2020 in paleomalacology

This list 2020 in paleomalacology is a list of new taxa of ammonites and other fossil cephalopods, as well as fossil gastropods, bivalves and other molluscs that are scheduled to be described during the year 2020, as well as other significant discoveries and events related to molluscan paleontology that are scheduled to occur in the year 2020.

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Ammonites

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Angolaites elegans[1]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Aulacostephanus camericensis[2]	Sp. nov	In press	Cope & Etches	Late Jurassic (Kimmeridgian)		United Kingdom
Baidites obesus[3]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Borissjakoceras falcatum[4]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Tunisia	A member of the family Binneyitidae.
?Bouleiceras betetensis[5]	Sp. nov	In press	Martínez & Joral	Early Jurassic (Toarcian)		Spain	A member of the family Hildoceratidae.
Bouleiceras ibericum[5]	Sp. nov	In press	Martínez & Joral	Early Jurassic (Toarcian)		Spain	A member of the family Hildoceratidae.
Cagliceras baldii[6]	Sp. nov	Valid	Kovács, Dunai & Evanics	Early Jurassic (Toarcian)		Hungary	A member of the family Erycitidae.
Cagliceras wegeneri[6]	Sp. nov	Valid	Kovács, Dunai & Evanics	Early Jurassic (Toarcian)		Hungary	A member of the family Erycitidae.
Carthaginites elegans[4]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Turrilitidae.
Churkites warei[3]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Collignoniceras badilleti[7]	Sp. nov	Valid	Amédro & Châtelier in Amédro et al.	Late Cretaceous (Turonian)		France
Collignoniceras hourqueigi[7]	Sp. nov	Valid	Amédro & Châtelier in Amédro et al.	Late Cretaceous (Turonian)		France
Collignoniceras vigennum[7]	Sp. nov	Valid	Amédro & Châtelier in Amédro et al.	Late Cretaceous (Turonian)		France
Conlinites evolutum[4]	Sp. nov	Valid	Kennedy	Early Cretaceous (Albian)		Algeria	A member of the family Brancoceratidae.
Coquandiceras[4]	Gen. et comb. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria  Tunisia	A member of the family Acanthoceratidae belonging to the subfamily Mantelliceratinae. The type species is "Ammonites" jubae Coquand (1880); genus also includes "Ammonites" villei Coquand (1862).
Cryptoturrilites[4]	Gen. et sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Tunisia	A member of the family Turrilitidae. The type species is C. tenuicostatus.
Elobiceras (Craginites) simplex[1]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Elobiceras (Craginites) sparcicostatum[1]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Flemingites lidakensis[3]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Goniatites abaiensis[8]	Sp. nov	Valid	Korn et al.	Carboniferous (Viséan)	Baktysai Formation	Kazakhstan
Goniatites zhankurganensis[8]	Sp. nov	Valid	Korn et al.	Carboniferous (Viséan)	Baktysai Formation	Kazakhstan
Graysonites elegans[4]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Acanthoceratidae.
Gregoryceras boreale[9]	Sp. nov	Valid	Mitta & Stupachenko	Late Jurassic (Oxfordian)		Russia	A member of the family Aspidoceratidae.
Homeomorphites[10]	Gen. et sp. et comb. nov	In press	Bert, Bersac & Canut	Early Cretaceous (Hauterivian to Barremian)		Argentina  Chile	A member of the family Neocomitidae. The type species is H. aguirreurretae; genus also includes H. ploszkiewiczi (Riccardi and Aguirre-Urreta, 1989) and H. varicostatus (Riccardi and Aguirre-Urreta, 1989).
Homoceratoides wangchengi[11]	Sp. nov	In press	Korn & Wang in Korn et al.	Carboniferous (Bashkirian)		China
Howarthiceras[12]	Gen. et comb. nov	In press	Zakharov & Rogov	Upper Jurassic (Tithonian)		United States ( California)	A new genus for "Groebericeras" baylei Imlay & Jones.
Hypowaagenia[13]	Gen. et sp. nov	Valid	Schweigert & Schlampp	Late Jurassic (Kimmeridgian)	Arzberg Formation	Germany  Pakistan?	A member of the family Aspidoceratidae. The type species is H. endressi; genus might also include "Aspidoceras" acanthomphalum Zittel (1870).
Kosmoclymenia ebbighauseni[14]	Sp. nov	Valid	Korn	Devonian (Famennian)		Morocco
Lasswitzia[1]	Gen. et comb. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek Formation	United States ( Texas)	A member of the family Brancoceratidae. The type species is "Schloenbachia austinensis" Roemer var. minima Lasswitz (1904) (raised to the rank of a separate species Lasswitzia minima).
Lenicostites[10]	Gen. et comb. nov	In press	Bert, Bersac & Canut	Early Cretaceous (Barremian)		France	A member of Ancyloceratoidea belonging to the new family Lenicostitidae. The type species is "Hemihoplites" rusticus Vermeulen (1996).
Litophragmatoceras curiosus[4]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Flickiidae.
Mittaites[15]	Gen. et comb. nov	In press	Frau et al.	Early Cretaceous (Berriasian)		Russia	The type species is "Mazenoticeras" ceccai.
Mortoniceratoides elegans[1]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Kiamichi	United States ( Texas)	A member of the family Brancoceratidae.
Muessenbiaergia bockwinkeli[14]	Sp. nov	Valid	Korn	Devonian (Famennian)		Morocco	A member of the family Kosmoclymeniidae.
Neophlycticeras algeriense[4]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Lyelliceratidae.
Neotelodactylites[16]	Gen. et comb. nov	Valid	Jattiot, Gris & Trincal	Early Jurassic (Toarcian)		France  Italy	A new genus for "Peronoceras" eucosmum Lippi Boncambi (1947).
Paracraginites[1]	Gen. et sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae. The type species is P. conlini.
Paraspidites bicarinatus[3]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Pervinquieria (Deiradoceras) serpentiforme[1]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Pervinquieria (Pervinquieria) asper[1]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Protacanthoceras sottaraense[4]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Acanthoceratidae.
Ptychites embreei[17]	Sp. nov	Valid	Bischof & Lehmann	Middle Triassic (Anisian)	Favret Formation	United States ( Nevada)
Romaniceras (Romaniceras) marigniacum[7]	Sp. nov	Valid	Amédro & Châtelier in Amédro et al.	Late Cretaceous (Turonian)		France
Roopnarinites[3]	Gen. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Scaphites occlusus[4]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria  Tunisia	A member of the family Scaphitidae.
Stoliczkaia (Stoliczkaia) djerissaensis[4]	Sp. nov	Valid	Kennedy	Early Cretaceous (Albian)		Tunisia	A member of the family Lyelliceratidae.
Subflemingites bihatiense[3]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia

General research

• A study aiming to determine the drag force experienced by a wide range of ammonite shell shapes is published by Hebdon, Ritterbush & Choi (2020).[18]
• A study on the abundance and size distribution of small specimens of Baculites and Hoploscaphites at eight cold methane seep sites in the Upper Cretaceous Pierre Shale (South Dakota, United States) is published by Rowe et al. (2020), who interpret their findings as indicating that newly hatched ammonites lived in close proximity to seep fluids emerging at the sediment-water interface and the associated microbial food web.[19]
• A study on changes of hydrostatic properties of three species of Didymoceras during their ontogeny is published by Peterman et al. (2020).[20]

Other cephalopods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Ancistroceras ristnensis[21]	Sp. nov	In press	Aubrechtová & Meidla	Ordovician		Estonia
Ancistroceras vahikuelaensis[21]	Sp. nov	In press	Aubrechtová & Meidla	Ordovician		Estonia
Chromatoceras[22]	Gen. et comb. nov	Valid	Turek & Manda	Silurian (Pridoli)	Kopanina	Czech Republic	A member of Oncocerida belonging to the family Oonoceratidae. The type species is "Cyrtoceras" veteranum Barrande (1866); genus also includes "Cyrtoceras" constringens Barrande (1866).
Lituites nehatuensis[21]	Sp. nov	In press	Aubrechtová & Meidla	Ordovician		Estonia
Tafadnatoceras[23]	Gen. et sp. nov	In press	Ebbestad et al.	Late Ordovician	Upper Tiouririne Formation	Morocco	A member of Orthocerida belonging to the family Stereoplasmoceratidae. The type species is T. tiouririnense.
Wadema tattai[23]	Sp. nov	In press	Ebbestad et al.	Late Ordovician	Upper Ktaoua Formation	Morocco	A member of Actinocerida belonging to the family Wademidae.

General research

• A study aiming to determine whether the development of chamber volume in fossil cephalopod phragmocones conveys information about their physiology is published by Tajika et al. (2020), who evaluate the implications of their findings for the knowledge of the ecology and extinction selectivity of fossil cephalopods.[24]
• A study on the spatial distribution of Givetian oncocerid fossils from the Hamar Laghdad ridge (eastern Anti-Atlas, Morocco), evaluating its implications for the knowledge of the life cycle of these cephalopods, is published by Pohle et al. (2020).[25]
• A study on the reproductive strategies of fossil coleoids, based on data from shells of belemnoids, spirulids and sepiids, is published by Fuchs et al. (2020).[26]
• A specimen of Plesioteuthis preserved with an associated tooth of a pterosaur is reported from the Upper Jurassic Altmühltal Formation (Germany) by Hoffmann et al. (2020), who evaluate the implications of this finding for the knowledge of the likely habitat of Plesioteuthis.[27]

Gastropods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Alanstukella[28]	Gen. et comb. nov	Valid	Mazaev	Carboniferous (Kasimovian) to Permian (Sakmarian)		Russia	A member of the family Trochonematidae. The type species is A. rossica (Stuckenberg, 1905).
Allossospira[23]	Gen. et comb. nov	In press	Ebbestad et al.	Late Ordovician	Upper Ktaoua Formation	Morocco	A member of the family Gosseletinidae belonging to the subfamily Euryzoninae. The type species is "Lophospira" debganensis Horný (1997).
Altotomaria[29]	Gen. et sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand	Genus includes new species A. reticulata.
Apachella thailandensis[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Assiminea striatura[30]	Sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Assiminea.
Balticopta[31]	Gen. et sp. nov	Valid	Balashov & Perkovsky	Eocene (Priabonian)	Baltic amber	Russia ( Kaliningrad Oblast)	A member of the family Gastrocoptidae. The type species is B. gusakovi.
Bania goehlichae[32]	Sp. nov	Valid	Mandic, Harzhauser & Neubauer	Middle Miocene		Bosnia and Herzegovina
Belgrandiella? nichterli[33]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain	Possibly a species of Belgrandiella.
Bellerophon erawanensis[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Biarmeaspira mazaevi[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Calculocochlea[34]	Gen. et sp. nov	Valid	Juárez-Ruiz, Matamales-Andreu & Kadolsky	Early Oligocene		Spain	A member of the family Vidaliellidae. Genus includes new species C. oliveri.
Calogoniodiscus pyrenaicus[33]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Cheilospicata cedrus[35]	Sp. nov	Valid	Garvie in Garvie, Goedert & Janssen			United States
Chilodonta tricostatula[36]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A species of Chilodonta.
Clanculus (Clanculopsis) krachi[37]	Sp. nov	In press	Nosowska	Middle Miocene		Poland  Ukraine	A species of Clanculus.
Clio gailae[35]	Sp. nov	Valid	Goedert & Janssen in Garvie, Goedert & Janssen			United States ( Washington)	A species of Clio.
Cochleochilus? spiralocostatus[36]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	Possibly a member of the family Ataphridae.
Currylimacina asperita[35]	Sp. nov	Valid	Garvie in Garvie, Goedert & Janssen			United States
Emarginula rara[36]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A species of Emarginula.
Enneopupa priabonica[33]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Eohispania[33]	Gen. et sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain	Genus includes new species E. dominicii.
Eotrichophorus[30]	Gen. et sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Genus includes new species E. kachin.
Fusulculus hanseni[38]	Nom. nov	Valid	Pacaud	Early Paleocene		Denmark	A species of Fusulculus; a replacement name for Epalxis ? rosenkrantzi Hansen (2019).
Gosseletina microstriata[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Headonia[33]	Gen. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Heliconoides hodgkinsoni[35]	Sp. nov	Valid	Garvie in Garvie, Goedert & Janssen			United States
Helvetataphrus[36]	Gen. et comb. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	France   Switzerland	A member of the family Ataphridae. The type species is "Turbo" subrugosus Buvignier (1852).
Heterogen praelongispira[39]	Sp. nov	Valid	Matsuoka & Hirano	Pleistocene	Katata Formation	Japan	A species of Heterogen.
Heterosubulites longusapertura[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Incisilabium trochiformis[40]	Sp. nov	Valid	Lozouet, Cahuzac & Charles	Eocene (Priabonian)		France	A member of the family Modulidae.
Katosira ? bicarinata[41]	Sp. nov	Valid	Ferrari, Little & Atkinson	Early Jurassic (Toarcian)	Blea Wyke Sandstone Formation	United Kingdom	A member of the family Zygopleuridae.
Laminifera nannodes[33]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Laxecostula[33]	Gen. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Limacina parvabrazensis[35]	Sp. nov	Valid	Garvie & Janssen in Garvie, Goedert & Janssen			United States	A species of Limacina.
Limacina pseudopygmaea[35]	Sp. nov	Valid	Garvie & Janssen in Garvie, Goedert & Janssen			United States	A species of Limacina.
Limacina texanopsis[35]	Sp. nov	Valid	Garvie in Garvie, Goedert & Janssen			United States	A species of Limacina.
Limacina yadongensis[42]	Sp. nov	Valid	Li et al.	Early Eocene	Zhepure Formation	China	A species of Limacina.
Lophospira latilabra[23]	Sp. nov	In press	Ebbestad et al.	Late Ordovician	Upper Ktaoua Formation	Morocco	A member of the family Lophospiridae.
Loriolotrema[43]	Gen. et sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Jurassic		Europe	A member of the family Brachytrematidae. Genus includes new species L. liriola.
Lozouetina naonedosca[44]	Sp. nov	Valid	Pacaud & Gómez	Eocene (Bartonian)		France  Spain	A member of the family Ovulidae belonging to the subfamily Pediculariinae.
Macropupina[30]	Gen. et sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Genus includes new species M. electricus.
Modulus benoisti[40]	Sp. nov	Valid	Lozouet, Cahuzac & Charles	Miocene (Serravallian)		France	A species of Modulus.
Neniopsilla[33]	Gen. et sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain	Genus includes new species N. amoena.
Niveria ledoni[45]	Sp. nov	Valid	Fehse & Vicián	Late Miocene		France	A species of Niveria.
Niveria miocarinata[45]	Sp. nov	Valid	Fehse & Vicián	Late Miocene		France	A species of Niveria.
Oxfordina[36]	Gen. et comb. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A member of the family Eucyclidae. The type species is "Trochus" crassicosta Buvignier (1852).
Paleodiplommatina[30]	Gen. et sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Genus includes new species P. spelomphalos.
Paracraticula? catalaunica[33]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Patella staceata[36]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A species of Patella.
Perissocyclos[30]	Gen. et sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Genus includes new species P. kyrtostoma.
Peruvispira brasilensis[46]	Sp. nov	In press	Simões et al.	Late Paleozoic	Taciba Formation	Brazil
Platyzona gradata[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Profundimitra lacuiensis[47]	Nom. nov	Valid	Nielsen & Ampuero	Early Miocene		Chile	A species of Profundimitra; a replacement name for Mitra martini Philippi (1887).
Prososthenia krijgsmani[32]	Sp. nov	Valid	Mandic, Harzhauser & Neubauer	Middle Miocene		Bosnia and Herzegovina
Prososthenia milosevici[48]	Sp. nov	Valid	Neubauer et al.	Miocene		Serbia	A member of the family Hydrobiidae belonging to the subfamily Pyrgulinae.
Prososthenia? naissensis[48]	Sp. nov	Valid	Neubauer et al.	Miocene		Serbia	A member of the family Hydrobiidae belonging to the subfamily Pyrgulinae.
Prososthenia rundici[48]	Sp. nov	Valid	Neubauer et al.	Miocene		Serbia	A member of the family Hydrobiidae belonging to the subfamily Pyrgulinae.
Pseudangaria[36]	Gen. et comb. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A member of the family Angariidae. The type species is "Delphinula" kobyi De Loriol in De Loriol & Koby (1895).
Pseudotectonica[33]	Gen. et sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain	Genus includes new species P. pulchra.
Radvanospira[23]	Gen. et comb. et sp. nov	In press	Ebbestad et al.	Late Ordovician		Czech Republic  Morocco  Spain	A member of the family Platyceratidae. The type species is "Naticopsis" antiquata Barrande in Perner (1903); genus also includes new species R. baniensis.
Rimulopsis torulosus[36]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A member of the family Fissurellidae.
Scissurella nesbittae[49]	Sp. nov	Valid	Geiger & Goedert	Eocene or Oligocene (latest Priabonian or earliest Rupelian)	Gries Ranch Formation	United States ( Washington)	A species of Scissurella.
Scutus mirus[50]	Sp. nov	Valid	Gard	Late Oligocene to early Miocene		New Zealand	A species of Scutus.
Seguenzia statiana[51]	Sp. nov	Valid	Sosso, Bertolaso & Dell’Angelo	Miocene (Tortonian)	Marne di S. Agata Fossili Formation	Italy	A species of Seguenzia.
Selvovum[34]	Gen. et comb. nov	Valid	Juárez-Ruiz, Matamales-Andreu & Kadolsky	Late Eocene		Spain	A member of the family Vidaliellidae; a new genus for "Ampullaria" selvensis.
Steenbergia wohlmuthorum[52]	Sp. nov	Valid	Walther & Groh	Late Pleistocene		Madeira	A member of the family Geomitridae.
Stephaniphera[53]	Gen. et sp. nov	Valid	Schneider, Kollmann & Pickford	Late Cretaceous	Al-Khodh Formation	Oman	A member of Cerithioidea belonging to the family Hemisinidae. The type species is S. coronata.
Strebloceras kobayashii[54]	Sp. nov	Valid	Haga	Middle Pleistocene	Toyohashi Formation	Japan	A member of the family Caecidae.
Streptacis? khaokhadensis[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Striactaenonina elegans[41]	Sp. nov	Valid	Ferrari, Little & Atkinson	Early Jurassic (Toarcian)	Blea Wyke Sandstone Formation	United Kingdom	A member of Architectibranchia belonging to the family Tubiferidae.
Subtemenia[53]	Gen. et comb. nov	Valid	Schneider, Kollmann & Pickford	Late Cretaceous	Al-Khodh Formation	Iran  Oman	A member of the family Pseudomelaniidae. The type species is "Paryphostoma" morgani Douvillé (1904).
Trachydomia suwanneeae[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Trachyspira eleganta[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Triptychia catalanica[33]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Triptychia tomerdingensis[55]	Sp. nov	Valid	Kadolsky	Miocene (Aquitanian)		Germany	A member of the family Filholiidae.
Tritonophon grandis[23]	Sp. nov	In press	Ebbestad et al.	Late Ordovician	Lower Second Bani Formation	Morocco	A member of the family Bellerophontidae.
Trochomodulus stampinicus[40]	Sp. nov	Valid	Lozouet, Cahuzac & Charles	Oligocene (Rupelian)		France	A member of the family Modulidae.
Truncatellina dilatatus[56]	Sp. nov	Valid	Yu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Truncatellina.
Trypanocochlea lopburiensis[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Turbo histrioides[57]	Sp. nov	Valid	Kase et al.	Miocene		Japan	A species of Turbo.
Turbo izuensis[57]	Sp. nov	Valid	Kase et al.	Miocene		Japan	A species of Turbo.
Turritelloidea stepheni[41]	Sp. nov	Valid	Ferrari, Little & Atkinson	Early Jurassic (Toarcian)	Blea Wyke Sandstone Formation	United Kingdom	A member of the family Gordenellidae.
Umbilia darryli[58]	Sp. nov	Valid	Hawke	Miocene		Australia	A species of Umbilia.
Umbilia hallani[58]	Sp. nov	Valid	Hawke	Miocene		Australia	A species of Umbilia.
Vermetujanulus[52]	Gen. et sp. nov	Valid	Walther & Groh	Late Pleistocene		Madeira	A member of the family Gastrodontidae. Genus includes new species V. boessnecki.
Worthenia humiligrada[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Xenostoma[30]	Gen. et sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Genus includes new species X. lophopleura.
Yunnania inflata[29]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand

Bivalves

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Adelfia[59]	Gen. et comb. et sp. nov	Valid	Alvarez & Río	Eocene	La Meseta Formation Loreto Formation Submeseta Formation	Antarctica  Chile	A member of the family Veneridae. The type species is "Eumarcia" austrolissa Stilwell & Zinsmeister (1992); genus also includes "Venus" arenosa Ortmann (1899), as well as new species A. omega.
Aguileria romualdoensis[60]	Sp. nov	In press	Rodrigues et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil	A member of the family Bakevelliidae.
Araripenaia[61]	Gen. et sp. nov	In press	Da Silva et al.	Early Cretaceous	Crato Formation	Brazil	An unionid bivalve with possible hyriid affinities. Genus includes new species A. elliptica.
Araripevellia[60]	Gen. et sp. nov	In press	Rodrigues et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil	A member of the family Bakevelliidae. Genus includes new species A. musculosa.
Astarte danningeri[62]	Sp. nov	Valid	Mandic et al.	Early Miocene		Austria	A species of Astarte.
Cratonaia[63]	Gen. et sp. nov	Valid	Da Silva et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil	A bivalve belonging to the order Unionida and to the group Silesunionoidea. Genus includes new species C. novaolindensis.
Eothyasira[64]	Gen. nov	Valid	Karapunar et al.	Early Jurassic (Pliensbachian)		Germany
Geloina amithoscutana[53]	Sp. nov	Valid	Schneider, Kollmann & Pickford	Late Cretaceous	Al-Khodh Formation	Oman	A member of the family Cyrenidae.
Gyrostrea bogdanovae[65]	Sp. nov	Valid	Kosenko & Metelkin	Early Cretaceous (Hauterivian–Barremian)	Kugusem Formation	Kazakhstan	An oyster.
Illyricocongeria forcakovici[32]	Sp. nov	Valid	Mandic, Harzhauser & Neubauer	Middle Miocene		Bosnia and Herzegovina
Lima allerdingensis[62]	Sp. nov	Valid	Mandic et al.	Early Miocene		Austria	A species of Lima.
Limopsis quenstedti[64]	Nom. nov	Valid	Karapunar et al.				A replacement name for Nucula aurita Quenstedt (1856).
Lyropecten terrysmithae[66]	Sp. nov	Valid	Powell, Millard & Garcia	Miocene	Monterey Formation Santa Margarita Sandstone	United States ( California)	A scallop.
Malletia tsubetsuensis[67]	Sp. nov	Valid	Matsubara in Matsubara et al.	Miocene (Aquitanian)	Tsubetsu Formation	Japan	A member of the family Malletiidae
Marciachlys[59]	Gen. et comb. nov	Valid	Alvarez & Río	Eocene	Submeseta Formation	Antarctica	A member of the family Veneridae. The type species is "Eurhomalea" inflata Zinsmeister (1984).
Mezounio[68]	Gen. et sp. nov	Valid	Zakharov in Zakharov et al.	Middle Jurassic (Bajocian)		Russia	An unionid bivalve. Genus includes new species M. zverkovi.
Monginellopsis[61]	Gen. et sp. nov	In press	Da Silva et al.	Early Cretaceous	Crato Formation	Brazil	A bivalve with possible trigonioidid affinities. Genus includes new species M. bellaradiata.
Monopleura alesta[69]	Sp. nov	Valid	Masse & Fenerci-Masse	Early Cretaceous (Barremian)		France	A rudist bivalve.
Monopleura jurensis[69]	Sp. nov	Valid	Masse & Fenerci-Masse	Early Cretaceous (Hauterivian)		France	A rudist bivalve.
Muscatella[53]	Gen. et sp. nov	Valid	Schneider, Kollmann & Pickford	Late Cretaceous	Al-Khodh Formation	Oman	A member of the family Cyrenidae. The type species is M. biszczukae.
Myonera inouei[70]	Sp. nov	Valid	Amano in Amano & Kurita	Paleocene (Selandian)	Tomikawa Formation	Japan	A member of the family Cuspidariidae.
Myonia aparicioi[71]	Sp. nov	In press	Díaz Saravia & González	Carboniferous (Pennsylvanian)	Santa Elena Formation	Argentina
Nanogyra brevisulcata[72]	Sp. nov	Valid	Toscano & Lazo	Early Cretaceous	Neuquén Basin	Argentina	A bivalve belonging to the family Gryphaeidae.
Nicaniella schoberti[64]	Sp. nov	Valid	Karapunar et al.	Early Jurassic (Pliensbachian)		Germany
Oesterheldia[73]	Gen. et sp. nov	Valid	Pérez & Giachetti	Late Oligocene – early Miocene		Argentina	A member of the family Carditidae. The type species is "Venericardia" cannada Ihering (1907); genus also includes "Cyclocardia" dalek Pérez & del Río (2017).
Palaeomutela australis[74]	Sp. nov	In press	Guerrini et al.	Permian	Rio do Rasto Formation	Brazil	A bivalve belonging to the group Actinodontida and the family Palaeomutelidae.
Paracuspidaria[64]	Gen. nov	Valid	Karapunar et al.	Early Jurassic (Pliensbachian)		Germany
Permophorus? yrigoyeni[71]	Sp. nov	In press	Díaz Saravia & González	Carboniferous (Pennsylvanian)	Santa Elena Formation	Argentina
Praenucula pojetai[23]	Sp. nov	In press	Ebbestad et al.	Late Ordovician		Morocco	A bivalve belonging to the family Praenuculidae.
Pratulum kurrawchipellu[75]	Sp. nov	In press	Rojas & Nielsen	Early Miocene	Ipún Beds	Chile	A species of Pratulum.
Pratulum terpoorteni[75]	Sp. nov	In press	Rojas & Nielsen	Early Miocene	Navidad Formation	Chile	A species of Pratulum.
Promytilus rochacamposi[71]	Sp. nov	In press	Díaz Saravia & González	Carboniferous (Pennsylvanian)	Santa Elena Formation	Argentina
Protocardia broilii[76]	Sp. nov	Valid	Yanin	Early Cretaceous (Berriasian)		Crimean Peninsula
Relogiincola[74]	Gen. et sp. nov	In press	Guerrini et al.	Permian	Rio do Rasto Formation	Brazil	A bivalve belonging to the group Edmondioidea and the family Pachydomidae. Genus includes new species R. delicata.
Retroceramus alliensis[68]	Sp. nov	Valid	Zakharov in Zakharov et al.	Middle Jurassic (Bajocian)		Russia
Retroceramus (?) inopinatus[68]	Sp. nov	Valid	Zakharov in Zakharov et al.	Middle Jurassic (Bajocian)		Russia
Rollieria franconica[64]	Sp. nov	Valid	Karapunar et al.	Early Jurassic (Pliensbachian)		Germany
Rollieria goldfussi[64]	Nom. nov	Valid	Karapunar et al.				A replacement name for Nucula subovalis Goldfuss (1837).
Shajia[77]	Gen. et sp. nov	Valid	Rao et al.	Early Cretaceous (Aptian or Albian)	Langshan Formation	China  India	A rudist bivalve. Genus includes new species S. tibetica.
Terraia decarinata[74]	Sp. nov	In press	Guerrini et al.	Permian	Rio do Rasto Formation	Brazil	A bivalve belonging to the family Astartidae.
Trigonipraxis madjerensis[48]	Sp. nov	Valid	Neubauer et al.	Miocene		Serbia	A member of the family Dreissenidae belonging to the subfamily Congeriinae.

Other mollusks

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acanthochitona interglacialis[78]	Sp. nov	Valid	Dell'Angelo et al.	Late Pleistocene		Saudi Arabia	A chiton, a species of Acanthochitona.
Apoptopegma joyceae[79]	Sp. nov	Valid	Smith, Brock & Paterson	Cambrian (Guzhangian)	Shannon Formation	Australia	A rostroconch.
Lucilina aqabaensis[78]	Sp. nov	Valid	Dell'Angelo et al.	Late Pleistocene		Saudi Arabia	A chiton, a species of Lucilina.
Qaleruaqia[80]	Gen. et sp. nov	Valid	Peel	Cambrian Stage 4	Aftenstjernesø Formation	Greenland	A member of the family Mattheviidae. The type species is Q. sodermanorum.

Research

• New specimens of Pelagiella exigua with associated impressions of two clusters of chaetae, providing new information on the anatomy of this taxon, are described from the Cambrian Kinzers Formation (Pennsylvania, United States) by Thomas, Runnegar & Matt (2020), who infer the occurrence of a pair of chaeta-bearing appendages extending from the anterior body wall or foot of the animal (unique among living and extinct molluscs), and interpret Pelagiella as the earliest known stem gastropod.[81]

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