptaulax? parvum[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Cerithioidea belonging to the family Cryptaulacidae.

Cryptaulax? triangulare[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Cerithioidea belonging to the family Cryptaulacidae.

Cryptoptyxis rarenodosa[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of the family Maoraxidae.

Cylichna stevniensis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Cylichna.

Cyllene satoi[116]

Sp. nov

Valid

Amano

Late Pliocene

Tentokuji Formation

 Japan

A species of Cyllene.

Cypraeorbis nadaii[106]

Sp. nov

Valid

Vicián, Kovács & Stein

Late Oligocene – early Miocene

 Hungary

A cowry.

Cystiscus pseudobrevis[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Cystiscus.

Dalliella elisae[101]

Sp. nov

Valid

Pacaud

Paleogene

 France

Denticulofusus[117]

Gen. et comb. nov

Valid

Vermeij & Pacaud

Eocene (Lutetian and Priabonian)

 France

A member of the family Columbariidae. The type species is "Fusus" gothicus Deshayes (1834); genus also includes "Coluzea" cavelieri Gain & Le Renard (2017) and "Coluzea" monicae Belliard, Gain & Le Renard (2017).

Dentimargo salvani[88]

Sp. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

A species of Dentimargo.

Dictyotomaria balabanovi[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Phymatopleuridae.

Discula cameroni[87]

Sp. nov

Valid

Teixeira et al.

Quaternary

 Madeira

A species of Discula.

Discula polymorpha docaensis[87]

Subsp. nov

Valid

Teixeira et al.

Quaternary

 Madeira

Doggerostra riedeli affinis[93]

Subsp. nov

Valid

Guzhov

Late Jurassic

 Russia

A member of Heterobranchia of uncertain phylogenetic placement.

Dolicholatirus lorenzi[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Dolicholatirus.

Donaldina erwini[115]

Sp. nov

Valid

Foster et al.

Early Triassic

 China

Dragonia[114]

Gen. et 2 sp. nov

Valid

Guzhov

Late Jurassic

 Russia

A member of the family Cerithiopsidae. Genus includes new species D. minuta and D. longa.

Eirlysia reticulatiformis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Phymatopleuridae.

Elymicyclus[100]

Gen. et 4 sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Genus includes new species E. alternatus, E. ietumensis, E. martae and E. garibaldii.

Emarginula acutidens[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

A species of Emarginula.

Emarginula burgioi[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

A species of Emarginula.

Eoplatyzona ratchaburiensis[91]

Sp. nov

Valid

Ketwetsuriya, Cook & Nützel

Permian (Wordian)

Ratburi Group

 Thailand

A member of the family Gosseletinidae.

Eovoluta adegokei[88]

Nom. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

Exelissa indiana[108]

Sp. nov

Valid

Szabó & Jaitly

Middle Jurassic (Callovian)

Gadhada Formation

 India

A member of the family Cryptaulacidae.

Epalxis? rosenkrantzi[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Transferred to the genus Fusulculus by Pacaud (2020), who coined a replacement name Fusulculus hanseni.[118]

Epetrium skeldervigensis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Epitonium (Epitonium) zhuoi[119]

Sp. nov

Valid

Yu, Wang & Jarzembowski

Late Cretaceous (Cenomanian

Burmese amber

 Myanmar

A gastropod of uncertain phylogenetic placement. Originally described as a species of Epitonium; Neubauer et al. (2019) transferred this species to the pupinid genus Pseudopomatias.[120]

Epitonium (Papyriscala) lyui[119]

Sp. nov

Valid

Yu, Wang & Jarzembowski

Late Cretaceous (Cenomanian

Burmese amber

 Myanmar

A gastropod of uncertain phylogenetic placement. Originally described as a species of Epitonium; Neubauer et al. (2019) transferred this species to the pupinid genus Pseudopomatias.[120]

Euconospira skuini[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Phymatopleuridae.

Eucyclomphalus? marenostrum[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Eucycloscala gracilicostatus[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Euspira jhuraensis[121]

Sp. nov

Valid

Das et al.

Late Jurassic

 India

A species of Euspira.

Euspira lakhaparensis[121]

Sp. nov

Valid

Das et al.

Late Jurassic

 India

A species of Euspira.

Euspira sirenkoi[113]

Sp. nov

Valid

Pedriali, Sosso & Dell'Angelo

Miocene

 Ukraine

A species of Euspira.

Euthema hesoana[96]

Sp. nov

Valid

Asato & Hirano in Hirano et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Diplommatinidae.

Euthema hesoana

Falsobuvigna[104]

Gen. et sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Rissooidea belonging to the new family Falsobuvignidae. The type species is F. peregrina.

Fischeriella sicula[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Fissurella? stantoni[122]

Sp. nov

Valid

Powell & Geiger

Miocene

Topanga Canyon Formation

 United States
( California)

A member of the family Fissurellidae.

Fusimorio salbriacensis[94]

Sp. nov

Valid

Lozouet

Miocene

 France

A member of the family Cancellariidae.

Fusinus schnetleri[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Fusinus.

Gibberula amurakucensis[88]

Sp. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

A species of Gibberula.

Gibberula bezoyensis[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Gibberula.

Gibberula pseudoaquitanica[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Gibberula.

Gibberula subovulatoides[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Gibberula.

Girtyspira oblonga[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Meekospiridae.

Globodoma magna[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Gosseletinidae.

Goniasma gerasimovi[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Goniasmatidae.

Goniobasis unilirata[103]

Sp. nov

Valid

Naranjo-García & Aguillón in Vega et al.

Late Cretaceous (late Campanian)

Cerro del Pueblo Formation

 Mexico

A member of the family Pleuroceridae.

Gymnocerithium? convexoconcavum[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Campaniloidea belonging to the family Gymnocerithiidae.

Gyrodes mahalanobisi[121]

Sp. nov

Valid

Das et al.

Late Jurassic

 India

A member of the family Naticidae.

Haitia taylori[103]

Sp. nov

Valid

Naranjo-García & Aguillón in Vega et al.

Late Cretaceous (late Campanian)

Cerro del Pueblo Formation

 Mexico

A member of the family Physidae.

Harmatia tokodensis[123]

Sp. nov

Valid

Kovács & Vicián

Eocene

 Hungary

A member of the family Muricidae.

Hemicerithium perrettai[101]

Sp. nov

Valid

Pacaud

Eocene

 United Kingdom

Hesperiella cyrtocostata[91]

Sp. nov

Valid

Ketwetsuriya, Cook & Nützel

Permian (Wordian)

Ratburi Group

 Thailand

A member of the family Porcelliidae.

Heteronatica[93]

Gen. et sp. nov

Valid

Guzhov

Jurassic

 Russia

A member of the family Cornirostridae. Genus includes new species H. globosa.

Hinea gaellae[101]

Sp. nov

Valid

Pacaud

Paleogene

 France

A species of Hinea.

Hirsuticyclus[124]

Gen. et sp. nov

Valid

Neubauer, Xing & Jochum

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Cyclophoridae. The type species is H. electrum.

Holospira thompsoni[103]

Sp. nov

Valid

Naranjo-García & Aguillón in Vega et al.

Late Cretaceous (late Campanian)

Cerro del Pueblo Formation

 Mexico

A species of Holospira.

Homalopoma acaste[90]

Sp. nov

Valid

Thivaiou, Harzhauser & Koskeridou

Miocene (Aquitanian)

Pentalofos Formation

 Greece

A species of Homalopoma.

Hydrobia galatoniana[125]

Sp. nov

Valid

Esu & Girotti

Oligocene (Chattian)

Galatone Formation

 Italy

A species of Hydrobia sensu lato.

Hydrobia ionica[125]

Sp. nov

Valid

Esu & Girotti

Oligocene (Chattian)

Galatone Formation

 Italy

A species of Hydrobia sensu lato.

Hyphantozyga? khaophrikensis[91]

Sp. nov

Valid

Ketwetsuriya, Cook & Nützel

Permian (Wordian)

Ratburi Group

 Thailand

A member of the family Pseudozygopleuridae.

Involuta coxi[126]

Sp. nov

Valid

Halder & Das

Eocene

 India

A volute.

Jponsia aryanamensis[101]

Sp. nov

Valid

Pacaud

 Iran

Jponsia yesimae[101]

Sp. nov

Valid

Pacaud

Paleogene

 Turkey

Jumaramaria[108]

Gen. et sp. nov

Valid

Szabó & Jaitly

Middle Jurassic (Bathonian)

Jhurio Formation

 India

A possible member of the family Pleurotomariidae. The type species is J. jumaraensis.

Jurassiscala[100]

Gen. et 2 sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Genus includes new species J. sturanii and possibly also J? tenuiretis.

Keilostoma nautagarona[101]

Sp. nov

Valid

Pacaud

Paleogene

 France

Laevitomaria babalusciae[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Lagocheilus cretaspira[96]

Sp. nov

Valid

Asato & Hirano in Hirano et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Lagocheilus.

Lagocheilus cretaspira

Lagocheilus electrospira[96]

Sp. nov

Valid

Asato & Hirano in Hirano et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Lagocheilus.

Lagocheilus electrospira

Leiostyla castanheiraensis[87]

Sp. nov

Valid

Teixeira et al.

Quaternary

 Madeira

A species of Leiostyla.

Leiostyla cooki[87]

Sp. nov

Valid

Teixeira et al.

Quaternary

 Madeira

A species of Leiostyla.

Leiostyla desertaensis[87]

Sp. nov

Valid

Teixeira et al.

Quaternary

 Madeira

A species of Leiostyla.

Leiostyla simulans[87]

Sp. nov

Valid

Teixeira et al.

Quaternary

 Madeira

A species of Leiostyla.

Leptaxis groviana antiqua[87]

Subsp. nov

Valid

Teixeira et al.

Quaternary

 Madeira

A species of Leptaxis.

Leptaxis isambertoi[87]

Sp. nov

Valid

Teixeira et al.

Quaternary

 Madeira

A species of Leptaxis.

Levizygopleura composita[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudozygopleuridae.

Levizygopleura magna[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudozygopleuridae.

Littoraria (Littorinopsis) massicardi[101]

Sp. nov

Valid

Pacaud

Paleogene

 France

A species of Littoraria.

Lyria peyrehoradensis[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Lyria.

Masaevia[93]

Gen. et sp. nov

Valid

Guzhov

Jurassic

 Russia

A member of Heterobranchia of uncertain phylogenetic placement. Genus includes new species M. sinistra.

Mathilda dania[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Mathilda.

Melanopsis neubaueri[101]

Sp. nov

Valid

Pacaud

Paleogene

 France

A species of Melanopsis.

Melarhaphe perminima[101]

Sp. nov

Valid

Pacaud

Paleogene

 France

A species of Melarhaphe.

Melarhaphe vanhyftei[101]

Sp. nov

Valid

Pacaud

Paleogene

 France

A species of Melarhaphe.

Merica admirabilis[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Merica.

Merica estotiensis[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Merica.

Merica mutabilis[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Merica.

Mesalia bohnhavasae[89]

Sp. nov

Valid

Harzhauser & Landau

Middle Miocene

 Hungary

A species of Mesalia.

Mesalia danica[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Mesalia.

Mesalia sovisi[89]

Sp. nov

Valid

Harzhauser & Landau

Middle Miocene

 Austria
 Czech Republic

A species of Mesalia.

Mesalia stryriaca[89]

Sp. nov

Valid

Harzhauser & Landau

Middle Miocene

 Austria

A species of Mesalia.

Monimirus[92]

Gen. et sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Stuoraxidae. Genus includes new species M. bonus.

Monodonta joetsuensis[127]

Sp. nov

Valid

Amano

Late Pliocene

Nadachi Formation

 Japan

A species of Monodonta.

Murexsul primigenius[128]

Sp. nov

Valid

Merle & Pacaud

Eocene (Priabonian)

 France

A species of Murexsul.

Neverita amerrukensis[88]

Nom. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

A species of Neverita.

Nipteraxis parilis[88]

Sp. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

Novlepatella uralensis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Daminilidae.

Ocinebrina houarti[99]

Sp. nov

Valid

Landau et al.

Miocene (Tortonian)

 France

A species of Ocinebrina.

Olssonia[95]

Gen. et comb. nov

Valid

Sanders, Merle & Puillandre

Eocene to early Miocene

Chira Formation
Talara Formation

 Peru

A member of the family Bursidae. The type species is "Bursa" chira Olsson (1930); genus also includes Bursa chira var. yasila Olsson (1930), raised to the rank of a separate species Olssonia yasila.

Opaliopsis carlsbergi[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Orecopia inopinata[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Omphalotrochidae.

Orthonema conspicuum[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Orthonematidae.

Orthonema nonnullum[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Orthonematidae.

Orthonema perspicuum[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Orthonematidae.

Orthonema repandum[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Orthonematidae.

Orthonychia korotkovae[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Platyceratidae.

Palaeoloxotoma[86]

Nom. nov

Valid

Hansen

Early-Middle Jurassic to Late Cretaceous (Maastrichtian)

 Denmark
 France
 Jamaica
 New Zealand
 Poland

A member of the family Fissurellidae belonging to the subfamily Emarginulinae; a replacement name for Loxotoma Fischer (1885).

Palaeostylus confragosus[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Palaeostylidae.

Palaeostylus nervosus[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Palaeostylidae.

Payraudeautia ermesi[113]

Sp. nov

Valid

Pedriali, Sosso & Dell'Angelo

Miocene

 Ukraine

A species of Payraudeautia.

Payraudeautia sabrinae[113]

Sp. nov

Valid

Pedriali, Sosso & Dell'Angelo

Miocene

 Ukraine

A species of Payraudeautia.

Payraudeautia varovtsiana[113]

Sp. nov

Valid

Pedriali, Sosso & Dell'Angelo

Miocene

 Ukraine

A species of Payraudeautia.

Paziella gallica[99]

Sp. nov

Valid

Landau et al.

Miocene (Tortonian)

 France

A species of Paziella.

Paziella gracilenta[99]

Sp. nov

Valid

Landau et al.

Miocene (Tortonian)

 France

A species of Paziella.

Paziella modesta[106]

Sp. nov

Valid

Vicián, Kovács & Stein

Late Oligocene – early Miocene

 Hungary

A species of Paziella.

Paziella parveenae[128]

Sp. nov

Valid

Merle & Pacaud

Eocene (Ypresian)

 Pakistan

A species of Paziella.

Paziella zsoldosi[123]

Sp. nov

Valid

Kovács & Vicián

Eocene

 Hungary

A species of Paziella.

Perakella[129]

Gen. et 2 sp. nov

Valid

Mazaev

Permian

 Malaysia
 Russia

Genus includes new species P. shakhtauensis and P. batteni.

Peruvispira uralensis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Eotomariidae.

Phragmolites lissoni[130]

Sp. nov

Valid

Ebbestad & Gutiérrez-Marco

Ordovician (Sandbian)

Calapuja Formation

 Peru

Pictavia lactera[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of the family Ampullinidae.

Planiturbo lerensis[108]

Sp. nov

Valid

Szabó & Jaitly

Middle Jurassic (Callovian)

Chari Formation

 India

A member of the family Metriomphalidae.

Platyceras sagum[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

Platyzona shikhanensis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pithodeidae.

Pleurocera gigantica[103]

Sp. nov

Valid

Naranjo-García & Aguillón in Vega et al.

Paleocene

Las Encinas Formation

 Mexico

A species of Pleurocera.

Plocezyga boikoi[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudozygopleuridae.

Plocezyga convexa[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudozygopleuridae.

Plocezyga minuta[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudozygopleuridae.

Plocezyga venusta[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudozygopleuridae.

Ponticypraea[131]

Gen. et sp. nov

Valid

Fehse et al.

Late Eocene

 Australia

A member of the family Cypraeidae. Genus includes new species P. egregia.

Procerithium? inaequetuberculata[91]

Sp. nov

Valid

Ketwetsuriya, Cook & Nützel

Permian (Wordian)

Ratburi Group

 Thailand

A member of the family Procerithiidae.

Propeucyclus[100]

Gen. et 3 sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Genus includes new species P. sicanus, P. obesus and possibly also P? semireticulatus.

Provalvata maior[132]

Sp. nov

Valid

Cataldo et al.

Early Cretaceous (Barremian)

Huitrín Formation

 Argentina

Provalvata minor[132]

Sp. nov

Valid

Cataldo et al.

Early Cretaceous (Barremian)

Huitrín Formation

 Argentina

Provanna pelada[98]

Sp. nov

Valid

Kiel et al.

Early Oligocene

Talara Basin

 Peru

A species of Provanna.

Pseudocochlespira transversa[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Pseudolivella inadspecta[88]

Sp. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

Pseudotoma danica[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Pseudozygopleura applicata[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudozygopleuridae.

Ptychidia austrorotundata[89]

Nom. nov

Valid

Harzhauser & Landau

Early Miocene

 Austria
 Slovakia

A member of the family Turritellidae; a replacement name for Turritella turris rotundata Schaffer (1912).

Ptychidia erynella[89]

Sp. nov

Valid

Harzhauser & Landau

Early and middle Miocene

 Austria
 Czech Republic
 Hungary

A member of the family Turritellidae.

Pyramistomia aliakmoni[90]

Sp. nov

Valid

Thivaiou, Harzhauser & Koskeridou

Miocene (Aquitanian)

Pentalofos Formation

 Greece

A member of the family Pyramidellidae belonging to the subfamily Odostomiinae.

Pyrazopsis mexcala[101]

Sp. nov

Valid

Pacaud

 Mexico

Pyrgotrochus vorosi[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Pyropelta seca[98]

Sp. nov

Valid

Kiel et al.

Early Oligocene

Talara Basin

 Peru

A species of Pyropelta.

Ramusatomaria[100]

Gen. et sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Genus includes new species R. nuda.

Retimusina[100]

Gen. et 2 sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Genus includes new species R. poseidoni and possibly also R? tritoni.

Retispira khaophrikensis[91]

Sp. nov

Valid

Ketwetsuriya, Cook & Nützel

Permian (Wordian)

Ratburi Group

 Thailand

A member of the family Bellerophontidae belonging to the subfamily Knightitinae.

Rimella aldjazairensis[101]

Sp. nov

Valid

Pacaud

Eocene

 Algeria

A species of Rimella.

Rissoina albaresensis[101]

Sp. nov

Valid

Pacaud

Paleogene

 France

A species of Rissoina.

Rotfanella gerasimovi[93]

Sp. nov

Valid

Guzhov

Jurassic

 Russia

A member of the family Cimidae.

Rotfanella reticulata[93]

Sp. nov

Valid

Guzhov

Jurassic

 Russia

A member of the family Cimidae.

Rugosacyclus[104]

Gen. et comb. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Caenogastropoda of uncertain phylogenetic placement. The type species is "Spirocyclina" rugosa Brösamlen (1909).

Saalensia[104]

Gen. et sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Sorbeoconcha belonging to the family Brachytrematidae. The type species is S. birugata.

Sallya bimaris[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudophoridae.

Sallya indigena[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudophoridae.

Scelidotoma aldersoni[122]

Sp. nov

Valid

Powell & Geiger

Miocene

Topanga Canyon Formation

 United States
( California)

A species of Scelidotoma.

Schistoloma electrothauma[96]

Sp. nov

Valid

Asato & Hirano in Hirano et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Pupinidae.

Schistoloma electrothauma

Seleukella[92]

Gen. et sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Palaeozygopleuridae. Genus includes new species S. costata.

Semisulcospira (Biwamelania) gamoensis[133]

Sp. nov

Valid

Matsuoka & Miura

Early Pleistocene

Gamo Formation

 Japan

A species of Semisulcospira.

Semisulcospira (Biwamelania) nojirina[133]

Sp. nov

Valid

Matsuoka & Miura

Late Pliocene

Ayama Formation
Koka Formation

 Japan

A species of Semisulcospira.

Semisulcospira (Biwamelania) reticulataformis[133]

Sp. nov

Valid

Matsuoka & Miura

Late Pliocene

Ayama Formation

 Japan

A species of Semisulcospira.

Semisulcospira (Biwamelania) tagaensis[133]

Sp. nov

Valid

Matsuoka & Miura

Early Pleistocene

Gamo Formation

 Japan

A species of Semisulcospira.

Shurovites robustus[104]

Sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Cerithioidea belonging to the family Cryptaulacidae.

Shwedagonia korolukae[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Phymatopleuridae.

Shwedagonia pagoda[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Phymatopleuridae.

Sigmesalia ganntourensis[88]

Sp. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

Sigmesalia imitatrix[88]

Sp. nov

Valid

Pacaud & Lebrun

Late Cretaceous (Maastrichtian)

 Madagascar

Siphonochelus tambacounda[128]

Sp. nov

Valid

Merle & Pacaud

Eocene (Lutetian)

 Senegal

A species of Siphonochelus.

Solarioconulus kachchhensis[108]

Sp. nov

Valid

Szabó & Jaitly

Middle Jurassic (Callovian)

Chari Formation

 India

A member of the family Ataphridae.

Spiroscala elegans[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Phymatopleuridae.

Stegocoelia abscisa[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Orthonematidae.

Stegocoelia arta[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Orthonematidae.

Stegocoelia centrosinuata[91]

Sp. nov

Valid

Ketwetsuriya, Cook & Nützel

Permian (Wordian)

Ratburi Group

 Thailand

A member of the family Goniasmatidae.

Stenozone spiralis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Eotomariidae.

Stephanozyga seminuda[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Pseudozygopleuridae.

Streptacis nalivkini[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Streptacididae.

Strobeus bashkiriensi[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Soleniscidae.

Strophostylus fortuitus[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Platyceratidae.

Stuoraxis crassa[93]

Sp. nov

Valid

Guzhov

Jurassic

 Russia

A member of the family Stuoraxidae.

Sulcoactaeon? haboensis[108]

Sp. nov

Valid

Szabó & Jaitly

Middle Jurassic (Bathonian)

Patcham Formation

 India

A member of the family Aplustridae.

Sveltia castellum[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Sveltia.

Sveltia ruginosa[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Sveltia.

Sveltia varonei[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Sveltia.

Tapinotomaria cancellata[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Portlockiellidae.

Tatara pseudosassia[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Tectonatica anistratenkorum[113]

Sp. nov

Valid

Pedriali, Sosso & Dell'Angelo

Miocene

 Ukraine

A species of Tectonatica.

Tectonatica pseudoprietoi[113]

Sp. nov

Valid

Pedriali, Sosso & Dell'Angelo

Miocene

 Ukraine

A species of Tectonatica.

Tectus? indecorus[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Possibly a species of Tectus.

Temnotropis stevniensis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Termihabena[134]

Gen. et sp. et comb. nov

Valid

Mazaev

Early Permian

 Russia

Genus includes new species T. lirata, as well as Т. nikitowkensis (Yakowlew, 1899), T. asiatica (Licharew, 1967), T. millegranosa (Girty, 1934) and T. pinegensis (Mazaev, 2006).

Thereitis carlsbergi[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Thereitis cretacea[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Thylacodes contendis[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Thylacodes.

Toronyella[100]

Gen. et 2 sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Genus includes new species T. lineata and T. margaritata.

Torquesia (Ispharina) atax[101]

Nom. nov

Valid

Pacaud

Paleogene

 France

Trachydomia shkurkoi[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Trachyspiridae.

Trapanimaria[100]

Gen. et 3 sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Genus includes new species T. gattoi, T. nicolosiensis and possibly also T? pallinii.

Tritia albrechti[135]

Sp. nov

Valid

Stein

Miocene

North Sea basin

A species of Tritia.

Tritia girondica kautskyi[135]

Subsp. nov

Valid

Stein

Miocene

North Sea basin

Tritia hemmoorica[135]

Sp. nov

Valid

Stein

Miocene

North Sea basin

A species of Tritia.

Tritia jyllandica[135]

Sp. nov

Valid

Stein

Miocene

North Sea basin

A species of Tritia.

Tritia karinwienrichae[135]

Nom. nov

Valid

Stein

Miocene

 Germany

A species of Tritia; a replacement name for Nassarius karinae Wienrich (2001).

Tritia kostejana krocki[135]

Subsp. nov

Valid

Stein

Miocene

North Sea basin

Tritia mostafavii[135]

Nom. nov

Valid

Stein

Miocene

 Germany

A species of Tritia; a replacement name for Hinia coronata Mostafavi (1978).

Tritia mothsi[135]

Sp. nov

Valid

Stein

Miocene

North Sea basin

A species of Tritia.

Tritia sorgenfreii[135]

Sp. nov

Valid

Stein

Miocene

North Sea basin

A species of Tritia.

Tritia westfalica[135]

Sp. nov

Valid

Stein

Miocene

North Sea basin

A species of Tritia.

Trochotomaria conoidea[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Trochotomaria polymorpha[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Tropacerithium[104]

Gen. et 2 sp. nov

Valid

Gründel, Keupp & Lang

Late Jurassic (Kimmeridgian)

 Germany

A member of Cerithioidea possibly belonging to the family Cryptaulacidae. The type species is T. cumaritum; genus also includes T. danubii.

Tychonia fortis[92]

Sp. nov

Valid

Mazaev

Early Permian

 Russia

A member of the family Anomphalidae.

Typhina cryptica[128]

Sp. nov

Valid

Merle & Pacaud

Eocene (Ypresian)

 France

A species of Typhina.

Typhina radulfiensis[128]

Sp. nov

Valid

Merle & Pacaud

Eocene (Priabonian)

 France

A species of Typhina.

Unitas aequalis[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Cancellariidae.

Unitas aquitanica[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Cancellariidae.

Unitas arenosa[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Cancellariidae.

Unitas decorticata[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Cancellariidae.

Unitas gradata[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Cancellariidae.

Unitas heinbergi[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Unitas sanctistephani[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Cancellariidae.

Unitas substephanensis[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Cancellariidae.

Urlocella undulata[93]

Sp. nov

Valid

Guzhov

Jurassic

 Russia

A member of the family Cimidae.

Unzhispira[93]

Gen. et sp. nov

Valid

Guzhov

Jurassic

 Russia

A member of the family Cimidae. Genus includes new species U. minuta.

Vatopsis heinbergi[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Vermeijia[116]

Gen. et sp. nov

Valid

Amano

Late Pliocene

Kuwae Formation

 Japan

A member of the family Capulidae. Genus includes new species V. japonica.

Vernelia samae[115]

Sp. nov

Valid

Foster et al.

Early Triassic

 China

Vexillum? rosenkrantzi[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

Possibly a species of Vexillum.

Vexillum tenestolidum[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Vexillum.

Vicinocerithium vynckei[101]

Nom. nov

Valid

Pacaud

Paleogene

 France

Vicnigoria[92]

Gen. et sp. nov

Valid

Mazaev

Early Permian

 Russia

Probably a member of the family Trochonematidae. Genus includes new species V. monstrata.

Viennella[89]

Gen. et comb. et sp. nov

Valid

Harzhauser & Landau

Middle Miocene

 Austria
 Hungary

A member of the family Turritellidae. The type species is "Turritella" incisaeformis Csepreghy-Meznerics (1956); genus also includes new species V. ignorata.

Viviparus riegensis[101]

Nom. nov

Valid

Pacaud

Paleogene

 France

A species of Viviparus.

Volvarina sibuzatiana[94]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Volvarina.

Wateletia custodia[101]

Nom. nov

Valid

Pacaud

Paleogene

 France

Worthenia? waterhouse[91]

Sp. nov

Valid

Ketwetsuriya, Cook & Nützel

Permian (Wordian)

Ratburi Group

 Thailand

A member of the family Lophospiridae.

Zarnglaffia palermitana[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Zarnglaffia polygonalis[100]

Sp. nov

Valid

Szabó et al.

Jurassic

 Italy

Zebinella selandica[86]

Sp. nov

Valid

Hansen

Cretaceous-Paleogene boundary

 Denmark

A species of Zebinella.

Zizipupa[93]

Gen. et sp. nov

Valid

Guzhov

Jurassic

 Russia

A member of the family Ampezzanildidae. Genus includes new species Z. costata.

Other molluscs

Research

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Acharax svalbardensis[148]

Sp. nov

Valid

Hansen et al.

Late Pleistocene

Fram Strait

A bivalve belonging to the family Solemyidae.

Atomodesma? hautmanni[115]

Sp. nov

Valid

Foster et al.

Early Triassic

 China

A bivalve.

Atrina multicostata[149]

Sp. nov

Valid

Friesenbichler et al.

Middle Triassic (Anisian)

Caerace Formation

 Romania

A bivalve, a species of Atrina.

Baxoitrigonia[150]

Gen. et sp. nov

Stiller & Chen

Late Jurassic (Tithonian)

Lagongtang Formation

 China

A bivalve. Genus includes new species B. baxoiensis.

Bengtsonella[151]

Gen. et comb. nov

Junior homonym

Cooper & Leanza

Early Cretaceous (Albian)

Riachuelo Formation

 Brazil

A bivalve belonging to the family Megatrigoniidae and the subfamily Megatrigoniinae. The type species is "Anditrigonia" britoi Hessel (2005). The generic name is preoccupied by Bengtsonella Muller & Hinz (1991) and Bengtsonella Mostler (1996).

Californigonia[151]

Gen. et comb. nov

Valid

Cooper & Leanza

Jurassic (Callovian–early Oxfordian)

Bicknell Sandstone

 Canada
( British Columbia)
 United States
( California)

A bivalve belonging to the subfamily Megatrigoniinae. The type species is "Trigonia" plumasensis Hyatt (1892).

Callocardia (Nitidavenus) youssoufia[88]

Sp. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

A bivalve.

Callucina itaporangensis[152]

Sp. nov

Valid

Ayoub-Hannaa et al.

Cretaceous

Cotinguiba Formation

 Brazil
 United Kingdom

A bivalve belonging to the family Lucinidae.

Cardiocardita peregrina[88]

Sp. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

A bivalve.

Carditella pitufina[153]

Sp. nov

Valid

Pérez

Early Pliocene

 Argentina

A bivalve belonging to the family Condylocardiidae.

Chlamys (Praechlamys) prima[149]

Sp. nov

Valid

Friesenbichler et al.

Middle Triassic (Anisian)

Caerace Formation

 Romania

A bivalve, a species of Chlamys.

Ckaraosippur[154]

Gen. et comb. et sp. nov

Valid

Santelli & del Río

MiocenePliocene

Bahía Inglesa Formation
Camarones Formation
Navidad Formation

 Argentina
 Chile

A scallop. The type specie is "Pecten" calderensis Möricke (1896); genus also includes new species C. camachoi.

Conchocele yatsuoensis[155]

Sp. nov

Valid

Amano et al.

Early Miocene

Kurosedani Formation

 Japan

A bivalve.

Costacallista discrepans[88]

Sp. nov

Valid

Pacaud & Lebrun

Eocene (Lutetian)

 Morocco

A bivalve.

Craginella[151]

Gen. et comb. nov

Valid

Cooper & Leanza

Late Jurassic (Kimmeridgian)

Malone Formation

 Mexico
 United States
( Louisiana
 Texas)

A bivalve belonging to the subfamily Megatrigoniinae. The type species is "Trigonia" goodellii Cragin (1897).

Damborenella[151]

Gen. et comb. nov

Valid

Cooper & Leanza

Late Jurassic (Tithonian)

 Argentina
 Chile
 Colombia
 Peru

A bivalve belonging to the subfamily Megatrigoniinae. The type species is "Trigonia" eximia Philippi (1899); genus also includes "Trigonia" discors Philippi (1899), "Anditrigonia" lamberti Levy (1967) and "Anditrigonia eximia" tesselicaudata Leanza (1993) (raised to the rank of a separate species Damborenella tessellicaudata).

Dentalium mcganna[156]

Sp. nov

Valid

Powell, Clites & Poust

Miocene

Sobrante Formation

 United States
( California)

A species of Dentalium.

Dietotenhosen[154]

Gen. et comb. nov

Valid

Santelli & del Río

MiocenePliocene

Bahía Inglesa Formation
Coquimbo Formation
Horcón Formation
Hornillos Formation
Huaricangana Formation
La Portada Formation
Pisco Formation
Taime Formation

 Chile
 Peru

A scallop. Genus includes D. hupeanus (Philippi, 1887) and D. remondi (Philippi, 1887).

Dosinia (Dosinisca) vredenburgi[157]

Sp. nov

Valid

Borkar & Kulkarni

Miocene

Chhasra Formation
Khari Nadi Formation
Lower Mekran Series

 India
 Pakistan

A bivalve belonging to the family Veneridae; a species of Dosinia.

Dosinia (Dosinisca) wynnei[157]

Sp. nov

Valid

Borkar & Kulkarni

Miocene

Chhasra Formation
Khari Nadi Formation
Lower Mekran Series

 India
 Pakistan

A bivalve belonging to the family Veneridae; a species of Dosinia.

Entolium reticulatum[149]

Sp. nov

Valid

Friesenbichler et al.

Middle Triassic (Anisian)

Caerace Formation

 Romania

A bivalve.

Eosphaera[158]

Gen. et comb. nov

Valid

Fürsich, Heinze & Ayoub-Hannaa

Jurassic (Callovian to Kimmeridgian)

 Egypt
 India
 Madagascar
 Portugal

A bivalve belonging to the family Lucinidae. The type species is "Sphaera" madagascariensis Newton (1889).

Chaetopleura abbessi[159]

Sp. nov

Valid

Cherns & Schwabe

Oligocene (Chattian) and Miocene (Burdigalian)[160]

 France

A chiton. Originally described as a species of Chaetopleura; subsequently transferred to the genus Ischnochiton by Dell’Angelo et al. (2018).[160]

Chaetopleura gaasi[159]

Sp. nov

Valid

Cherns & Schwabe

Oligocene (Rupelian)

 France

A chiton, Originally described as a species of Chaetopleura; subsequently transferred to the new genus Spinochiton by Dell’Angelo et al. (2018).[160]

Helcionella lemdadensis[161]

Sp. nov

Valid

Geyer, Valent & Meier

Cambrian

Tannenknock Formation

 Germany

Hoernesia? danisa[115]

Sp. nov

Valid

Foster et al.

Early Triassic

 China

A bivalve.

Homopleura[162]

Gen. et comb. et 2 sp. nov

Valid

Masse & Fenerci-Masse

Cretaceous

 Bulgaria
 France

A rudist bivalve. The type species is "Monopleura" affinis Matheron (1878); genus also includes "Monopleura" imbricata Matheron (1842) and "Monopleura" coquandi Matheron (1878), as well as new species H. rustrella and H. balkanica.

Inoceramus chiplonkari[27]

Sp. nov

Valid

Walaszczyk in Gale, Kennedy & Walaszczyk

Late Cretaceous (Cenomanian and Turonian)

Karai Formation

 India
 Japan

Ischnochiton crovatoi[163]

Sp. nov

Valid

Dell’Angelo, Sosso & Bonfitto

Pleistocene

 Italy

A chiton, a species of Ischnochiton.

Ischnochiton fehsei[159]

Sp. nov

Valid

Cherns & Schwabe

Eocene

 France

A chiton, a species of Ischnochiton.

Lasaea firma[164]

Sp. nov

Valid

Lesport & Lozouet in Lesport et al.

Miocene (Aquitanian)

 France

A bivalve belonging to the family Lasaeidae.

Leptostega frankenwaldensis[161]

Sp. nov

Valid

Geyer, Valent & Meier

Cambrian

Tannenknock Formation

 Germany

A helcionelloid.

Neilo altamirano[98]

Sp. nov

Valid

Kiel et al.

Early Oligocene

Talara Basin

 Peru

A bivalve belonging to the family Malletiidae.

Neocrassina (Pruvostiella) madagascariensis[158]

Sp. nov

Valid

Fürsich, Heinze & Ayoub-Hannaa

Late Jurassic (Oxfordian)

Morondava Basin

 Madagascar

A bivalve belonging to the family Astartidae.

Palaeonucula feruglioi[165]

Sp. nov

Valid

Damborenea & Pagani

Early Jurassic

 Argentina

A bivalve.

Parailsanella luonanensis[166]

Sp. nov

Valid

Li et al.

Cambrian Series 2

Xinji Formation

 China

A helcionellid.

Peronaea basteroti[164]

Sp. nov

Valid

Lesport, Lozouet & Pacaud in Lesport et al.

Miocene (Aquitanian)

 France

A bivalve, a species of Peronaea.

Phaenodesmia? piatnitzkyi[165]

Sp. nov

Valid

Damborenea & Pagani

Early Jurassic

 Argentina

A bivalve.

Pinna simionescui[149]

Sp. nov

Valid

Friesenbichler et al.

Middle Triassic (Anisian)

Caerace Formation

 Romania

A bivalve, a species of Pinna.

Platymyoidea morrisi[167]

Sp. nov

Valid

Hodges

Early Jurassic

 United Kingdom

A bivalve.

Pleurophopsis matsumotoi[168]

Sp. nov

Valid

Amano et al.

Late Oligocene to early Miocene

Nabae Group

 Japan

A bivalve belonging to the family Vesicomyidae.

Pleurophopsis talarensis[98]

Sp. nov

Valid

Kiel et al.

Early Oligocene

Talara Basin

 Peru

A bivalve belonging to the family Vesicomyidae.

Praedicerocardium[149]

Gen. et sp. nov

Valid

Friesenbichler et al.

Middle Triassic (Anisian)

Caerace Formation

 Romania

A bivalve. Genus includes new species P. vetulus.

Procardia inouei[169]

Sp. nov

Valid

Amano

Paleocene

Katsuhira Formation

 Japan

A bivalve belonging to the group Anomalodesmata and to the family Parilimyidae.

Pseudophopsis[98]

Gen. et comb. nov

Valid

Kiel et al.

Eocene and Early Oligocene

Talara Basin

 Cuba
 Peru

A bivalve belonging to the group Heterodonta and to the family Kalenteridae. The type species is "Pleurophopsis" peruviana Olsson (1931); genus also includes "Unio" bitumen Cooke (1919).

Quenstedtia madagascariensis[158]

Sp. nov

Valid

Fürsich, Heinze & Ayoub-Hannaa

Late Jurassic (Oxfordian)

Morondava Basin

 Madagascar

A bivalve belonging to the family Quenstedtiidae.

Ramentoides xixiangensis[170]

Sp. nov

Valid

Qin et al.

Cambrian

 China

A member of Maikhanellidae.

Romaniamya[149]

Gen. et sp. nov

Valid

Friesenbichler et al.

Middle Triassic (Anisian)

Caerace Formation

 Romania

A bivalve. Genus includes new species R. mahmudiaensis.

Rutitrigonia cintarojensis[132]

Sp. nov

Valid

Cataldo et al.

Early Cretaceous (Barremian)

Huitrín Formation

 Argentina

A bivalve.

Ryderia tehuelchana[165]

Sp. nov

Valid

Damborenea & Pagani

Early Jurassic

 Argentina

A bivalve.

Scythentolium anisicum[149]

Sp. nov

Valid

Friesenbichler et al.

Middle Triassic (Anisian)

Caerace Formation

 Romania

A bivalve.

Sogdianagonia[171]

Nom. nov

Valid

Schneider & Kelly

Late Cretaceous (Cenomanian to Turonian)

 Kyrgyzstan
 Uzbekistan

A bivalve belonging to the family Megatrigoniidae; a replacement name for Turkestanella Tashiro (1979). The type species is "Trigonia" turkestanensis Arkhangelsky (1916).

Solemya lucifuga[172]

Sp. nov

Valid

Pérez-Barría & Nielsen

Early Miocene

Ranquil Formation

 Chile

A bivalve, a species of Solemya.

Stenoplax monila[159]

Sp. nov

Valid

Cherns & Schwabe

Oligocene (Rupelian)

 France

A chiton, a species of Stenoplax.

Tindaria kretensis[173]

Sp. nov

Valid

Koskeridou, La Perna & Giamali

Early Pliocene

 Greece

A bivalve.

Tonicella lira[159]

Sp. nov

Valid

Cherns & Schwabe

Oligocene (Chattian)

 France

A chiton, a species of Tonicella.

Vaccinites alceotarlaoi[174]

Sp. nov

Valid

Pons et al.

Late Cretaceous (early Campanian)

 Serbia
 Spain

A rudist bivalve.

Vaccinites bilottei[174]

Sp. nov

Valid

Pons et al.

Late Cretaceous (late Campanian)

 Bulgaria

A rudist bivalve.

Vaccinites pirenaicus[174]

Sp. nov

Valid

Pons et al.

Late Cretaceous (late Campanian)

 Spain

A rudist bivalve.

Wareniconcha mercenarioides[175]

Sp. nov

Valid

Kase et al.

Pliocene

 Philippines

A bivalve belonging to the family Vesicomyidae.

Xylophagella littlei[176]

Sp. nov

Valid

Hryniewicz in Hryniewicz et al.

Late Paleocene

Basilika Formation

 Norway

A bivalve belonging to the family Pholadidae.

Yoldiella spitsbergensis[176]

Sp. nov

Valid

Amano in Hryniewicz et al.

Late Paleocene

Basilika Formation

 Norway

A bivalve belonging to the family Yoldiidae.

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References

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  2. Carlie Pietsch; Kathleen A. Ritterbush; Jeffrey R. Thompson; Elizabeth Petsios; David J. Bottjer (2019). "Evolutionary models in the Early Triassic marine realm". Palaeogeography, Palaeoclimatology, Palaeoecology. 513: 65–85. Bibcode:2019PPP...513...65P. doi:10.1016/j.palaeo.2017.12.016.
  3. Nicholas A. Freymueller; Jason R. Moore; Corinne E. Myers (2019). "An analysis of the impacts of Cretaceous oceanic anoxic events on global molluscan diversity dynamics". Paleobiology. 45 (2): 280–295. doi:10.1017/pab.2019.10.
  4. Tingting Yu; Richard Kelly; Lin Mu; Andrew Ross; Jim Kennedy; Pierre Broly; Fangyuan Xia; Haichun Zhang; Bo Wang; David Dilcher (2019). "An ammonite trapped in Burmese amber". Proceedings of the National Academy of Sciences of the United States of America. 116 (23): 11345–11350. doi:10.1073/pnas.1821292116. PMC 6561253. PMID 31085633.
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