2020 in paleobotany

This article records new taxa of fossil plants that are scheduled to be described during the year 2020, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2020.

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New taxa

Flowering plants

Arecales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Echimonocolpites chicxulubensis[1]	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of a flowering plant, probably a member of the family Arecaceae.
Palmoxylon dindoriensis[2]	Sp. nov	Valid	Khan, Roy & Bera in Khan et al.	Late Cretaceous (Maastrichtian)-Paleocene (Danian)	Deccan Intertrappean Beds	India	A petrified palm stem.
Sabalites dawsonii[3]	Sp. nov	Valid	Greenwood & Conran	Eocene		Canada ( British Columbia)
Spinizonocolpites riochiquensis[4]	Sp. nov	Valid	Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal	Late Cretaceous (Maastrichtian)	Lago Colhué Huapí Formation	Argentina	A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020.

Alismatales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Limnobiophyllum pedunculatum[5]	Sp. nov	Valid	Low, Su & Xing in Low et al.	Late Oligocene		China	A member of the family Araceae.
Natantisphyllum[6]	Gen. et sp. nov	In press	Puebla, Vento & Prámparo	Late Cretaceous		Argentina	A member of the family Araceae. Genus includes new species N. crenae.

Buxales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Pachysandra europaea[7]	Sp. nov	In press	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Pachysandra.

Chloranthales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Sarcandraxylon[8]	Gen. et sp. nov	Valid	Pipo, Iglesias & Bodnar	Late Cretaceous (early–middle Campanian)	Santa Marta Formation	Antarctica	A member of the family Chloranthaceae. Genus includes new species S. sanjosense.

Cornales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Amersinia littletonensis[9]	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation	United States ( Colorado)
Langtonia parva[9]	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation	United States ( Colorado)	A member of the family Mastixiaceae.
Mastixicarpum hoodii[9]	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation	United States ( Colorado)	A member of the family Mastixiaceae.
Nyssa gergoei[10]	Sp. nov	Valid	Hably	Miocene		Hungary	A tupelo.
Nyssa gyoergyi[10]	Sp. nov	Valid	Hably	Miocene		Hungary	A tupelo.
Portnallia alexanderi[9]	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation	United States ( Colorado)	A member of the family Mastixiaceae.

Crossosomatales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Staphylea woodworthensis[11]	Sp. nov	In press	Zhu & Manchester	Oligocene	Renova Formation	United States ( Montana)	A species of Staphylea.

Cucurbitales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Coriaripites goodii[12]	Sp. nov	Valid	Barreda, Palazzesi & Tellería in Renner et al.	Late Cretaceous (Campanian–early Maastrichtian)	Lopez de Bertodano Formation Santa Marta Formation Snow Hill Island Formation	Antarctica	Pollen grains similar to those of extant members of the genus Coriaria.

Ericales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Sladenia zhengyii[13]	Sp. nov	In press	Jia & Zhou in Jia et al.	Early Miocene	Maguan Basin	China	A member of the family Sladeniaceae.

Fabales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Cercis zekuensis[14]	Sp. nov	In press	Li et al.	Early Miocene		China	A species of Cercis.
Gleditsia pliocaenica[7]	Sp. nov	In press	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Gleditsia.
Prioria martineziorum[15]	Sp. nov	Valid	Rodríguez-Reyes & Estrada-Ruiz	Oligocene-Miocene	Santiago Formation	Panama	A species of Prioria.
Striatopollis grahamii[1]	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of an eudicot, probably a member of the family Fabaceae.

Fagales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Alnus chaybulakensis[16]	Sp. nov	Valid	Averyanova & Xing	Paleogene		Kazakhstan	An alder.
Carpinus asymmetrica[17]	Sp. nov	In press	Xue & Jia in Xue et al.	Early Miocene	Maguan	China	A species of Carpinus.
Carpinus symmetrica[17]	Sp. nov	In press	Xue & Jia in Xue et al.	Early Miocene	Maguan Basin	China	A species of Carpinus
Castanopsis bulgarica[18]	Sp. nov	In press	Mantzouka, Ivanov & Bozukov	Late Miocene–early Pliocene (late Messinian–early Zanclean)	Pokrovnik	Bulgaria	A species of Castanopsis.
Quercus borissovii[16]	Sp. nov	Valid	Averyanova & Xing	Paleogene		Kazakhstan	An oak.

Garryales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Eucommia szaferi[7]	Sp. nov	In press	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Eucommia.

Gentianales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Asclepiadospermum[19]	Gen. et 2 sp. nov	Valid	Del Rio et al.	Early Eocene	Niubao Formation	China	A member of the family Apocynaceae belonging to the subfamily Asclepiadoideae. Genus includes new species A. marginatum and A. ellipticum.

Icacinales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Iodes passiciensis[20]	Sp. nov	Valid	Del Rio & De Franceschi	Early Eocene		France	A member of the family Icacinaceae.
Pyrenacantha simonsii[21]	Sp. nov	Valid	Stull et al.	Early Oligocene		Egypt	A species of Pyrenacantha.

Laurales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Mezilaurinoxylon oleiferum[22]	Sp. nov	Valid	Ruiz, Brea & Pujana in Ruiz et al.	Paleocene (Danian)	Salamanca Formation	Argentina	A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Patagonoxylon[22]	Gen. et sp. nov	Valid	Ruiz, Brea & Pujana in Ruiz et al.	Paleocene (Danian)	Salamanca Formation	Argentina	A member of Laurales of uncertain phylogenetic placement. Genus includes new species P. scalariforme. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Thymolepis[23]	Gen. et sp. nov	Valid	Chambers & Poinar	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Possibly an early representative of Monimiaceae. Genus includes new species T. toxandra.

Liliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Smilax fujianensis[24]	Sp. nov	In press	Dong et al.	Middle Miocene		China	A species of Smilax.
Smilax zhangpuensis[24]	Sp. nov	In press	Dong et al.	Middle Miocene		China	A species of Smilax.

Magnoliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Magnolia nanningensis[25]	Sp. nov	Valid	Huang et al.	Late Oligocene	Nanning Basin	China	A species of Magnolia.
Magnolia waltheri[7]	Sp. nov	In press	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Magnolia.

Malvales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Veraguasoxylon[15]	Gen. et sp. nov	Valid	Rodríguez-Reyes & Estrada-Ruiz	Oligocene-Miocene	Santiago Formation	Panama	A member of the family Malvaceae. Genus includes new species V. panamense.

Myrtales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Mangroveoxylon[26]	Gen. et comb. nov	In press	Moya & Brea	Late Miocene?	Ituzaingó Formation	Argentina	A member of the family Combretaceae; a new genus for "Menendoxylon" areniensis Lutz (1979).
Myrceugenellites grandiporosum[22]	Sp. nov	Valid	Ruiz, Brea & Pujana in Ruiz et al.	Paleocene (Danian)	Salamanca Formation	Argentina	A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Nymphaeales

Name	Novelty	Status	Authors	Type locality	Type locality	Location	Notes	Images
Praenymphaeapollenites[27]	Gen. et sp. nov	Valid	Barrón, Peris & Labandeira in Peris et al.	Late Cretaceous Cenomanian)	Burmese amber	Myanmar	Pollen of a member of Nymphaeaceae. Genus includes new species P. cenomaniensis.

Poales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Rhizomatites[28]	Gen. et sp. nov	In press	Robledo & Anzótegui in Robledo et al.	Miocene-Pliocene		Argentina	A member of Cyperaceae. Genus includes new species R. cyperoides.

Proteales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Banksia microphylla[29]	Sp. nov	Valid	Carpenter in Carpenter & Milne	Late Eocene		Australia	A species of Banksia.
Banksieaeidites zanthus[29]	Sp. nov	Valid	Milne in Carpenter & Milne	Late Eocene		Australia	A Banksia-like pollen.
Platanus emryi[30]	Sp. nov	Valid	Huegele, Spielbauer & Manchester	Miocene		United States	A species of Platanus.

Ranunculales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Clematis csabae[10]	Sp. nov	Valid	Hably	Miocene		Hungary	A species of Clematis.
Diploclisia praeaffinis[31]	Sp. nov	In press	Jia et al.	Late Miocene		China	A member of the family Menispermaceae.
Paleoorbicarpum[32]	Gen. et sp. nov	Valid	Han et al.	Paleocene	Sanshui Basin	China	A member of the family Menispermaceae. Genus includes new species P. parvum.
Stephania geniculata[32]	Sp. nov	Valid	Han et al.	Paleocene	Sanshui Basin	China	A species of Stephania
Stephania ornamenta[32]	Sp. nov	Valid	Han et al.	Paleocene	Sanshui Basin	China	A species of Stephania

Rosales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Berhamniphyllum junrongii[33]	Sp. nov	Valid	Zhou, Wang & Huang in Zhou et al.	Late Eocene	Markam Basin	China	A member of the family Rhamnaceae
Crataegus pentagynoides[7]	Sp. nov	In press	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Crataegus
Hemiptelea kryshtofovichii[16]	Sp. nov	Valid	Averyanova & Xing	Paleogene		Kazakhstan	A member of the family Ulmaceae.
Scabrastephanoporites[1]	Gen. et sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of an eudicot, probably a member of the family Ulmaceae or Cannabaceae. Genus includes new species S. variabilis.

Sapindales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Acer dombeyopsis[7]	Sp. nov	In press	Kvaček, Teodoridis & Denk	Pliocene		Germany	A maple.
Acer viburnoides[7]	Sp. nov	In press	Kvaček, Teodoridis & Denk	Pliocene		Germany	A maple.
Acer vitiforme[7]	Sp. nov	In press	Kvaček, Teodoridis & Denk	Pliocene		Germany	A maple.
Brosipollis reticulatus[1]	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of a flowering plant, probably a member of the family Burseraceae.
Llanodelacruzoxylon[34]	Gen. et sp. nov		Rodríguez-Reyes, Estrada-Ruiz & Gasson	Oligocene–Miocene	Santiago Formation	Panama	A member of the family Anacardiaceae. Genus includes new species L. sandovalii.
Manchestercarpa[35]	Gen. et sp. nov	Valid	Atkinson	Late Cretaceous (Campanian)		Canada ( British Columbia)	A member of the family Meliaceae described on the basis of a fossil fruit. Genus includes new species M. vancouverensis.
Parametopioxylon[36]	Gen. et sp. nov	Valid	Franco et al.	Miocene	Ituzaingó Formation	Argentina	A member of the family Anacardiaceae described on the basis of fossil wood. Genus includes new species P. crystalliferum.
Psilastephanocolporites hammenii[1]	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of a flowering plant, probably a member of the family Meliaceae

Saxifragales

Name	Novelty	Status	Authors	Type locality	Type locality	Location	Notes	Images
Corylopsis grisea[37]	Sp. nov	In press	Quirk & Hermsen	Early Pliocene	Gray Fossil Site	United States ( Tennessee)	A species of Corylopsis.
Protoaltingia[38]	Gen. et sp. nov	Valid	Scharfstein, Stockey & Rothwell	Late Cretaceous (Coniacian)		Canada ( British Columbia)	A member of the family Altingiaceae. Genus includes new species P. comoxense.

Trochodendrales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Eotrochion[39]	Gen. et sp. nov	In press	Manchester, Kvaček & Judd	Paleocene		United States ( Wyoming)	A member of the family Trochodendraceae. Genus includes new species E. polystylum.
Paraconcavistylon[39]	Gen. et comb. nov	In press	Manchester, Kvaček & Judd	Early Eocene		Canada ( British Columbia)  United States ( Washington)	A member of the family Trochodendraceae; a new genus for "Concavistylon" wehrii Manchester et al. (2018)
Trochodendron infernense[39]	Sp. nov	In press	Manchester, Kvaček & Judd	Paleocene		United States ( Wyoming)	A species of Trochodendron.

Other angiosperms

Name	Novelty	Status	Authors	Type locality	Location	Notes	Images
Aextoxicoxylon kawasianus[40]	Sp. nov	Valid	Vera et al.	Late Cretaceous	Puntudo Chico Formation	Argentina	A fossil dicot wood
Carpolithes gergoei[10]	Sp. nov	Valid	Hably & Erdei in Hably	Miocene		Hungary	A fossil fruit of a flowering plant of uncertain phylogenetic placement.
Cavilignum[41]	Gen. et sp. nov	Valid	Siegert & Hermsen	Early Pliocene	Gray Fossil Site	United States ( Tennessee)	A flowering plant of uncertain phylogenetic placement, described on the basis of fossil endocarps. Genus includes new species C. pratchettii.
Dasykothon[42]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species D. leptomiscus.
Phantophlebia[43]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A flowering plant of uncertain phylogenetic placement, possibly related to myrsinoid members of the family Primulaceae. Genus includes new species P. dicycla.
Rasenganus[44]	Gen. et sp. nov	Valid	Xing & Gu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A possible epizoochorous fruit. Genus includes new species R. auricularus.
Sinoherba[45]	Gen. et sp. nov	In press	Liu & Wang in Liu, Chen & Wang	Early Cretaceous (Barremian–Aptian)	Yixian Formation	China	An early monocot. Genus includes new species S. ningchengensis.

Pinales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Agathis ledongensis[46]	Sp. nov	Valid	Oskolski et al.	Late Oligocene–early Miocene	Qiutangling Formation	China	A species of Agathis
Agathoxylon cozzoi[47]	Sp. nov	Valid	Gnaedinger & Zavattieri	Late Triassic	Chihuido Formation	Argentina
Araucaria famii[48]	Sp. nov	Valid	Stockey & Rothwell	Late Cretaceous (Campanian)		Canada ( British Columbia)	A species of Araucaria
Araucaria fildesensis[49]	Sp. nov	Valid	Shi et al.	Eocene	Fossil Hill Formation	Antarctica (King George Island)	A species of Araucaria
Araucaria huncoensis[50]	Sp. nov	Valid	Rossetto‐Harris in Rossetto‐Harris et al.	Early Eocene	Laguna del Hunco Formation	Argentina	A species of Araucaria.
Araucarites pachacuteci[51]	Sp. nov	In press	Martínez in Martínez et al.	Early Cretaceous (Berriasian–Valanginian)	Huancané Formation	Peru	A member of the family Araucariaceae
Brachyoxylon zhouii[52]	Sp. nov	In press	Jiang et al.	Early Cretaceous	Guantou Formation	China	A conifer wood.
Brachyphyllum sattlerae[53]	Sp. nov	In press	Batista et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Pinidae.
Circoporoxylon tibetense[54]	Sp. nov	In press	Xia et al.	Middle Jurassic (Callovian)	Xiali Formation	China	Possibly a member of the family Podocarpaceae.
Comoxostrobus[55]	Gen. et sp. nov	Valid	Stockey, Rothwell & Atkinson	Late Cretaceous (early Coniacian)		Canada ( British Columbia)	A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species C. rossii.
Cupressinoxylon llantenesense[47]	Sp. nov	Valid	Gnaedinger & Zavattieri	Late Triassic	Llantenes Formation	Argentina
Cupressinoxylon manuelii[56]	Sp. nov	In press	Ríos-Santos, Cevallos-Ferriz & Pujana	Late Cretaceous (Campanian-Maastrichtian)	Cabullona Group	Mexico
Dacrycarpus guipingensis[57]	Sp. nov	In press	Wu et al.	Miocene	Erzitang Formation	China	A species of Dacrycarpus
Friisia[58]	Gen. et sp. nov	In press	Mendes & Kvaček	Early Cretaceous (late Aptian – early Albian)	Lusitanian Basin	Portugal	An ovuliferous cone of a member of the family Podocarpaceae. Genus includes new species F. lusitanica.
Marskea cuspidata[59]	Sp. nov	Valid	Frolov & Mashchuk	Middle Jurassic	Prisayan Formation	Russia	A member of the family Taxaceae.
Mukawastrobus[60]	Gen. et sp. nov	In press	Stockey, Nishida & Rothwell	Late Cretaceous (late Campanian—early Maastrichtian)		Japan	A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species M. satoi.
Pinuxylon selmeierianum[61]	Sp. nov	Valid	Dolezych & Reinhardt	Paleogene	Eureka Sound Group	Canada ( Nunavut)	A member of the family Pinaceae described on the basis of fossil wood
Podocarpus pliomacrophyllus[62]	Sp. nov	In press	Chen et al.	Early Pliocene		China	A species of Podocarpus
Tsuga asiatica[63]	Sp. nov	In press	Wu & Zhou in Wu et al.	Late Paleogene		China	A species of Tsuga

Other seed plants

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Amyelon turpanense[64]	Sp. nov	In press	Shi, Yu & Sun	Permian (Lopingian)	Wutonggou	China	A root of a member of Cordaitales
Androstrobus obovatus[65]	Sp. nov	In press	Bodnar et al.	Late Triassic	Potrerillos Formation	Argentina	A member of Cycadales.
Battenispermum[66]	Gen. et sp. nov	In press	Mendes, Pedersen & Friis	Early Cretaceous		Portugal	A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species B. hirsutum.
Carpolithus volantus[67]	Sp. nov	Valid	Gómez et al.	Early Cretaceous (Aptian)	La Cantera Formation	Argentina	A fossil seed, possibly produced by a member of Gnetales.
Ductolobatopitys[68]	Gen. et sp. nov	Valid	Conceição & Crisafulli in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm described on the basis of fossil wood. Genus includes new species D. mussae.
Jianchangia[69]	Gen. et sp. nov	Valid	Yang, Wang & Ferguson	Early Cretaceous (Aptian)	Jiufotang Formation	China	A member of Ephedraceae. Genus includes new species J. verticillata.
Johniphyllum[70]	Gen. et sp. nov	Valid	Looy & Duijnstee	Permian (Guadalupian)		United States ( Texas)	A member of Voltziales. Genus includes new species J. multinerve.
Jordaniopteris[71]	Gen. et comb. nov	Valid	Anderson in Anderson et al.	Permian (possibly Lopingian)	Um Irna Formation	Jordan	A seed fern. A new genus for "Dicroidium" irnensis Abu Hamad et al. (2008); genus also includes "Dicroidium" jordanensis Abu Hamad et al. (2008), "Dicroidium" robustum Kerp & Vörding (2008) and "Dicroidium" bandelii Abu Hamad et al. (2017).
Kaokoxylon brasiliensis[68]	Sp. nov	Valid	Conceição, Neregato & Iannuzzi in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A conifer described on the basis of fossil wood.
Mariopteris hexiensis[72]	Sp. nov	In press	Wang et al.	Permian (Cisuralian)	Shanxi Formation	China
Mariopteris yongchangensis[72]	Sp. nov	In press	Wang et al.	Permian (Cisuralian)	Shanxi Formation	China
Novaiorquepitys[73]	Gen. et sp. nov	In press	Conceição & Crisafulli in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm stem. Genus includes new species N. maranhensis Conceição, Neregato & Iannuzzi.
Palaeocupressinoxylon[74]	Gen. et sp. nov	In press	Wan, Yang & Wang	Late Permian	Turpan–Hami Basin	China	A silicified gymnospermous fossil wood. Genus includes new species P. uniseriale.
Pseudovoltzia sapflorensis[70]	Sp. nov	Valid	Looy & Duijnstee	Permian (Guadalupian)		United States ( Texas)	A member of Voltziales
Pterostoma neehoffii[75]	Sp. nov	Valid	Conran et al.	Middle Miocene		New Zealand	A cycad
Pteruchus frenguellii[76]	Sp. nov	In press	Blomenkemper et al.	Late Permian	Umm Irna Formation	Jordan	A pollen organ of a seed fern
Pteruchus lepidus[76]	Sp. nov	In press	Blomenkemper et al.	Late Permian	Umm Irna Formation	Jordan	A pollen organ of a seed fern
Samaropsis jinchangensis[77]	Sp. nov	In press	Hua & Sun in Hua et al.	Early Permian		China	A seed fossil
Umaltolepis involuta[78]	Sp. nov	In press	Nosova	Middle Jurassic	Angren Formation	Uzbekistan
Umaltolepis sogdianica[78]	Sp. nov	In press	Nosova	Middle Jurassic	Angren Formation	Uzbekistan
Umkomasia aequatorialis[76]	Sp. nov	In press	Blomenkemper et al.	Late Permian	Umm Irna Formation	Jordan	A cupulate structure of a seed fern
Wantus[70]	Gen. et sp. nov	Valid	Looy & Duijnstee	Permian (Guadalupian)		United States ( Texas)	A member of Voltziales. Genus includes new species W. acaulis.
Wudaeophyton[79]	Gen. et sp. nov	Valid	Pšenička et al.	Early Permian	Taiyuan Formation	China	A small vine, most similar to pteridosperms from the group Callistophytales. Genus includes new species W. wangii.
Yangopteris[80]	Gen. et comb. nov	In press	Zhou et al.	Permian (Asselian)		China	A seed fern; a new genus for "Alethopteris" ascendens Halle.
Yvyrapitys[73]	Gen. et sp. nov	In press	Conceição & Crisafulli in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm stem. Genus includes new species Y. novaiorquensis Conceição, Neregato & Iannuzzi.

Other plants

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Annularia noronhai[81]	Sp. nov	In press	Correia et al.	Carboniferous (Gzhelian)	Douro Basin	Portugal	A member of the family Calamitaceae
Annularia paisii[82]	Sp. nov	Valid	Correia et al.	Carboniferous (Gzhelian)		Portugal
Botryopteris multifolia[83]	Sp. nov	In press	He et al.	Permian (Lopingian)	Junlian Formation	China	A fern
Calamites cambrensis[84]	Sp. nov	Valid	Thomas	Carboniferous (Pennsylvanian)		United Kingdom
Chlamydomonas hanublikanus[85]	Sp. nov		Vršanská & Hinkelman	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Chlamydomonas
Circinites[28]	Gen. et sp. nov	In press	Robledo & Anzótegui in Robledo et al.	Miocene-Pliocene		Argentina	A fern belonging to the family Pteridaceae. Genus includes new species C. pteridoides.
Clavatisporites cenomaniana[86]	Sp. nov	Valid	Santamarina in Santamarina et al.	Late Cretaceous (Cenomanian)	Mata Amarilla Formation	Argentina	Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Collarecodium? nezpercae[87]	Sp. nov	Valid	Bucur & Rigaud in Bucur et al.	Late Triassic (Norian)		United States ( Idaho)	A green alga belonging to the group Bryopsidales and possibly to the family Udoteaceae.
Collarisporites minor[86]	Sp. nov	Valid	Santamarina in Santamarina et al.	Late Cretaceous (Cenomanian)	Mata Amarilla Formation	Argentina	Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Coniopteris sandaolingensis[88]	Sp. nov	Valid	Yuan & Sun in Yuan et al.	Middle Jurassic	Xishanyao Formation	China
Drynaria diplosticha[89]	Sp. nov	In press	Yu & Xie in Yu et al.	Late Miocene	Bangmai Formation	China	A fern belonging to the family Polypodiaceae.
Electrophycus[90]	Gen. et sp. nov	In press	Poinar & Brown	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A green alga, possibly a member of the family Chaetophoraceae. Genus includes new species E. astroplethus.
Equicalastrobus pusillus[91]	Sp. nov	In press	Zhang & Yan in Zhang et al.	Late Triassic	Baojishan Basin	China	A member of Equisetales
Equisetum yenbaiense[92]	Sp. nov	Valid	Aung et al.	Late Miocene		Vietnam	A species of Equisetum
Equisetum yongpingense[92]	Sp. nov	Valid	Aung et al.	Late Pliocene	Sanying Formation	Vietnam	A species of Equisetum
Frullania partita[93]	Sp. nov	Valid	Li et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A liverwort, a species of Frullania
Frullania vanae[94]	Sp. nov	Valid	Mamontov et al.	Eocene	Rovno amber	Ukraine	A liverwort, a species of Frullania
Gippslandites[95]	Gen. et sp. nov	In press	McSweeney, Shimeta & Buckeridge	Late Silurian–early Devonian		Australia	A member of Zosterophyllaceae. Genus includes new species G. minutus.
Gmujij[96]	Gen. et sp. nov	In press	Pfeiler & Tomescu	Devonian (Emsian)	Battery Point	Canada ( Quebec)	An early euphyllophyte. Genus includes new species G. tetraxylopteroides.
Griphoporella minuta[87]	Sp. nov	Valid	Bucur & Peybernes in Bucur et al.	Late Triassic		Japan	A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Holosporella magna[87]	Sp. nov	Valid	Bucur & Fucelli in Bucur et al.	Late Triassic (Norian)		United States ( Nevada)	A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Holosporella? rossanae[87]	Sp. nov	Valid	Bucur & Del Piero in Bucur et al.	Late Triassic (Norian)		Canada ( Yukon)	A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Jurafructus[97]	Gen. et sp. nov	Valid	Chen et al.	Middle−Late Jurassic	Jiulongshan Formation	China	A plant of uncertain phylogenetic placement, possibly a flowering plant described on the basis of a probable fossil drupe. Genus includes new species J. daohugouensis.
Keraphyton[98]	Gen. et sp. nov	Valid	Champreux, Meyer-Berthaud & Decombeix	Devonian (Famennian)	Mandowa Mudstone Formation	Australia	A member of Iridopteridales of uncertain phylogenetic placement. Genus includes new species K. mawsoniae.
Lygodium sanshuiense[99]	Sp. nov	In press	Naugolnykh et al.	Paleocene	Buxin Formation	China	A species of Lygodium
Maiaspora[100]	Gen. et sp. nov	In press	Mamontov et al.	Carboniferous (Viséan)	Moscow Syneclise	Russia	A miospore. Genus includes new species M. panopta.
Neoarthropitys[101]	Gen. et sp. nov	In press	Gnaedinger et al.	Middle Triassic	Quebrada de los Fósiles	Argentina	A member of Equisetales. Genus includes new species N. gondwanaensis.
Osmundacaulis asiatica[102]	Sp. nov	Valid	Cheng et al.	Cretaceous		China	A member of the family Osmundaceae
Osmundacaulis sinica[102]	Sp. nov	Valid	Cheng et al.	Cretaceous		China	A member of the family Osmundaceae
Ovoidites circumplicatus[103]	Sp. nov	In press	Zavattieri, Gutiérrez & Monti	Middle Triassic	Quebrada de los Fósiles Formation	Argentina	A green alga belonging to the group Zygnematales.
Ovoidites tripartitus[103]	Sp. nov	In press	Zavattieri, Gutiérrez & Monti	Middle Triassic	Quebrada de los Fósiles Formation	Argentina	A green alga belonging to the group Zygnematales.
Palaeostachya guanglongii[104]	Sp. nov	In press	Liu et al.	Permian (Asselian)	Taiyuan Formation	China	A member of the family Calamitaceae.
Parazosterophyllum[95]	Gen. et sp. nov	In press	McSweeney, Shimeta & Buckeridge	Late Silurian–early Devonian		Australia	A member of Zosterophyllaceae. Genus includes new species P. timsiae.
Patruliuspora oregonica[87]	Sp. nov	Valid	Bucur & Rigaud in Bucur et al.	Late Triassic (Norian)		United States ( Oregon)	A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Patruliuspora pacifica[87]	Sp. nov	Valid	Bucur, Del Piero & Peyrotty in Bucur et al.	Late Triassic (Norian)		Canada ( Yukon)	A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Pellites hamiensis[105]	Sp. nov	In press	Li et al.	Middle Jurassic	Xishanyao	China	A liverwort belonging to the family Pelliaceae.
Plenasium (Aurealcaulis) elegans[106]	Sp. nov	In press	Hiller et al.	Eocene	Na Duong Formation	Vietnam	A member of Osmundaceae
Plenasium xiei[107]	Sp. nov	In press	Cheng et al.	Cretaceous		China	A member of Osmundaceae
Pleuromeia shaolinii[108]	Sp. nov	In press	Zhang & Wang in Zhang et al.	Middle Triassic	Linjia	China
Polycingulatisporites multiverrucata[86]	Sp. nov	Valid	Santamarina in Santamarina et al.	Late Cretaceous (Cenomanian)	Mata Amarilla	Argentina	Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Polysporia baetica[109]	Sp. nov	In press	Álvarez-Vázquez, Bek & Drábková	Carboniferous (Pennsylvanian)	Peñarroya-Belmez-Espiel Coalfield	Spain	A member of Isoetales
Polystichum pacltovae[110]	Sp. nov	Valid	Kvaček in Kvaček & Teodoridis	Oligocene		Czech Republic	A fern, a species of Polystichum
Proodontosoria[111]	Gen. et sp. nov	Valid	Li et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A fern belonging to the family Lindsaeaceae. Genus includes new species P. myanmarensis.
Proterocladus antiquus[112]	Sp. nov	Valid	Tang et al.	Mesoproterozoic circa 1 Ga	Nanfen	China	An early siphonocladalean chlorophyte
Psilochara monevaensis[113]	Sp. nov	Valid	Sanjuan & Soulié-Märsche	Middle Miocene		Spain	A charophyte.
Qianshouia[114]	Gen. et sp. nov	In press	Huang et al.	Late Devonian	Wutong Formation	China	A plant of uncertain phylogenetic placement, possibly a lycopsid or a sphenopsid. Genus includes new species Q. mira.
Scolecopteris minuta[115]	Sp. nov	In press	Wan et al.	Early Permian	Taiyuan Formation	China	A fern belonging to the group Marattiales.
Sigillaria pfefferkornii[116]	Sp. nov	In press	D'Antonio, Boyce & Wang	Permian (Asselian)		China
Sigillaria wudensis[116]	Sp. nov	In press	D'Antonio, Boyce & Wang	Permian (Asselian)		China
Sphaerochara miocenica[113]	Sp. nov	Valid	Sanjuan & Soulié-Märsche	Miocene		Lebanon  Spain	A charophyte.
Thyrsopteris cretacea[117]	Sp. nov	Valid	Li et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Thyrsopteris
Tomiostrobus sinensis[118]	Sp. nov	Valid	Feng in Feng et al.	Early Triassic (Induan)	Kayitou Formation	China	A member of the family Isoetaceae.
Triskelia[119]	Gen. et sp. nov	In press	Strullu‐Derrien et al.	Devonian	Rhynie Chert	United Kingdom	A green alga. Genus includes new species T. scotlandica.
Ufadendron elongatum[120]	Sp. nov	Valid	Tang et al.	Late Permian	Linxi	China	A lycopsid belonging to the family Tomiodendraceae
Zeilleria fosteri[121]	Sp. nov	Valid	Thomas et al.	Carboniferous (Bashkirian)		United Kingdom	A fern

General research

• A study on the evolutionary history of green plants is published by Nie et al. (2020).[122]
• Description of new fossil material of Yurtusia uniformis from the Cambrian Yanjiahe Formation (China) and a study on the phylogenetic relationships and possible life cycle of this organism is published by Shang et al. (2020), who consider Y. uniformis to be a likely green microalga.[123]
• Evidence of development of dichotomous roots in euphyllophytes that were extant during the Devonian and Carboniferous periods is presented by Hetherington, Berry & Dolan (2020), who interpret their findings as indicating that dichotomous root branching evolved in both lycophytes and euphyllophytes.[124]
• An early land plant producing multiple spore size classes is described from the Lower Devonian Campbellton Formation (Canada) by Bonacorsi et al. (2020).[125]
• A study on the impact of the appearance and evolution of herbivorous tetrapods on the evolution of land plants from the Carboniferous to the Early Triassic is published by Brocklehurst, Kammerer & Benson (2020).[126]
• A study on the production of periderm in Late Paleozoic arborescent lycopsids is published by D'Antonio & Boyce (2020), who argue that these lycopsids did not grow from sporelings into large trees through the production of a periderm cylinder, because physiological limitations would have prohibited the production of thick periderm.[127]
• A study on the architecture and development of the Carboniferous arborescent lycopsid Paralycopodites is published by DiMichele & Bateman (2020).[128]
• A study on the phylogenetic placement of the extinct fern genus Coniopteris is published by Li et al. (2020).[129]
• New information on the morphology of Paleoazolla patagonica is presented by Benedetti et al. (2020), who evaluate the implications of this taxon for the knowledge of the evolution of water ferns.[130]
• A study on the morphology and development of Genomosperma, and on its implications for the knowledge of the evolutionary origins of seed development, is published by Meade, Plackett & Hilton (2020).[131]
• A study on the anatomy of the seed cone scales of Krassilovia mongolica is published by Herrera et al. (2020), who argue that K. mongolica and Podozamites harrisii are the seed cones and leaves of the same extinct plant, and name a new family Krassiloviaceae within the order Voltziales.[132]
• A putative bamboo "Chusquea" oxyphylla from the early Eocene Laguna del Hunco biota (Argentina) is reinterpreted as a conifer by Wilf (2020), who transfers this species to the genus Retrophyllum.[133]
• Presence of secretory tissues is reported in extinct flowers from the Cretaceous amber from Myanmar and Cenozoic Dominican amber (including specimens preserved while in the process of emitting compounds) by Poinar & Poinar (2020).[134]
• Fossil pollen of flowering plants is reported from the Aptian and Albian of Australia by Korasidis & Wagstaff (2020), who evaluate the implications of their findings for the knowledge of the timing of the appearance and diversification of the flowering plants in the high-latitude southern basins of Australia.[135]
• A study on the morphology of palm and palm-like pollen from the Eocene Yaw Formation (Myanmar), and on the implications of these fossils for the knowledge of distribution and diversity of Eocene palms across the globe, is published by Huang et al. (2020).[136]
• A study on the extinction of plants from south polar terrestrial ecosystems during the Permian–Triassic extinction event and on their recovery after this extinction event, based on data from the Sydney Basin (Australia), is published by Mays et al. (2020).[137]
• A study on the age of the Paleogene Kanaka Creek fossil flora (Huntingdon Formation; British Columbia, Canada) and on its implications for reconstructions of the contemporaneous paleoclimate and paleoenvironment will be published by Mathewes, Greenwood & Love (2020).[138]
• Fossil fruits (mericarps) of the neoendemic Apiaceae Melanoselinum (≡ Daucus) decipiens were reported from the lacustrine and fluvial sediments of Porto da Cruz, Madeira, dated 1.3 Ma, by Góis-Marques et al. 2020[139]. This paper not only reports the oldest Daucus s.l. fossil known to date but also the first fossil evidence of a plant with insular woodiness (see Island gigantism).
• The leaf fossil Mesodescolea plicata from the Early Cretaceous of Patagonia, first interpreted as a cycad with affinities with extant Stangeria, is reinterpreted as an angiosperm leaf with affinities with Austrobaileyales or Chloranthales by Coiro et al. 2020[140], with implications for the evolution of leaf shape in the early radiation of the angiosperms.

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47. Silvia C. Gnaedinger; Ana María Zavattieri (2020). "Coniferous woods from the Upper Triassic of southwestern Gondwana, Tronquimalal Group, Neuquén Basin, Mendoza Province, Argentina". Journal of Paleontology. 94 (3): 387–416. doi:10.1017/jpa.2020.1.
48. Ruth A. Stockey; Gar W. Rothwell (2020). "Diversification of crown group Araucaria: the role of Araucaria famii sp. nov. in the mid‐Cretaceous (Campanian) radiation of Araucariaceae in the Northern Hemisphere". American Journal of Botany. 107 (7): 1072–1093. doi:10.1002/ajb2.1505. PMID 32705687.
49. Gongle Shi; Haomin Li; Andrew B. Leslie; Zhiyan Zhou (2020). "Araucaria bract-scale complex and associated foliage from the early-middle Eocene of Antarctica and their implications for Gondwanan biogeography". Historical Biology: An International Journal of Paleobiology. 32 (2): 164–173. doi:10.1080/08912963.2018.1472255.
50. Gabriella Rossetto‐Harris; Peter Wilf; Ignacio H. Escapa; Ana Andruchow‐Colombo (2020). "Eocene Araucaria Sect. Eutacta from Patagonia and floristic turnover during the initial isolation of South America". American Journal of Botany. 107 (5): 806–832. doi:10.1002/ajb2.1467. PMID 32388874.
51. L.C.A. Martínez; E. Pacheco Huacallo; R.R. Pujana; H. Padula (2020). "A new megaflora (leaves and reproductive structures) from the Huancané Formation (Lower Cretaceous), Peru". Cretaceous Research. 110: Article number 104426. doi:10.1016/j.cretres.2020.104426.
52. Zikun Jiang; Hao Wu; Ning Tian; Yongdong Wang; Aowei Xie (2020). "A new species of conifer wood Brachyoxylon from the Cretaceous of Eastern China and its paleoclimate significance". Historical Biology: An International Journal of Paleobiology. in press. doi:10.1080/08912963.2020.1755282.
53. Maria Edenilce P. Batista; Lutz Kunzman; Artur A. A. Sá; Antônio Álamo F. Saraiva; Maria Iracema B. Loiola (2020). "A new species of Brachyphyllum from the Crato Formation (Lower Cretaceous), Araripe Basin, Brazil". Ameghiniana. in press. doi:10.5710/AMGH.23.06.2020.3333.
54. Guoqing Xia; Ning Tian; Marc Philippe; Haisheng Yi; Chihua Wu; Gaojie Li; Zhiqiang Shi (2020). "Oldest Jurassic wood with Gondwanan affinities from the Middle Jurassic of Tibetan Plateau and its paleoclimatological and paleoecological significance". Review of Palaeobotany and Palynology. 281: Article 104283. doi:10.1016/j.revpalbo.2020.104283.
55. Ruth A. Stockey; Gar W. Rothwell; Brian A. Atkinson (2020). "Late Cretaceous diversification of cupressaceous conifers: a taiwanioid seed cone from the Eden Main, Vancouver Island, British Columbia, Canada". International Journal of Plant Sciences. 181 (5): 529–541. doi:10.1086/708383.
56. César Ríos-Santos; Sergio R.S. Cevallos-Ferriz; R.R. Pujana (2020). "Cupressaceous woods in the Upper Cretaceous Cabullona Group in Fronteras, Sonora, Mexico". Journal of South American Earth Sciences. in press: Article 102756. doi:10.1016/j.jsames.2020.102756.
57. Xin‐Kai Wu; Natalia E. Zavialova; Tatiana M. Kodrul; Xiao‐Yan Liu; Natalia V. Gordenko; Natalia P. Maslova; Cheng Quan; Jian‐Hua Jin (2020). "Northern Hemisphere megafossil of Dacrycarpus (Podocarpaceae) from Miocene of South China and its evolutionary and palaeoecological implication". Journal of Systematics and Evolution. in press. doi:10.1111/jse.12534.
58. Mário Miguel Mendes; Jiří Kvaček (2020). "Friisia lusitanica gen. et sp. nov., a new podocarpaceous ovuliferous cone from the Lower Cretaceous of Lusitanian Basin, western Portugal". Cretaceous Research. 108: Article 104352. doi:10.1016/j.cretres.2019.104352.
59. Andrey O. Frolov; Irina M. Mashchuk (2020). "Discovery of isolated leaves of Marskea (Taxaceae) in the Middle Jurassic sediments of Irkutsk Basin (East Siberia, Russia)". Phytotaxa. 449 (2): 164–172. doi:10.11646/phytotaxa.449.2.4.
60. Ruth A. Stockey; Harufumi Nishida; Gar W. Rothwell (2020). "Evolutionary diversification of taiwanioid conifers: evidence from a new Upper Cretaceous seed cone from Hokkaido, Japan". Journal of Plant Research. in press. doi:10.1007/s10265-020-01214-y. PMID 32686035.
61. Martina Dolezych; Lutz Reinhardt (2020). "First evidence for the conifer Pinus, as Pinuxylon selmeierianum sp. nov., during the Paleogene on Wootton Peninsula, northern Ellesmere Island, Nunavut, Canada". Canadian Journal of Earth Sciences. 57 (1): 25–39. Bibcode:2020CaJES..57...25D. doi:10.1139/cjes-2018-0163.
62. Hui Chen; De-Liang Tang; Yu Zhang; Peng-Cheng An; Xin-Yu Yan; Su-Ting Ding; Jing-Yu Wu (2020). "Fossil Podocarpus (Podocarpaceae) from the lower Pliocene of Tengchong, Yunnan Province, China and its biogeographic significance". Historical Biology: An International Journal of Paleobiology. in press: 1–10. doi:10.1080/08912963.2019.1697254.
63. Meng-Xiao Wu; Jian Huang; Tao Su; Qin Leng; Zhe-Kun Zhou (2020). "Tsuga seed cones from the late Paleogene of southwestern China and their biogeographical and paleoenvironmental implications". Palaeoworld. in press. doi:10.1016/j.palwor.2019.07.005.
64. Xiao Shi; Jianxin Yu; Yuewu Sun (2020). "Tyloses in the Lopingian cordaitalean root from Xinjiang, Northwest China". Review of Palaeobotany and Palynology. 273: Article 104134. doi:10.1016/j.revpalbo.2019.104134.
65. Josefina Bodnar; Eduardo M. Morel; Eliana P. Coturel; Daniel G. Ganuza (2020). "New plant fossil records and biostratigraphic analysis from the Uspallata Group (Late Triassic) at Cacheuta Hill, Cuyo Basin, west-central Argentina". Geobios. in press. doi:10.1016/j.geobios.2020.04.002.
66. Mário Miguel Mendes; Kaj Raunsgaard Pedersen; Else Marie Friis (2020). "Battenispermum hirsutum gen. et sp. nov., a new Early Cretaceous seed from Portugal with chlamydospermous organisation". Cretaceous Research. 109: Article 104376. doi:10.1016/j.cretres.2020.104376.
67. María A. Gómez; Gabriela G. Puebla; Mercedes B. Prámparo; Andrea B. Arcucci (2020). "Fossil seeds from the La Cantera Formation, Early Cretaceous, San Luis Province, Argentina". Acta Palaeobotanica. 60 (1): 181–198. doi:10.35535/acpa-2020-0008.
68. Domingas Maria da Conceição; Luiz Saturnino de Andrade; Rodrigo Neregato; Roberto Iannuzzi; Alexandra Crisafulli; Juan Carlos Cisneros (2020). "New petrified gymnosperms from the Permian of Maranhão (Pedra de Fogo Formation), Brazil: Ductolobatopitys nov. gen. and Kaokoxylon". Geobios. 60: 47–59. doi:10.1016/j.geobios.2020.04.003.
69. Yong Yang; Yingwei Wang; David Kay Ferguson (2020). "A new macrofossil ephedroid plant with unusual bract morphology from the Lower Cretaceous Jiufotang Formation of northeastern China". BMC Evolutionary Biology. 20 (1): Article number 19. doi:10.1186/s12862-019-1569-y. PMC 7001366. PMID 32019502.
70. Cindy V. Looy; Ivo A. P. Duijnstee (2020). "Voltzian conifers of the South Ash Pasture flora (Guadalupian, Texas): Johniphyllum multinerve gen. et sp. nov., Pseudovoltzia sapflorensis sp. nov., and Wantus acaulis gen. et sp. nov". International Journal of Plant Sciences. 181 (3): 363–385. doi:10.1086/706853.
71. Heidi M. Anderson; Maria Barbacka; Marion K. Bamford; W. B. Keith Holmes; John M. Anderson (2020). "Dicroidium (foliage) and affiliated wood Part 3 of a reassessment of Gondwana Triassic plant genera and a reclassification of some previously attributed". Alcheringa: An Australasian Journal of Palaeontology. 44 (1): 64–92. doi:10.1080/03115518.2019.1622779.
72. Xue-Lian Wang; Yan-Zhao Ji; Yi-Fan Hua; Cong-Hui Xiong; Bai-Nian Sun (2020). "New materials of Mariopteris from the Cisuralian of northwestern China and their implications for palaeogeographic diversification". Historical Biology: An International Journal of Paleobiology. in press: 1–15. doi:10.1080/08912963.2019.1675054.
73. Domingas Maria da Conceição; Alexandra Crisafulli; Roberto Iannuzzi; Rodrigo Neregato; Juan Carlos Cisneros; Luiz Saturnino de Andrade (2020). "New petrified gymnosperms from the Permian of Maranhão (Pedra de Fogo Formation), Brazil: Novaiorquepitys and Yvyrapitys". Review of Palaeobotany and Palynology. 276: Article 104177. doi:10.1016/j.revpalbo.2020.104177.
74. Ming-Li Wan; Wan Yang; Jun Wang (2020). "Palaeocupressinoxylon uniseriale n. gen. n. sp., a gymnospermous wood from the upper Permian of Central Taodonggou, southern Bogda Mountains, northwestern China". Palaeoworld. 29 (1): 117–125. doi:10.1016/j.palwor.2019.06.002.
75. John G. Conran; Jennifer M. Bannister; Uwe Kaulfuss; Daphne E. Lee (2020). "Pterostoma neehoffii (cf. Zamiaceae): a new species of extinct cycad from the middle Miocene of New Zealand and an overview of fossil New Zealand cycads". New Zealand Journal of Botany. 58 (1): 30–47. doi:10.1080/0028825X.2019.1653939.
76. Patrick Blomenkemper; Hans Kerp; Abdalla Abu Hamad; Benjamin Bomfleur (2020). "Contributions towards whole-plant reconstructions of Dicroidium plants (Umkomasiaceae) from the Permian of Jordan". Review of Palaeobotany and Palynology. 278: Article 104210. doi:10.1016/j.revpalbo.2020.104210.
77. Yifan Hua; Xuelian Wang; Junlin Dong; Yanzhao Ji; Bainian Sun (2020). "A number of new seed fossils from the lower Permian of Gansu, Northwest China: implication for research on arils". Historical Biology: An International Journal of Paleobiology. in press: 1–10. doi:10.1080/08912963.2019.1566323.
78. Natalya Nosova (2020). "Female reproductive structures of Umaltolepis Krassilov and associated short shoots, buds and leaves of Pseudotorellia Florin from the Middle Jurassic of Angren, Uzbekistan". Review of Palaeobotany and Palynology. 281: Article 104266. doi:10.1016/j.revpalbo.2020.104266.
79. Josef Pšenička; Jun Wang; Jason Hilton; Weiming Zhou; Jiří Bek; Stanislav Opluštil; Jana Votočková Frojdová (2020). "A small heterophyllous vine climbing on Psaronius and Cordaites trees in the earliest Permian forests of North China". International Journal of Plant Sciences. 181 (6): 616–645. doi:10.1086/708814.
80. Weiming Zhou; Shan Wan; Mingli Wan; Jason Hilton; Josef Pšenička; Jun Wang (2020). "Yangopteris ascendens (Halle) gen. et comb. nov., a climbing alethopterid pteridosperm from the Asselian (earliest Permian) Wuda Tuff Flora". Review of Palaeobotany and Palynology. in press: Article 104282. doi:10.1016/j.revpalbo.2020.104282.
81. Pedro Correia; Zbynĕk Šimůnek; Christopher J. Cleal; Bruno Vallois; Rúben Domingos; Artur A. Sa (2020). "On a new species of the calamitalean fossil-genus Annularia from the Douro Basin (lower Gzhelian; NW Portugal)". Historical Biology: An International Journal of Paleobiology. in press: 1–10. doi:10.1080/08912963.2019.1613391.
82. Pedro Correia; Arden R. Bashforth; Zbynĕk Šimůnek; Christopher J. Cleal; Artur A. Sá; Conrad C. Labandeira (2020). "The history of herbivory on sphenophytes: a new calamitalean with an insect gall from the Upper Pennsylvanian of Portugal and a review of arthropod herbivory on an ancient lineage". International Journal of Plant Sciences. 181 (4): 387–418. doi:10.1086/707105.
83. Xiao-Yuan He; Shi-Jun Wang; Jason Hilton; Jean Galtier; Hong-Guan Jiang (2020). "An advanced species of the fern Botryopteris Renault from the Permian of southwestern China". Review of Palaeobotany and Palynology. 273: Article 104136. doi:10.1016/j.revpalbo.2019.104136.
84. Barry A. Thomas (2020). "A new species of leafy calamite stem from the Pennsylvanian (Bolsovian) of the South Wales Coalfield". Acta Palaeobotanica. 60 (1): 207–211. doi:10.35535/acpa-2020-0010.
85. Jan Hinkelman; Lucia Vršanská (2020). "A Myanmar amber cockroach with protruding feces contains pollen and a rich microcenosis". The Science of Nature. 107 (2): Article number 13. doi:10.1007/s00114-020-1669-y. PMID 32125545.
86. Patricio Emmanuel Santamarina; Viviana Dora Barreda; Ari Iglesias; Augusto Nicolás Varela (2020). "Palynology from the Cenomanian Mata Amarilla Formation, southern Patagonia, Argentina". Cretaceous Research. 109: Article 104354. doi:10.1016/j.cretres.2019.104354.
87. Ioan I. Bucur; Sylvain Rigaud; Nicolò Del Piero; Andrea Fucelli; Eric Heerwagen; Camille Peybernes; Giovan Peyrotty; Christian Verard; Jérôme Chablais; Rossana Martini (2020). "Upper Triassic calcareous algae from the Panthalassa Ocean". Rivista Italiana di Paleontologia e Stratigrafia. 126 (2): 499–540. doi:10.13130/2039-4942/13681.
88. Xiu-Cai Yuan; Cong-Hui Xiong; Fan-Kai Sun; Zi-Xi Wang; Teng Mao; Yi-Jie Li; Chun-Hui Liu; Ming-Xuan Sun; Jun-Ling Dong; Bai-Nian Sun (2020). "The geological significance of a new species of Coniopteris from the Middle Jurassic of northwestern China". Historical Biology: An International Journal of Paleobiology. 32 (2): 267–280. doi:10.1080/08912963.2018.1488251.
89. Yang Yu; San-Ping Xie; John Devaney; Si-Hang Zhang; Tian-Yu Chen; Xu Zeng; Bing Wang; Yu Zhang (2020). "A new species of Drynaria (Polypodiaceae) from the late Miocene of Yunnan, Southwest China and implications on the genus evolution". Palaeobiodiversity and Palaeoenvironments. in press. doi:10.1007/s12549-020-00429-0.
90. George Poinar; Alex E. Brown (2020). "A green algae (Chaetophorales: Chaetophoraceae) in Burmese amber". Historical Biology: An International Journal of Paleobiology. in press: 1–5. doi:10.1080/08912963.2019.1616719.
91. Li Zhang; Yong Wang; Hong-Yu Chen; Lei Han; Yu-Xin Zhang; Wen-Jia Li; Tao Yang; Hao-Jian Wang; Lin Bao; De-Fei Yan (2020). "New fossil material of Equicalastrobus (Equisetales) and associated leaves from the Late Triassic of Baojishan basin, Gansu Province, China". Historical Biology: An International Journal of Paleobiology. in press: 1–12. doi:10.1080/08912963.2020.1716747.
92. Aye Thida Aung; Jian Huang; Truong Van Do; Ai Song; Jia Liu; Zhe-Kun Zhou; Tao Su (2020). "Three new fossil records of Equisetum (Equisetaceae) from the Neogene of south-western China and northern Vietnam". PhytoKeys. 138: 3–15. doi:10.3897/phytokeys.138.38674. PMC 6969027. PMID 31988601.
93. Ya Li; Yong-Dong Wang; Harald Schnerder; Peng-Cheng Wu (2020). "Frullania partita sp. nov. (Frullaniaceae, Porellales), a new leafy liverwort from the mid-Cretaceous of Myanmar". Cretaceous Research. 108: Article 104341. doi:10.1016/j.cretres.2019.104341.
94. Yuriy S. Mamontov; John J. Atwood; Evgeny E. Perkovsky; Michael S. Ignatov (2020). "Hepatics from Rovno amber (Ukraine): Frullania pycnoclada and a new species, F. vanae". The Bryologist. 123 (3): 421–430. doi:10.1639/0007-2745-123.3.421.
95. Fearghus R. McSweeney; Jeff Shimeta; John St. J. S. Buckeridge (2020). "Two new genera of early Tracheophyta (Zosterophyllaceae) from the upper Silurian–Lower Devonian of Victoria, Australia". Alcheringa: An Australasian Journal of Palaeontology. in press. doi:10.1080/03115518.2020.1744725.
96. Kelly C. Pfeiler; Alexandru M. F. Tomescu (2020). "An Early Devonian actinostelic euphyllophyte with secondary growth from the Emsian of Gaspé (Canada) and the importance of tracheid wall thickening patterns in early euphyllophyte systematics". Papers in Palaeontology. in press. doi:10.1002/spp2.1335.
97. Li-jun Chen; Ye-mao Hou; Peng-fei Yin; Xin Wang (2020). "An edible fruit from the Jurassic of China". China Geology. 3 (1): 8–15. doi:10.31035/cg2020010.
98. Antoine Champreux; Brigitte Meyer-Berthaud; Anne-Laure Decombeix (2020). "Keraphyton gen. nov., a new Late Devonian fern-like plant from Australia". PeerJ. 8: e9321. doi:10.7717/peerj.9321. PMC 7304422. PMID 32587800.
99. Sergey V. Naugolnykh; Ming Tu; Xiao-Yan Liu; Jian-Hua Jin (2020). "A new species of Lygodium (Schizaeaceae) from the Buxin Formation (Paleocene), Sanshui Basin, South China". Palaeoworld. in press. doi:10.1016/j.palwor.2019.07.003.
100. Dmitriy A. Mamontov; Duncan McLean; Olga A. Orlova; Olga A. Gavrilova (2020). "Maiaspora: a new miospore genus with enigmatic sculpture from the late Visean of European Russia". Papers in Palaeontology. in press. doi:10.1002/spp2.1278.
101. Silvia Gnaedinger; Bárbara Cariglino; Ana María Zavattieri; Mariana Monti; Pedro R. Gutiérrez (2020). "Neoarthropitys gondwanaensis gen. et sp. nov. from the Middle Triassic of Gondwana: An intermediate stage in the anatomical trend of Equisetalean stems". Review of Palaeobotany and Palynology. in press: Article 104298. doi:10.1016/j.revpalbo.2020.104298.
102. Yeming Cheng; Fengxiang Liu; Xiaonan Yang; Tongxing Sun (2020). "Two new species of Mesozoic tree ferns (Osmundaceae: Osmundacaulis) in Eurasia as evidence of long-term geographic isolation". Geoscience Frontiers. 11 (5): 1875–1888. doi:10.1016/j.gsf.2020.01.019.
103. Ana M. Zavattieri; Pedro R. Gutiérrez; Mariana Monti (2020). "Middle Triassic freshwater green algae and fungi of the Puesto Viejo Basin, central-western Argentina: palaeoenvironmental implications". Alcheringa: An Australasian Journal of Palaeontology. in press. doi:10.1080/03115518.2020.1749302.
104. Li Liu; Josef Pšenička; Jiří Bek; Mingli Wan; Hermann W. Pfefferkorn; Jun Wang (2020). "A whole calamitacean plant Palaeostachya guanglongii from the Asselian (Permian) Taiyuan Formation in the Wuda Coalfield, Inner Mongolia, China". Review of Palaeobotany and Palynology. in press: Article 104245. doi:10.1016/j.revpalbo.2020.104245.
105. Ruiyun Li; Xiaoqiang Li; Shenghui Deng; Bainian Sun (2020). "Morphology and microstructure of Pellites hamiensis nov. sp., a Middle Jurassic liverwort from northwestern China and its evolutionary significance". Geobios. in press. doi:10.1016/j.geobios.2020.07.003.
106. P. Hiller; M. Böhme; S. Schneider; J. Prieto; B. Bomfleur (2020). "Plenasium (Aurealcaulis) elegans sp. nov. from the Eocene of Vietnam – a connecting link in the evolution of modern Royal Ferns (Osmundeae, Osmundaceae)". Journal of Systematic Palaeontology. 18 (8): 703–715. doi:10.1080/14772019.2019.1683771.
107. Ye‐Ming Cheng; Feng‐Xiang Liu; Ning Tian; Yue‐Gao Jin; Tong‐Xing Sun (2020). "A new Cretaceous species of Plenasium from China Plenasium xiei sp. nov. from the Cretaceous of Northeast China: additional evidence for the longevity of osmundaceous ferns". Journal of Systematics and Evolution. in press. doi:10.1111/jse.12532.
108. Yi Zhang; Yong-Dong Wang; Yue Hong; Liu Cao; Fu-liang Gao (2020). "Pleuromeia discovered from the Middle Triassic Linjia Formation of Benxi, Northeast China". Palaeoworld. in press. doi:10.1016/j.palwor.2020.02.004 (inactive 2020-03-06).
109. Carmen Álvarez-Vázquez; Jiří Bek; Jana Drábková (2020). "Polysporia baetica sp. nov., a new heterosporous sub-arborescent isoetalean from lower Bolsovian (Middle Pennsylvanian) strata of the Peñarroya-Belmez-Espiel Coalfield (Córdoba, SW Spain)". Review of Palaeobotany and Palynology. 272: Article 104115. doi:10.1016/j.revpalbo.2019.104115.
110. Zlatko Kvaček; Vasilis Teodoridis (2020). "A new Oligocene fern of Dryopteridaceae from the Českéstředohoří Mts (Czech Republic)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 295 (1): 9–16. doi:10.1127/njgpa/2020/0864.
111. Chunxiang Li; Robbin C. Moran; Junye Ma; Bo Wang; Jiasheng Hao (2020). "A new fossil record of Lindsaeaceae (Polypodiales) from the mid-Cretaceous amber of Myanmar". Cretaceous Research. 105: Article 104040. doi:10.1016/j.cretres.2018.12.010.
112. Qing Tang; Ke Pang; Xunlai Yuan; Shuhai Xiao (2020). "A one-billion-year-old multicellular chlorophyte". Nature Ecology & Evolution. 4 (4): 543–549. doi:10.1038/s41559-020-1122-9. PMID 32094536.
113. Josep Sanjuan; Ingeborg Soulié-Märsche (2020). "New charophyte assemblage from middle Miocene lacustrine deposits of Moneva (Ebro Basin, Spain)". Geobios. 59: 79–90. doi:10.1016/j.geobios.2020.03.001.
114. Pu Huang; Le Liu; Lu Liu; Min Qin; De-Ming Wang; Jin-Zhuang Xue (2020). "A new plant with novel leaves from the Upper Devonian of Zhejiang Province, China". Palaeoworld. in press. doi:10.1016/j.palwor.2020.03.003.
115. Mingli Wan; Wenjun Sun; Jiří Bek; Feng Liu; Christopher Hill; Jun Wang (2020). "Scolecopteris minuta sp. nov., a marattialean fern from the early Permian Wuda Tuff Flora of Inner Mongolia, China". Review of Palaeobotany and Palynology. in press: Article 104246. doi:10.1016/j.revpalbo.2020.104246.
116. Michael P. D'Antonio; C. Kevin Boyce; Jun Wang (2020). "Two new species of Sigillaria Brongniart from the Wuda Tuff (Asselian: Inner Mongolia, China) and their implications for lepidodendrid life history reconstruction". Review of Palaeobotany and Palynology. in press: Article 104203. doi:10.1016/j.revpalbo.2020.104203.
117. Chunxiang Li; Robbin C. Moran; Junye Ma; Bo Wang; Jiasheng Hao; Qun Yang (2020). "A mid-Cretaceous tree fern of Thyrsopteridaceae (Cyatheales) preserved in Myanmar amber". Cretaceous Research. 105: Article 104050. doi:10.1016/j.cretres.2019.01.002.
118. Zhuo Feng; Hai-Bo Wei; Yun Guo; Xiao-Yuan He; Qun Sui; Yu Zhou; Hang-Yu Liu; Xu-Dong Gou; Yong Lv (2020). "From rainforest to herbland: New insights into land plant responses to the end-Permian mass extinction". Earth-Science Reviews. 204: Article 103153. doi:10.1016/j.earscirev.2020.103153.
119. Christine Strullu‐Derrien; Alain Le Hérissé; Tomasz Goral; Alan R.T. Spencer; Paul Kenrick (2020). "The overlooked aquatic green algal component of early terrestrial environments: Triskelia scotlandica gen. et sp. nov. from the Rhynie cherts". Papers in Palaeontology. in press. doi:10.1002/spp2.1303.
120. Zhen Tang; Yi Zhang; Serge V. Naugolnykh; Changqing Zheng; Lu Shi; Tao Qin; Junhao Huang (2020). "Ufadendron elongatum sp. nov., an Angaran lycopsid from the Upper Permian of Inner Mongolia, China". Journal of Earth Science. 31 (1): 1–8. doi:10.1007/s12583-019-1230-0.
121. Barry A. Thomas; Peter Appleton; Christopher J. Cleal; Leyla J. Seyfullah (2020). "The distribution of plant fossils and their palaeoecology in Duckmantian (Bashkirian, Lower Pennsylvanian) strata at Brymbo, North Wales, UK". Geological Journal. 55 (4): 3179–3207. doi:10.1002/gj.3529.
122. Yuan Nie; Charles S. P. Foster; Tianqi Zhu; Ru Yao; David A. Duchêne; Simon Y. W. Ho; Bojian Zhong (2020). "Accounting for uncertainty in the evolutionary timescale of green plants through clock-partitioning and fossil calibration strategies". Systematic Biology. 69 (1): 1–16. doi:10.1093/sysbio/syz032. PMID 31058981.
123. Xiaodong Shang; Pengju Liu; Małgorzata Moczydłowska; Ben Yang (2020). "Algal affinity and possible life cycle of the early Cambrian acritarch Yurtusia uniformis from South China". Palaeontology. in press. doi:10.1111/pala.12491.
124. Alexander J. Hetherington; Christopher M. Berry; Liam Dolan (2020). "Multiple origins of dichotomous and lateral branching during root evolution". Nature Plants. 6 (5): 454–459. doi:10.1038/s41477-020-0646-y. PMID 32366983.
125. Nikole K. Bonacorsi; Patricia G. Gensel; Francis M. Hueber; Charles H. Wellman; Andrew B. Leslie (2020). "A novel reproductive strategy in an Early Devonian plant". Current Biology. 30 (9): R388–R389. doi:10.1016/j.cub.2020.03.040. PMID 32369746.
126. Neil Brocklehurst; Christian F. Kammerer; Roger J. Benson (2020). "The origin of tetrapod herbivory: effects on local plant diversity". Proceedings of the Royal Society B: Biological Sciences. 287 (1928): Article ID 20200124. doi:10.1098/rspb.2020.0124. PMC 7341937. PMID 32517628.
127. Michael P. D'Antonio; C. Kevin Boyce (2020). "Arborescent lycopsid periderm production was limited". New Phytologist. in press. doi:10.1111/nph.16727. PMID 32506426.
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