Paleobiota of the Posidonia Shale

The Posidonia Shale or Posidonienschiefer Formation is a geological formation of southwestern Germany, northern Switzerland, western Czech Republic, northwestern Austria, southeast Luxembourg and the Netherlands, that spans about 3 million years during the Early Jurassic period (early Toarcian stage). It is known for its detailed fossils, especially sea fauna, listed below.[1] Composed mostly by black shale, the formation is a Lagerstätte, where fossils show exceptional preservation (Including exquisite soft tissues), with a thickness that varies from about 1 m to about 40 m on the Rhine level, being on the main quarry at Holzmaden between 5 and 14 m.[1] Some of the preserved material has been transformed into fossil hydrocarbon Jet, specially wood remains, used for jewelry.[2] The exceptional preservation seen on the Posidonia Shale has been studied since the late 1800s, finding that a cocktail of chemical and environmental factors let to such an impressive conservation of the marine fauna.[2] The most common theory is the changes on the oxygen level, where the different anoxic events of the Toarcian left oxygen-depleted bottom waters, with the biota dying and falling to the bottom without any predator able to eat the dead bodies.[3]

18x6 m fossilized floating wood (Araucarioxylon), with Crinoids attached (Pentacrinites & Seirocrinus). It is one of the most emblematic fossils of the formation, where the anoxic seas of the lower toarcian lead to an exquisite preservation.

Microbial Activity

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Genus Species Location Material Notes Images

Stromatolites[4]

  • Non assignable Species
  • Teufelsgraben, Hetzles

Traces of Microbial Activity

Non-fenestrate stromatolite crusts formed in Aphotic deep-water environments during intervals of very low sedimentation.[4] Abundant on the Precambrian, but after it, and concretely on the Jurassic-Cretaceous, the appearance of the Corallinaceae algae and related biota forced the stromatolite-forming microbes to withdraw to extreme habitats such as hypersaline lagoons and possibly to deep-water settings. On the Posidonia Shale are related with plankton, mainly coccoliths and the problematic Schizosphaerella (A Haptophytan Alga), but also typical deep-sea forms including various groups of cephalopods, and articulated skeletons of fishes and reptiles.[4] The Stromatolites of this region have evidence of live on a deeper shelf environment with a quietwater deposit which suffered repeated phases of stagnant bottom waters, where a depth water habitat developed, probably at more than 100 meters depth.[4] There is a thin, southern widespread Stromatolite crust on the Top of the Posidonia Shale, called "Wittelshofener Bank", that has made rethink the depth of the major southern basin of the formation, where with the absence of phototrophic calcareous benthic organisms (probably due to the lack of light), shows the deph character of the Basin.[4] On the "Wittelshofener Bank" there is also the only occurrence of Ooids, presumably formed in the same deep-water environment.[4]

Example of Stromatolites fossil trace, done probably by Cyanobacteria

Frutexites[4]

  • Frutexites arboriformis
  • Teufelsgraben, Hetzles

Possible traces of Microbial Activity

Probably related with Archaea activity.[4] Altrought Frutexites is a cryptic microfossil and an important element of many deep water stromatolites, with an Inorganic origin proposed, where are interpreted as dendritic shrubs to purely inorganic growth of Aragonitic crystals, but also resemble shrubs of the cyanobacteria Angulocellularia.[4] On the Posidonia a cryptoendopelitic mode of life is assumed, being only possible for Heterotrophic bacteria or Fungi.[4] As seen on the Stromatolites of the Posidonia, Frutexites acted mainly as a dweller or secondary binder of the deep-water stromatolites, not as their major constructor.[4]

Rhizaria

Foraminifera

Genus Species Location Stratigraphic position Material Notes Images

Astacolus[5]

  • Astacolus bochardi
  • Astacolus primus
  • Buttenheim
  • All the levels

Shells

A benthonic Foraminiferan, member of Vaginulinidae inside the family Vaginulinida (Lagenina). An extant genus. Its shell resemble a mixture between an ammonite conch, due to having a lower spiral, and a mussel.

Drawing of an Astacolus shell

Flabellinella[5]

  • Flabellinella sp.
  • Buttenheim
  • All the levels

Shells

A benthonic Foraminiferan, member of Vaginulinidae inside the family Vaginulinida (Lagenina). Its conch has a Myriapod-like segmented built.

Lenticulina[5][6]

  • Lenticulina acutiangulata
  • Lenticulina gottingensis
  • Lenticulina sp.
  • Buttenheim
  • Unterstürmig
  • All the levels

Shells

A benthonic Foraminiferan, member of Vaginulinidae inside the family Vaginulinida (Lagenina).

Vaginulina[5][6]

  • Vaginulina simplex
  • Vaginulina sp.
  • Buttenheim
  • Unterstürmig
  • All the levels

Shells

A benthonic Foraminiferan, type member of Vaginulinidae inside the family Vaginulinida (Lagenina).

Palmula[5]

  • Palmula cuneiformis
  • Buttenheim
  • All the levels

Shells

A benthonic Foraminiferan, member of Vaginulinidae inside the family Vaginulinida (Lagenina).

Marginulina[6]

  • Marginulina oolithica
  • Marginulina prima
  • Unterstürmig
  • All the levels

Shells

A benthonic Foraminiferan, member of Marginulininae inside the family Vaginulinida (Lagenina).

Saracenaria[6]

  • Saracenaria aragonensis
  • Unterstürmig
  • All the levels

Shells

A benthonic Foraminiferan, member of Lenticulininae inside the family Vaginulinida (Lagenina).

Cornuspira[5]

  • Cornuspira involvens
  • Buttenheim
  • All the levels

Shells

A benthonic Foraminiferan, type member of Cornuspiridae inside the family Cornuspirida (Lagenina). Round-spiral shell morphology

Dentalina[5][6]

  • Dentalina terquiemi
  • Dentalina matutina
  • Dentalina vetusta
  • Dentalina sp.
  • Buttenheim
  • Unterstürmig
  • All the levels

Shells

A benthonic Foraminiferan, member of Nodosariidae inside the family Nodosariacea (Lagenina). Dentalina is an extant genus, with an elongated shell, that resemble a small worm.

Pseudonodosaria[6]

  • Pseudonodosaria melo
  • Pseudonodosaria quinquecostata
  • Unterstürmig
  • All the levels

Shells

A benthonic Foraminiferan, member of Nodosariidae inside the family Nodosariacea (Lagenina).

Ichthyolaria[5]

  • Ichthyolaria squamosa
  • Ichthyolaria sp.
  • Buttenheim
  • All the levels

Shells

A benthonic Foraminiferan, type member of Ichthyolariidae inside the family Lagenina. Another genus with a Myriapod-like segmented built.

Lingulina[6]

  • Lingulina pupa
  • Lingulina tenera
  • Unterstürmig
  • All the levels

Shells

A benthonic Foraminiferan, type member of Lingulininae inside the family Nodosariidae (Lagenina). Dentalina is an extant genus, with an elongated shell, that resemble a small worm.

Reinholdella[5]

  • Reinholdella sp.
  • Buttenheim
  • All the levels

Shells

A benthonic Foraminiferan, member of Ceratobuliminidae inside the family Robertinida . It resembles a small ammonite.

Dinoflagellata

Dinoflagellate cysts

Genus Species Location Stratigraphic position Material Notes Images

Nannoceratopsis[7][8]

  • Nannoceratopsis gracilis
  • Nannoceratopsis senex
  • Nannoceratopsis ridingii
  • Nannoceratopsis tricornuta[9]
  • Nannoceratopsis deflandrei
  • Nannoceratopsis triceras
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst, member of Dinophyceae of the family Nannoceratopsiaceae. On the Lias Epsylon Interval (Lowermost Toarcian), most of the assemblages are dominated by Nannoceratopsis gracilis. Nannoceratopsis senex becomes highly abundant until the uppermost Tenuicostatum.[7]

Comparodinium[9]

  • Comparodinium koessenium
  • Comparodinium lineatum
  • Comparodinium punctatum
  • Comparodinium scalatum
  • Comparodinium stipulatum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Comparodiniaceae.

Valvaeodinium[7][8]

  • Valvaeodinium punctatum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Comparodiniaceae.

Apodinium[10]

  • Apodinium fioccosum
  • Apodinium glabrum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Apodiniaceae. An Ectoparasitic dinoflagellate, whose hosts are normally Tunicates

Eyachia[10]

  • Eyachia priscus
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Scriniocassiaceae.

Argentiella[10]

  • Argentiella bifuminosa
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Scriniocassiaceae.

Scriniocassis[10]

  • Scriniocassis weberi
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Scriniocassiaceae.

Balechiodinium[10]

  • Balechiodinium concicum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Scriniocassiaceae.

Moesiodinium[10]

  • Moesiodinium cingulatum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Heterocapsaceae.

Morgenrothia[10]

  • Morgenrothia junior
  • Morgenrothia tenera
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Heterocapsaceae.

Susadinium[10]

  • Susadinium cristatum
  • Susadinium flaccum
  • Susadinium saetosum
  • Susadinium scrofoides
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Heterocapsaceae.

Parvocysta[10]

  • Parvocysta nasuta
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Heterocapsaceae.

Surculosphaeridium[11][12]

  • Surculosphaeridium longifurcatum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Gonyaulacaceae.

Phallocysta[11][12]

  • Phallocysta minuta
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst from the family Peridiniphycidae.

Mancodinium[7][8]

  • Mancodinium semitabulatum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst, type member of Mancodiniaceae. Dominant genera on some layers of the Lias Delta Stage.[7]

Maturodinium[8]

  • Maturodinium inornatum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst, member of Mancodiniaceae. Commonly found along the genus Beaumontella.[8]

Luehndea[7][8]

  • Luehndea spinosa
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst, type member of Luehndeoideae. Luehndea spinosa is common on the medium layers of the lower Posidonia Shale, while restricted to some areas on the Lias delta.[7]

Beaumontella[8]

  • Beaumontella caminuspina
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A Dinoflagellate cyst, member of Suessiaceae. Common on Pliensbachian levels, become present but rare on lower Toarcian.[8]

Algae

Includes abundant variety of algae, such as the genus of colonial Green algae Botryococcus,[13] or the unicellular algal bodies Tasmanites, and other small examples. Algae are a good reference for changes on the oxygen conditions along the Toarcian.[14]

Algae Acritarchs

Genus Species Location Stratigraphic position Material Notes Images

Veryhachium[7][8]

  • Veryhachium brevispinum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

An Acritarch probably from Algal origin. Veryhachium fossils represent open marine and transgressive conditions. It has high presence on most of the samples studied from the Posidonia Shale, being nearly the 50% of the Acritarch fraction on some locations.

Micrhystridium[7][8]

  • Micrhystridium inconspicuum
  • Micrhystridium spinuliferum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

An Acritarch probably from Algal origin. Its fossils indicate nearshore or estuarine to shallow lagoon and/or slightly brackish-water environments. It is the dominant on the nearshore sections.

Leiofusa[8]

  • Leiofusa jurassica
  • Leiofusa fusiformis
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

An Acritarch probably from Algal origin. Related to estuarine deposits.

Pterospermopsis[8]

  • Pterospermopsis pelagica
  • Pterospermopsis goslarensis
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

An Acritarch probably from Algal origin. Related to open shelf deposits

Cymatiosphaeropsis[15]

  • Cymatiosphaeropsis punctiferus
  • Cymatiosphaeropsis stigmatus
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

An Acritarch probably from Algal origin. Related to open shelf deposits

Pterosphaeridia[8][15]

  • Pterosphaeridia undulata
  • Pterosphaeridia eisenackii
  • Pterosphaeridia intersignata
  • Pterosphaeridia nodosa
  • Pterosphaeridia pachytheca
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

An Acritarch probably from Algal origin. Related to open shelf deposits

Haptophyta

Genus Species Location Stratigraphic position Material Notes Images

Mitrolithus[16]

  • Mitrolithus jansae
  • Lower Saxony Basin
  • All the levels

Millions of specimens

A member of the family Parhabdolithaceae inside Stephanolithiales. Shore deposits genus. The abundance drop of M. jansae further characterise the T-OAE perturbation, where becomes the dominant Genus on most of the Saxony Basin.

Parhabdolithus[16]

  • Parhabdolithus liasicus
  • Lower Saxony Basin
  • All the levels

Millions of specimens

Type member of the family Parhabdolithaceae inside Stephanolithiales.

Schizosphaerella[16]

  • Schizosphaerella punctulata
  • Lower Saxony Basin
  • All the levels

Millions of specimens

Type member of the family Schizosphaerellaceae inside Parhabdolithaceae. Towards the Pliensbachian-Toarcian extincion this genus gets a decrease in abundance and size that shows the change and biotic crisis.

Biscutum[16]

  • Biscutum grandis
  • Lower Saxony Basin
  • All the levels

Millions of specimens

Type member of the family Biscutaceae inside Parhabdolithaceae.

Crepidolithus[16]

  • Crepidolithus granulatus
  • Lower Saxony Basin
  • All the levels

Millions of specimens

A member of the family Chiastozygaceae inside Eiffellithales.

Chlorophyta

Genus Species Location Stratigraphic position Material Notes Images

Scriniocassis[11][12]

  • Scriniocassis limbatus
  • Scriniocassis limbicavatus
  • Scriniocassis priscus
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of Dinophyceae.

Palaeohystrichophora[11][12]

  • Palaeohystrichophora infusorioides
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of Peridiniaceae inside Dinophyceae.

Dissiliodinium[11][12]

  • Dissiliodinium giganteum
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of Gonyaulacaceae inside Dinophyceae.

Pleurozonaria[8][15][17]

  • Pleurozonaria globulus
  • Pleurozonaria chondrota
  • Pleurozonaria concinna
  • Pleurozonaria digitata
  • Pleurozonaria distans
  • Pleurozonaria diversipora
  • Pleurozonaria macropora
  • Pleurozonaria media
  • Pleurozonaria polyporosa
  • Pleurozonaria spongiosa
  • Pleurozonaria stellulata
  • Pleurozonaria suevica
  • Pleurozonaria wetzelii
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of Prasinophyceae. It the main genus present on silt and sand horizons, trending to be absent on black argillaceous layers.

Campenia[8][15][17]

  • Campenia minor
  • Campenia gigas
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of Prasinophyceae. A genus common on green clays and other upper strata on the formation.

Tasmanites[15][17][18]

  • Tasmanites mourai
  • Tasmanites tardus
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of Prasinophyceae. A genus common on green clays and other upper strata on the formation.

Cyrnatiosphaera[7][15][17]

  • Cyrnatiosphaera densisepta
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of Prasinophyceae. A genus common on green clays and other upper strata on the formation.

Leiosphaera[7][15]

  • Leiosphaera globosa
  • Leiosphaeridia deflandrei
  • Leiosphaeridia pusilla
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of the Prasinophyceae. Basinal deposits genus

Lancettopsis[15]

  • Lancettopsis lanceolata
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of the Prasinophyceae. Basinal deposits genus

Nostocopsis[15]

  • Nostocopsis saprolithica
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of the Prasinophyceae. Basinal deposits genus

Granodiscus[15]

  • Granodiscus granulatus
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of the Prasinophyceae. Basinal deposits genus

Tytthodiscus[15]

  • Tytthodiscus chondrotus
  • Tytthodiscus schandelahensis
  • Tytthodiscus suevicus
  • Tytthodiscus cf. suevicus
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of the Prasinophyceae. Basinal deposits genus

Cymatiosphaera[7]

  • Cymatiosphaera areolata
  • Cymatiosphaera punctifera
  • Cymatiosphaera stigmata
  • Cymatiosphaera tecta
  • Cymatiosphaera pachytheca
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of the family Pyramimonadales inside Prasinophyceae. Basinal deposits genus

Halosphaeropsis[7]

  • Halosphaeropsis liassica
  • All the epicontinental, basinal & nearshore sections of the formation
  • All the levels

Millions of specimens

A member of the family Halosphaeraceae inside Chlorodendrales. Basinal deposits genus

Botryococcus[15]

  • Botryococcus braunii
  • Botryococcus luteus
  • Deltaic Influenced Zones of the Formation
  • All the levels

Millions of specimens

Type member of the family Botryococcaceae inside Trebouxiales. Freshwater or Deltaic Genus

Plantae

The Flora is dominated by horsetails, what can be due to a similar ecological niche to modern genus Phragmites, able to resist saline conditions. Storms or floods maybe where the major events that transported this flora to the sea.

The macroflora of the Posidonia slate can be described as extremely poor in species.[19] Apart from the remains of Horsetails, it is without exception the remains of coarse branches and fronds from gymnosperms, in which one has a certain can assume transport resistance. Remains of Ferns are completely missing, except for tall arboreal ferns (Peltaspermales).[20] Mostly of the flora was reported from the area of Braunschweig.[19] The major explanation for the flora could be that the plants in question are mono-or oligotypic stands on the edge of the waters that flow into the Posidonienschiefer sea, probably tear away in the course of flood events, easily fragmented during transport and wave waves, possibly especially in the occasional storm events postulated.[21] In terms of taphonomy, this would result in a comparison with today's reed Phragmites, which can form extensive stocks on the edge of shallower and slowly flowing waters ("Reed belts").[19] The Wood remnants clearly indicate one higher diversity of Coniferous flora in the delivery area than the remains of leafy branches.[19] This fact is likely to be proportionate, similar to that frequent occurrence of charcoalized or gagged trunks, mostly of them are believed to be "driftwoods" that only take a long time drifting also suggests a frequent settlement with mussels and full-grown Sea Lilies.[19][21] The deposition settings are at large distance from the nearest coastline (for southern Germany about 100 kilometers), making only plants strong to transportation able to resist enougth to get deposited.[22][23]

Palynology

Genus Species Location Stratigraphic position Material Notes Images

Aratrisporites[24][25]

  • Appendicisporites tricuspidatus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Lycopsida. Most Abundant Pollen on the Bohemian realm

Heliosporites[25]

  • Heliosporites altmarkensis
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Lycopsida.

Leptolepidites[25]

  • Leptolepidites bossus
  • Leptolepidites macroverrucosus
  • Leptolepidites major
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Lycopsida.

Lycospora[25]

  • Lycospora salebrosacea
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Lycopsida.

Neochomotriletes[25]

  • Neochomotriletes triangularis
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Lycopsida.

Uvaesporites[25]

  • Uvaesporites argenteaeformis
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Lycopsida.

Zebrasporites[25]

  • Zebrasporites interscriptus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Lycopsida.

Retitriletes[25]

  • Retitriletes clavatoides
  • Retitriletes globosus
  • Retitriletes gracilis
  • Retitriletes subrotundus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Lycopodiaceae.

Simozonotriletes[25]

  • Simozonotriletes arcuatus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Lycopodiaceae.

Verrucosisporites[25]

  • Verrucosisporites obscurilaesuratus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Isoetales.

Foraminisporis[25]

  • Foraminisporis jurassicus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Bryophyta.

Appendicisporites[11][12]

  • Appendicisporites tricuspidatus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Schizaeales.

Densoisporites[25]

  • Densoisporites velatus
  • Densosporites variabilis
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Schizaeales.

Trilites[25]

  • Trilites minutus
  • Trilites lygodioides
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Selaginellaceae.

Crassipollenites[15][26]

  • Crassipollenites diffusus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Pollen

Affinities with Gymnospermophyta. Non concreted affinities

Ovalipollis[15][26]

  • Ovalipollis ovalis
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Pollen

Affinities with Gymnospermophyta. Non concreted affinities

Baculatisporites[25]

  • Baculatisporites primarius
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Pteridopsida.

Polycingulatisporites[25]

  • Polycingulatisporites liassicus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Pteridopsida.

Leiotriletes[25]

  • Leiotriletes mesozoicus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Pteridopsida.

Converrucosisporites[25]

  • Converrucosisporites luebbenensis
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Pteridopsida.

Contignisporites[25]

  • Contignisporites dunrobinensis
  • Contignisporites problematicus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Pteridopsida.

Lycopodiacidites[25]

  • Lycopodiacidites rugulatus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Ophioglossaceae.

Gleicheniidites[25]

  • Gleicheniidites umbonatus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Gleicheniaceae.

Spheripollenites[7][8][26]

  • Spheripollenites subgranulatus
  • Spheripollenites classopolloides
  • Spheripollenites laceratus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Pollen

Affinities with Bennettitales. Abundant on the Lower Jurassic of NW Europe.

Quadraeculina[25]

  • Quadraeculina anellaeformis
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Pollen

Affinities with Bennettitales.

Eucommiidites[25]

  • Eucommiidites granulosus
  • Eucommiidites troedssonii
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Bennettitales.

Ephedripites[25]

  • Ephedripites tortuosus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Ginkgoopsida.

Callialasporites[11][12]

  • Callialasporites dampieri
  • Callialasporites turbatus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Pinopsida.

Inaperturopollenites[7][26]

  • Inaperturopollenites orbiculatus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Pollen

Affinities with Pinidae. Abundant on the Lower Jurassic of NW Europe, represents pollen of medium to large arboreal plants, specially coniferales.

Cerebropollenites[25]

  • Cerebropollenites mesozoicus
  • Cerebropollenites macroverrucosus
  • Cerebropollenites thiergartii
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Pollen

Affinities with Cupressaceae inside Pinaceae.

Classopollis[15]

  • Classopollis classoides
  • Classopollis dellassis
  • Classopollis reclusus
  • Classopollis striatus
  • Classopollis tetradenverband
  • Classopollis torosus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Pollen

Affinities with Cheirolepidiaceae. Abundant on the Lower Jurassic of North and Southern Europe, represents pollen of medium to large arboreal plants, specially coniferales. The abundance of pollen of Classopollis and other thermophile plants was observed in this region in the lower Toarcian from the end of the antiquum (= tenuicostatum) zone to the middle of commune zone.[27]

Circumpollis[15][26]

  • Circumpollis pharisaeus
  • Circumpollis philosophus
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Pollen

Affinities with Cheirolepidiaceae. Pollen of medium to large arboreal plants, specially coniferales.

Disaccites[15][26]

  • Disaccites grandis
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Spores

Affinities with Podocarpaceae. Pollen of medium to large arboreal plants, specially coniferales.

Clavatipollenites[25]

  • Clavatipollenites hughesii
  • Grimmen
  • Mistelgau
  • Niedersachsen
  • Schandelah
  • All the levels

Pollen

Shares morphology with Chloranthaceae pollen. Can be an example of primigenial Angiosperms pollen, more probably pollen related to genera such as Sanmiguelia and other primigenial Flower-Plant like flora.

Equisetaceae

Genus Species Location Stratigraphic position Material Notes Images

Equisetites[19]

  • cf. Equisetites bunburyanus
  • Altdorf
  • Holzmaden
  • Ohmden
  • Niedersachsen
  • Schandelah
  • Tenuicostatum
  • Falciferum
  • Bifrons

Stems

Affinities with Equisetaceae inside Equisetopsida. Number of mostly very fragmented and not particularly well preserved, but clear horsetail remains described. So far Recognizable, leaf sheaths were developed in most cases, but the state of preservation does not allow a more precise determination.[28]

Neocalamites[23][29][30]

  • Neocalamites merianii
  • Altdorf
  • Holzmaden
  • Ohmden
  • Kerkhofen
  • Niedersachsen
  • Schandelah
  • Bascharange
  • Tenuicostatum
  • Falciferum
  • Bifrons

Stems and incomplete axes

Affinities with Equisetaceae inside Equisetopsida. Neocalamites is the most common more distributed of all the Posidonia Shale, being even found on Luxembourg Posidonia Strata.[30] Mostly of the Stems reported come from Aeolian-Dunar related deposits, or from nearshore-basinal deposition. Probably was related to the seashore.[30] Some stems are big, resembling the rates of growth seen on modern Bamboo specimens, suggesting +6–7 m tall Equisetopsids.[30]

Neocalamites merianii specimen

Palaeostachya?[31]

  • Palaeostachya? sp.
  • Dotternhausen
  • Tenuicostatum

Pollen Cones

Affinities with Calamitaceae inside Equisetopsida. It was an approximately 2–3 m high shrub-like plant related to swamp environments. Calamitaceans are common in late Palaeozoic wetland plant communities, so the find of a jurassic specimen is rare.[31] Maybe is a mistaken new genera of Pollen cone, but definitely come from an Equisetalean. This genus is found associated with Annularia, being both part of the Calamites plant. It can be related with the Equisetites stems found on the formation.

Palaeostachya drawing of a specimen

Pteridospermatophyta

Genus Species Location Stratigraphic position Material Notes Images

Pachypteris[32]

  • Pachypteris nordenskioeldii
  • Braunschweig
  • Falciferum
  • One specimen of pteridosperm frond.

Affinities with Umkomasiaceae inside Corystospermaceae. Is based on bipinnate leaves, rachis longitudinally striated, with a long petiole and secondary rachises. It belongs to large tree ferns. Posidonia Shale Specimen is characterized for its large size and probably where attached to trunks similar in built to the Cretaceous genus Tempskya.[32]

The only Pachypteris specimen know from the Posidonia Shale

Cycadeoidophyta

Genus Species Location Stratigraphic position Material Notes Images

Otozamites[33][34]

  • Otozamites gracilis
  • Otozamites bechei
  • Banz
  • Holzmaden
  • Ohmden[19]
  • Tenuicostatum
  • Falciferum
  • Bifrons
  • Leaflets

Affinities with Cycadeoidaceae inside Bennettitales. It is the most abundant medium-sized plant on the environment. Found specially on seashore depositional settings, but also on deltaic and lagoonar environments. It was a low arbustive-arboreal Bennetite related to arid environments, with a leave similar of that of the modern genus Encephalartos, specially Encephalartos munchii, but also Dioon mejiae. Otozamites has been considered synonym with Otopteris, but since the 1990s everybody used the name Otozamites, and Otopteris was forgotten.[35]

Otozamites gracilis specimen from the Posidonia Shale

Pterophyllum[28][33]

  • Pterophyllum acutifolium
  • Banz
  • Holzmaden
  • Altdorf[19]
  • Tenuicostatum
  • Falciferum
  • Bifrons
  • Leaflets

Affinities with Cycadeoidaceae inside Bennettitales. This Bennettitalean is related to Shrub built. Some specimens where assigned to Dioonites acutifolium (Junior synonym). Consits on leaves with pinnate, leaflets perpendicular or oblique to the rhachis, on the top of the leaf axis. It comes from fragments of fairly large fronds. In its external form it closely follows the modern genus Dioon, only the leaflets are of our kind wider and shorter, they are further apart, but the tendency to rhachis is the same.

Ptilophyllum[28][28][23]

  • Ptilophyllum gracilis
  • Ptilophyllum sp.
  • Banz
  • Ohmden
  • Holzmaden
  • Tenuicostatum
  • Kerkhofen
  • Falciferum
  • Bifrons
  • Leaflets

Affinities with Williamsoniaceae inside Bennettitales. Arboreal Cycadaceans, some with the presence of flower-like structures. It resembles the leaf of the modern Microcycas calocoma, and probably had a similar arboreal built, being the leave of tall Bennetite trees such as Bucklandia, found on the middle jurassic of England.

Ptilophyllum sp. specimen from the Posidonia Shale

Zamites[28][33]

  • Zamites mandelslohi
  • Zamites oblongifolius
  • Banz
  • Irlbach
  • Holzmaden
  • Altdorf[19]
  • Tenuicostatum
  • Falciferum
  • Bifrons
  • Leaflets

A member of Williamsoniaceae inside Bennettitales. It has been interpreted as a cycad in the family Cycadaceae or a Bennettitalean plant. Leaflets somewhat removed, oval-oblong, a little narrower near the base, rounded at the tip, nerves partially diverging from the base towards the edge. It was assigned to Pterophyllum oblongifolium and on the genus Glossozamites. This genus was the leaf of arboreal Bennetites, similar in appearance to the modern Encephalartos woodii.

Zamites mandelslohi specimen

Ginkgoales

Genus Species Location Stratigraphic position Material Notes Images

Baiera[28][24][20]

  • Baiera furcata
  • Baiera gracilis
  • Ohmden
  • Holzmaden
  • Banz
  • Tenuicostatum
  • Falciferum
  • Bifrons
  • Leave Compressions

Affinities with Ginkgoaceae inside Ginkgoales. Various leaves are known from Ohmden, coming from marine deposits and where identified as Ginko digitata, identified originally as the only Ginko specimen reported on the Posidonia Shale.

Baiera reconstruction

Pinophyta

Genus Species Location Stratigraphic position Material Notes Images

Pagiophyllum[33][36]

  • Pagiophyllum kurri
  • Pagiophyllum araucarinum[37]
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen[19]
  • Tenuicostatum
  • Falciferum
  • Bifrons
  • Fragmentary axis compressions with preserved leaves

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. Pagiophyllum araucarinum predominates among the two types of leafy coniferous branches that have become known from the Posidonia. However, there is no indication whether this fact reflects their respective share in the vegetation of the delivery area. Cheirolepidiaceae Pollen is the most abundant and diverse found on the formation, what is correlated with the abundance of this genus. Other factor that puts local Pagiophyllum on Cheirolepidiaceae is the dominance of an arid climate, the preferred for this type of conifers.

Pagiophyllum kurri specimen from Banz

Brachyphyllum[38]

  • Brachyphyllum sp.
  • Holzmaden
  • Tenuicostatum
  • Fragmentary axis

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. Specimens whose spiral foliage of the branch, in which the individual leaves open about 2/3 of their length (without the tip) are fused with the branch they hold in their free part fits tightly, suggests belonging to the genus Brachyphyllum.

Widdringtonites[38][39]

  • Widdringtonites liasinus
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Tenuicostatum
  • Falciferum
  • Bifrons
  • Fragmentary axis compressions with preserved leaves

A possible ancestral member of the Sequoioideae or Callitroideae inside Cupressaceae. Was originally related to the genus "Quasisequoia" couttsiae, as a sister taxa to Sequoia jeholensis. The later studies suggest affinities with large modern redwoods, but others find relationships with the modern genus Fitzroya cupressoides. Both Genera are related with relatively humid ecosystems, while Widdringtonites is known from an Arid setting.

Widdringtonites liasinus specimen from the Posidonia Shale

Taxodioxylon[40]

  • cf. Taxodioxylon
  • Holzmaden.[19]
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with the Taxodioideae family inside Cupressaceae. Resembles the modern genus Cryptomeria, but others resemble the southern genus Fitzroya, from the family Callitroideae. Large trees probably related to the coastal settings.

Protelicoxylon[41]

  • Protelicoxylon lepenneeii
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with the Taxaceae family inside Cupressaceae. Resembles the modern genus Austrotaxus.

Cupressinoxylon[20]

  • Cupressinoxylon sp.
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with Sequoioideae inside Cupressaceae. Large wood with a morphology similar to the modern genus Sequoia sempervirens. Probably related to Widdringtonites liasinus, representing the earliest representatives of the Sequoia tribe.

Circoporoxylon[42]

  • Circoporoxylon grandiporosum
  • Banz
  • Holzmaden
  • Ohmden
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with Podocarpaceae. It shares characters with modern Sciadopitys, Microcachrys, Dacrydium and Acmopyle.

Podocarpoxylon[20]

  • cf. Podocarpoxylon sp.
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil Wood

Affinities with Podocarpaceae, resembling modern genera such as Dacrycarpus, with other specimens resembling Juniperus. Includes wood more related to nearshore arbustive Conifers (columnar or low-spreading Shrubs with long, trailing branches), being the most abundant, but also medium to large arboreal conifers from nearshore forests.

Podocarpoxylon specimen

Phyllocladoxylon[20]

  • Phyllocladoxylon sp.
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Altforf[20]
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with Podocarpaceae. Similar to the modern Phyllocladus aspleniifolius.

Protophyllocladoxylon[20]

  • Protophyllocladoxylon franconicum
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with Podocarpaceae. Resembles the modern Phyllocladus hypophyllus.

Araucarioxylon[43]

  • cf. Araucarioxylon
  • Dotternhausen
  • Ohmden
  • Holzmaden.[43]
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood.

Affinities with Araucariaceae. The largest known rafting wood on the fossil record is assigned to this genus, with a length of 18 m. The rafts were populated with Crinoid colonies, and a wide variety of organisms.[43]

Araucarioxylon reconstruccion

Dadoxylon[20]

  • Dadoxylon lungriense
  • Banz
  • Holzmaden
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with Araucariaceae, resembling the modern Wollemia.

Dadoxylon sp. from Banz

Agathoxylon[20]

  • Agathoxylon agathiforme[44]
  • Agathoxylon continii
  • Banz
  • Irlbach
  • Altdorft
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with Araucariaceae, resembling the modern Agathis.

Bivalves added to an Agathoxylon driftwood

Protocupressinoxylon[45]

  • Protocupressinoxylon catenatum
  • Protocupressinoxylon liasinum
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with Cheirolepidiaceae. Includes large sized trunks up to 1.7 m tall and 115 cm wide. Large medium to large sized trees (25 m) that extent along the coastal lines of the Vindelician land. The wood from tose trunks shows insect activity, such as wood Vasps and Beetles, that had been found on the Posidonia Shale.

Protocupressinoxylon catenatum specimen from Irlbach

Simplicioxylon[46][41][47]

  • Simplicioxylon wurtembergicum[48]
  • Simplicioxylon cf.hungaricum
  • Dotternhausen
  • Altdorf
  • Irlbach
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with Cheirolepidiaceae

"Protopinaceae"[49]

  • Protopinaceae sp.
  • Holzmaden[19]
  • Ohmden
  • Banz
  • Altdorf
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Dubious genera with possible affinities with the triassic wood Woodworthia. Protopinaceae is an invalid group of mostly Paleozoic Woods.[50]

Woodworthia, example of Protopinaceae

Xenoxylon[51][52]

  • Xenoxylon cf. barberi
  • Xenoxylon ellipticum
  • Xenoxylon latiporosum
  • Xenoxylon cf. latiporosum
  • Holzmaden.[19][53]
  • Dotternhausen
  • Banz
  • Altdorf
  • Irlbach
  • Schandelah
  • Tenuicostatum
  • Falciferum
  • Bifrons

Fossil wood

Affinities with Coniferales, concretely is closer to the Podocarpaceae, Cupressaceae and in a lesser extend to the Cheirolepidiaceae. Finally can be a member of the extinct family Miroviaceae. It is the more abundant genus of wood present on the Bohemian Realm of the Posidonia Shale.

Invertebrata

Ichnofossils

Genus Species Location Material Notes Images

Thalassinoides[54]

  • Thalassinoides sp.
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Altdorf
  • Banz
  • Hemmikon
  • Maurach

Burrowing and track ichnofossils

Burrow-like ichnofossils, that can be related to Crustaceans, Annelids and Fishes.[54] The presence of this burrows changed along the different depositional layers, interpreted as result of relative magnitudes and durations of a series oxygenation events.[54] Increased Oxygen conditions eventually led to a level that permitted both the survival of larger Chondrites and Thalassinoides producing organisms, as well the depth of the Burrow-like structures.[54] The changes on the layers are detailed enough to know that oxygenation-change events duration was sufficient to allowe the migration and establishment of trace-producing organisms, establishing an "equilibrium" with bottom-water oxygen conditions.[54]

Thalassinoides found on the Posidonia Shale.

Chondrites[55]

  • Chondrites bollensis
  • Banz
  • Aichelberg
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach
  • Hemmikon
  • Dudelange-Zoufftgen

Burrowing and track ichnofossils

Burrow-like ichnofossils. Interpreted as the feeding burrow of a sediment-ingesting animal.[56] A more recent study has find that Scoloplos armiger and Heteromastus filiformis, occurring in the German Wadden Sea in the lower parts of tidal flats, make burrows that are homonymous with numerous trace fossils of the ichnogenus.[57]

Illustration of Chondrites bollensis

Phymatoderma[58]

  • Phymatoderma granulata
  • Banz
  • Altdorf
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach
  • Hemmikon

Burrowing and track ichnofossils.

Burrow-like ichnofossils. It consists commonly on a subhorizontal branching burrow system consisting of radiating tunnels filled with fecal pellets.[58] It has been interpreted as a product made by an Endobenthic deposit-feeding animal, specially a Fodinichnia, burrows produced by benthonic subsurface food-mining activity, as is proved by the tunnels and pelletal infill.[58] The study of the Fecal Pellets has revelated that the maker of this ichnogenus was an epicontinental shelf setting non-selective deposit feeder, ingesting particles on the sediment surface without selection. A mode of feeding common on aquatic Benthos, reported on modern animals such as Spionid Polychaete worms, tropical Holothurians and Spatangoid urchins.[59]

Zoophycos[58][60]

  • Zoophycos sp.
  • Banz
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach
  • Hemmikon

Burrowing and track ichnofossils.[61]

Burrow-like ichnofossils. It has been related to Echiuran annelids,[62] but also from moving and feeding polychaete worms.[63]

Example of Zoophycos fossil

Fucoides[61]

  • Fucoides bollensis
  • Banz
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach
  • Hemmikon

Burrowing and track ichnofossils.

Burrow-like ichnofossils. Considered an "Algae incertae sedis" fossil, it was recovered subdivided into 16 different groups, being in 1880 along certain ichnofossil genera, such as Cruziana with a massive nomenclatural complexity. Considered now a feeding burrow of a sediment-ingesting animal, sometimes synonymized with Chondrites.

Planolites[64]

  • Planolites montanus
  • Planolites beverleyensis
  • Banz
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach
  • Hemmikon

Burrowing and track ichnofossils.

Burrow-like ichnofossils. It is controversial, since is considered a strictly a junior synonym of Palaeophycus.[65]

Example of Planolites fossil

Palaeophycus[66]

  • Palaeophycus tubularis
  • Dotterhausen
  • Irlbach

Burrowing and track ichnofossils.

Burrow-like ichnofossils. Palaeophycus is considered related with Planolites, being a litoral fodichnia, probably from a Priapulidan.

Example of Palaeophycus fossil

Helminthopsis[66]

  • Helminthopsis isp.
  • Irlbach

Burrowing and track ichnofossils.

Burrow-like ichnofossils. It is interpreted as a grazing trail or Fodinichnia, produced at shallow depth in sediment by Polychaetes and Priapulids.[66]

Example of Helminthopsis fossil

Gyrochorte[66]

  • Gyrochorte isp.
  • Irlbach

Burrowing and track ichnofossils.

Burrow-like ichnofossils. Gyrochorte is interpreted as a result of active digging on the sediment by deposit-feeding worm-like animal, probably an Annelid or similar kinds of creatures, such as Crustaceans, Sea Urchins, nearshore fishes, etc.

Cylindrichnus[66]

  • Cylindrichnus isp.
  • Altdorf
  • Irlbach

Burrowing and track ichnofossils.

Burrow-like ichnofossils. Cylindrichnus isp. was found only on seashore-related sections, and probably represents litoral Polychaete Burrows.[67]

Asterosoma[66]

  • Asterosoma ludwigae
  • Holzmaden
  • Ohmden

Burrowing and track ichnofossils.

Burrow-like ichnofossils. Vertical or oblique complex trace fossil composed of a bunch of spindle-shaped structures and associated tubes, typical of a restricted environment (?estuarine/lagoonal).

Spongeliomorpha[66]

  • Spongeliomorpha isp.
  • Holzmaden
  • Dotterhausen
  • Irlbach

Burrows and associated traces

Burrow-like ichnofossils. Spongeliomorpha is believed to come from the domicile of Crustaceans: Anomuras (Probably Eocarcinoidea) and Decapodans (Probably Glypheidae), created as they dig in a firm, semiconsolidated substrate.

Diplocraterion[66]

  • Diplocraterion parallelum
  • Holzmaden
  • Ohmden
  • Altdorf
  • Dotternhausen
  • Irlbach

Burrows and associated traces

Burrow-like ichnofossils. Most Diplocraterion show only protrusive spreit, like the local ones, produced under predominantly erosive conditions where the organism was constantly burrowing deeper into the substrate as sediment was eroded from the top. It can be Made by Crustaceans, Annelids or other benthic fauna.[66]

Diplocraterion parallelum diagram

Anthozoa

Genus Species Location Material Notes Images

Thecocyathus[31][68]

  • Thecocyathus mitrae
  • Thecocyathus tintinabulum
  • Thecocyathus mactra
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Irlbach
  • Maurach
  • Hemmikon
  • Dudelange-Zoufftgen

Polyps

A stone cora of the family Caryophylliidae inside Hexacorallia. Related to shallow waters, this genus is the main coral found on the Posidonia Shale, resembling the modern Polycyathus muellerae. Its fossils are related with near-land facies, Coralline Islands and relatively small landmases shuch as the Bohemian Massif.

Thecocyathus mitrae from Banz

Porifera

Genus Species Location Material Notes Images

Cribrospongia[31]

  • Cribrospongia (Tremadictyon) rugatum
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Dotternhausen
  • Hemmikon

Specimens

A sea sponge of the family Cribrospongiidae inside Sceptrulophora. Found on Shallow and basinal waters, some specimens get 22 cm wide, with a funnel-like morphology. It is relatively common on nearshore strata, but generally rare.

Cribrospongia specimen

Stauroderma[31]

  • Stauroderma lochensis
  • Stauroderma sp.
  • Aichelberg
  • Dotternhausen

Specimens

A sea sponge (Glass sponge) of the family Staurodermatidae inside Hexactinellida. Found only on depth Basinal deposits, with a funnel like morphology attaining a diameter of at least 15 cm, with the exception of a large specimen of 30 cm diameter found on Dotternhausen.

Stauroderma, a 30 cm diameter specimen from Dotternhausen

Annelida

Genus Species Location Material Notes Images

Serpula?[69]

  • Serpula sp.
  • Banz
  • Aichelberg
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach
  • Hemmikon

Multiple Specimens.

A sessile, marine annelid tube worm of the family Serpulidae. Its affinities with the genus Serpula are controversial, since the genus is known mostly since Creataceus strata. Although there are other fossils assigned to the genus on same age deposits of France.[70]

Head of a modern Serpula vermicularis

Filogranula[71][72]

  • Filogranula tricristata
  • Banz
  • Aichelberg
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach
  • Hemmikon

Multiple Specimens.

A sessile, marine annelid tube worm of the family Serpulidae

Tetraserpula[73]

  • Tetraserpula quadrisulcata
  • Banz
  • Aichelberg
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach
  • Hemmikon

Multiple Specimens.

A sessile, marine annelid tube worm of the family Serpulidae.

Mucroserpula[74][75][76]

  • Mucroserpula quinquecristata
  • Banz
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Hemmikon

Multiple Specimens.

A sessile, marine annelid tube worm of the family Serpulidae. It show the characteristic features to live on soft mud ground.

Cementula[74][75][76]

  • Cementula complanata
  • Banz
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Hemmikon

Multiple Specimens.

A sessile, marine annelid tube worm of the family Serpulidae. It show the characteristic features to live on soft mud ground.

Pentaditrupa[74][75][76]

  • Pentaditrupa quinquesulcata
  • Pentaditrupa cylindracea
  • Banz
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Hemmikon

Multiple Specimens.

A sessile, marine annelid tube worm of the family Serpulidae. Pentaditrupa managed to lie freely on the mud, as shows that its tube's curvature provides stability to its position.

Propomatoceros[74][75][76]

  • Propomatoceros segmentata
  • Banz
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Hemmikon

Multiple Specimens.

A sessile, marine annelid tube worm of the family Serpulidae. Denominated "Serpula" segmentata, it lacks the two longitudinal edges characteristic on Mucroserpula.

Glomerula[77]

  • Glomerula plexus
  • Banz
  • Altdorf
  • Dotternhausen

Multiple Specimens.

A polychaete worm of the family Sabellidae.

Example of Glomerula specimen

Dictyothylakos[78]

  • Dictyothylakos pesslerae
  • Dictyothylakos sp.
  • Irlbach

Leech cocoons

Hirudinea cocoons, identified with palynological residues. The cocoons Dictyothylakos are common on flooded basin sediments, and implies not only the presence of parasitic leeches, but also the presence of large hosts nearby.

Example of Leech Cocoon

Mollusca

Brachiopoda

Genus Species Location Material Notes Images

Gibbirynchia[79][80][81][82][83][84]

  • Gibbirynchia amalthei
  • All the Formation

Thousands of Specimens.

A pennospiriferinid rhynchonellatan.[85]

Discina[79][80][81][82][83][84]

  • Discina papyracea
  • All the Formation

Thousands of Specimens.

A Discinidae rhynchonellatan. This genus was found had a planktotrophic larval stage, that adapted while growing to the local redox boundary, when this fluctuated near the sediment–water interface and oxygen availability prevailed, allowing benthic colonization. Is found on associations with Grammatodon and Pseudomytiloides.[85]

Lingula[79][80][81][82][83][84]

  • Lingula ovalis
  • All the Formation

Thousands of Specimens.

A Lingulidae rhynchonellatan. Associations of bioturbation infauna are dominated on certain sections by Palaeonucula/Lingula agrupations, developed under longer-term oxygenated conditions within the substrate and bottom waters.[85]

Rhynchonella[79][80][81][82][83][84]

  • Rhynchonella amalthei
  • Rhynchonella contraria
  • Rhynchonella batalleri
  • Rhynchonella rimosa
  • Rhynchonella bouchardi
  • Rhynchonella jurensis
  • All the Formation

Thousands of Specimens.

A Rhynchonellidae rhynchonellatan. Found assciated with Plicatula on long-term well-oxygenated conditions within the substrate and bottom waters.[85]

Bivalvia

Genus Species Location Material Notes Images

Meleagrinella[79][80][83]

  • Meleagrinella substriata
  • All the Formation[81]

Thousands of Specimens.

An oxytomid scallop.

Colony of specimens

Oxytoma[79][80]

  • Oxytoma inequivalvis
  • All the Formation

Thousands of Specimens.

An oxytomid scallop.

Chlamys[79][80][81][82][83]

  • Chlamys textoria
  • All the Formation

Thousands of Specimens.

A pectinoid scallop.

Single specimen

Eopecten[79][80][81][82][83]

  • Eopecten velatus
  • All the Formation

Thousands of Specimens.

A pectinoid scallop.

Propeamussium[79][80][81][82][83]

  • Propeamussium nonarium
  • Propeamusium pumilus
  • All the Formation

Thousands of Specimens.

A propeamussiid mud scallop.

Various specimens on the same rock

Plicatula[79][80][81][82][83]

  • Plicatula spinosa
  • All the Formation

Thousands of Specimens.

A plicatulid mud scallop.

Bositra[82][79][80][81][82][83][84][86]

  • Bositra buchii
  • All the Formation

20.000 specimens/m2

A "posidoniid" ostreoidan. It is the type fossil of the Posidonia Shale. Originally it was named "Posidonia bronni", thought to be a new genus, and the strata was denominated the Posidonia layers after it. Years later it turned out to be a junior synonym of Bositra, and thus, it was reassigned. However, the name of the layers was retained. The habitat and mode of life of Bositra has been debated for more than a century. There have been different interpretations, such as a pseudoplanktonic organism,[87] a benthic organism[88] related to open marine floor, where it was the main inhabitant of the basinal settings,[89][90] a free swimming mode of life filtering phytoplankton,[86] and a hybrid mode, where it has a life cycle with holopelagic reproduction controlled by the change on Oxygen levels,[91] and even a chemosymbiotic lifestile, related to the large crinoid rafts, being the main "Safe conduct" to evade anoxic events.[92] All the opinions along the years led to a large study in 1998, where the size/frequency distribution, the density of growth thanks to the lines related to the shell size and the position of the redox boundary by total organic carbon diagrams has revealed that Bositra probably had a benthic mode of life.[93]

Thousands of specimens in one matrix

Steinmannia[79][80][83][87]

  • Steinmannia bronni
  • Steinmannia radiata
  • All the Formation

Thousands of Specimens.

A "posidoniid" ostreoidan. Another Genera mistaken with "Posidonia bronni".

Various specimens in one matrix

Gervillella[79][80][81][82][83][84][87]

  • Gervillella lanceolata
  • All the Formation

Thousands of Specimens.

A bakevelliid mud oyster.

Liostrea[79][80][81][82][83][84][87]

  • Liostrea falcifera
  • All the Formation

Thousands of Specimens.

A gryphaeid mud oyster.

Various specimens

Gryphaea[79][80][81][82][83]

  • Gryphaea arcuata
  • All the Formation

Thousands of Specimens.

A gryphaeid mud oyster.

Various specimens

Pseudomytiloides[94][79][80][81][82][83]

  • Pseudomonotis substriata
  • Pseudomytiloides dubius
  • All the Formation

Thousands of Specimens.

An inoceramid clam. Being the second most common genera of Bivalve on the Formation, it had been object to several studies to find its ecological niche, like Bositra. Several opinions include a pseudoplanktonic-only organism, able to live in open sea,[90] or a benthonic-only organism.[89] On the 1998 evaluation with Bositra, was found that probably has a benthic early life that translated to a faculatively pseudoplanktonic mode of adult life.[93]

Single specimen

Inoceramus[94][79][80][81][82][83]

  • Inoceramus dubius
  • Inoceramus amygdaloides
  • All the Formation

Thousands of Specimens.

An inoceramid clam.

Thousands of specimens on a single rock

Nicaniella[79][80][81][82][83]

  • Nicaniella (Nicaniella) pumila
  • Nicaniella striatosulcata
  • All the Formation

Thousands of Specimens.

An astartid clam.

Solemya[95]

  • Solemya bollensis
  • Solemya voltzi
  • All the Formation

Thousands of Specimens.

A Clam, type member of the family Solemyidae inside Solemyida.

Single specimen

Palaeonucula[79][80][81][82][83]

  • Palaeonucula ungulella
  • Palaeonucula aff. hammeri
  • All the Formation

Thousands of Specimens.

A nuculidae nut clam.

Cucullaea[79][80][81][82][83][84]

  • Cucculea galathea
  • All the Formation

Thousands of Specimens.

A cucullaeid clam.

Mesomiltha[79][80][81][82][83][84]

  • Mesomiltha pumila
  • Mesomiltha pulchra
  • All the Formation

Thousands of Specimens.

A lucinid clam.

Goniomya[79][80][81][82][83][87]

  • Goniomya rhombifera
  • All the Formation

Thousands of Specimens.

A pholadomyid clam.

Grammatodon[79][80][82]

  • Grammatodon taylori
  • Grammatodon jurianus
  • All the Formation

Thousands of Specimens.

A Grammatodontinae clam. This Genus had a lecithotrophic and planktotrophic larval development.[85]

Unicardium[79][80][81][82][83][84]

  • Unicardium bollense
  • All the Formation

Thousands of Specimens.

A mactromyid clam.

Pleuromya[79][80][81][82][83]

  • Pleuromya costata
  • Pleuromya ovata
  • Pleuromya unioides
  • Pleuromya uniformis
  • All the Formation

Thousands of Specimens.

A pleuromyid clam.

Pleuromya uniformis, two specimens

Plagiostoma[79][80][81][82][83][31]

  • Plagiostoma giganteum
  • All the Formation

Thousands of Specimens.

A Limidae clam.

Plagiostoma giganteum, specimen multiview

Gibbirynchia[79][80][81][82][83][84]

  • Gibbirynchia amalthei
  • All the Formation

Thousands of Specimens.

A pennospiriferinid rhynchonellatan.

Gastropoda

Genus Species Location Material Notes Images

Coelodiscus[96][83]

  • Coelodiscus minutus
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Hemmikon
  • Dudelange-Zoufftgen

Docens of Specimens.

A Coelodiscidae sea Snail. The is the oldest known holoplanktonic gastropod, thanks to a bilateral symmetrical shells as an adaption to active swimming. Also the most common of the sea snails of the Formation, it is also one of the most varied in size terms, with some of the biggest specimens of snail from the Lower Toarcian know.[96] It has been related to large floating driftwood as one of the primary settlers.[96]

Tatediscus[97]

  • Tatediscus aratus
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Holzmaden
  • Ohmden

Docens of Specimens.

A Coelodiscidae sea Snail. Possible holoplanktonic gastropod.[96]

Procerithium[79][80][81][82][83]

  • Procerithium brandi
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen

Docens of Specimens.

A Procerithiidae sea Snail.

Toarctocera[98]

  • Toarctocera subpunctata
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen

Docens of Specimens.

An Aporrhaidae sea Snail. Among the latest described from the formation, is one of the earliest certain aporrhaidae. Characterised by large spines growing at the head of the cunch.

Cryptaulax[79][80][81][82][83]

  • Cryptaulax armatum
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Dotternhausen

Docens of Specimens.

A Cryptaulacidae sea Snail.

Neritaria[79][80][81][82][83]

  • Neritaria papilio
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Hemmikon
  • Dudelange-Zoufftgen

Docens of Specimens.

A Neritariidae sea Snail.

Single specimen

Zygopleura[79][80][81][82][83]

  • Zygopleura septemcincta
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Holzmaden
  • Ohmden
  • Hemmikon
  • Dudelange-Zoufftgen

Docens of Specimens.

A Zygopleuridae sea Snail.

Specimens

Levipleura[79][80][81][82][83]

  • Levipleura blainvillei
  • Banz
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Hemmikon

Docens of Specimens.

A Zygopleuridae sea Snail.

Trochus[79][80][81][82][83]

  • Trochus subduplicatus
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Hemmikon
  • Dudelange-Zoufftgen

Docens of Specimens.

A Trochidae sea Snail.

Specimens

Tylotrochus[79][80][97]

  • Tylotrochus subimbricatus
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Hemmikon

Docens of Specimens.

A Trochidae sea Snail. Characterised by a rhomboid scaly pattern on the teleoconch whorls.

Pleurotomaria[79][80][97]

  • Pleurotomaria debuchii
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Dudelange-Zoufftgen

Docens of Specimens.

A Pleurotomariidae sea Snail.

Specimen

Ptychomphalus[79][80][97]

  • Ptychomphalus expansus
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Dotternhausen

Docens of Specimens.

An Eotomariidae sea Snail.

Pterotrachea[99]

  • Pterotrachea liassica
  • Pterotrachea ceratophagus
  • Holzmaden
  • Ohmden
  • Aldortf
  • Dotternhausen

Various specimens.

A Pterotracheidae sea Slug. Among the oldest pelagic floating Slugs, Pterotrachea liassica had a more extended larval period than modern extant Pterotrachea coronata, because one additional whorl is Present.[99]

Modern Specimen

Cephalopoda

Genus Species Location Material Notes Images

Cenoceras[100][101][102]

  • Cenoceras intermedium
  • Cenoceras astacoides
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Irlbach
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Nautilidae Nautilidan. Includes the largest specimen of Cenoceras known, with 80 cm width.

Nautilidae shell from Banz, probably Cenoceras

Lytoceras[6][101][102]

  • Lytoceras ceratophagum
  • Lytoceras onychograptum
  • Lytoceras cornucopia
  • Lytoceras sublineatum
  • Lytoceras germaini
  • Lytoceras mucronatum
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Irlbach
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Lytoceratidae Ammonite. Lytoceras can get quite big, with nearly 50 cm in diameter.

Pachylytoceras[6][101]

  • Pachylytoceras hircinum
  • Pachylytoceras cf. hircinum
  • Pachylytoceras torulosum
  • Pachylytoceras wrighti
  • Pachylytoceras dilucidum
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg

Multiple Specimens.

A Lytoceratidae Ammonite.

Erycites[6][101]

  • Erycites labrosus
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg

Multiple Specimens.

A Hammatoceratidae Ammonite.

Hammatoceras[6][101][102]

  • Hammatoceras insigne
  • Hammatoceras compressum
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Irlbach

Multiple Specimens.

A Hammatoceratidae Ammonite.

Grammoceras[103][102]

  • Grammoceras thouarsense
  • Grammoceras orhignyi
  • Grammoceras doerntense
  • Grammoceras dispatisum
  • Grammoceras saemanni
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Irlbach
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen

Multiple Specimens.

A Hildoceratidae Ammonite.

Grammoceras specimen from Banz

Phlyseogrammoceras[103][101]

  • Phlyseogrammoceras dispansum
  • Phlyseogrammoceras cf. dispansiforme
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg

Multiple Specimens.

A Hildoceratidae Ammonite.

Pseudogrammoceras[103]

  • Pseudogrammoceras bingmanni
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg

Multiple Specimens.

A Hildoceratidae Ammonite.

Hudlestonia[101]

  • Hudlestonia serrodens
  • Hudlestonia affinis
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg

Multiple Specimens.

A Hildoceratidae Ammonite.

Catulloceras[101]

  • Catulloceras dumortieri
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg

Multiple Specimens.

A Hildoceratidae Ammonite.

Cotteswoldia[101]

  • Cotteswoldia distans
  • Cotteswoldia lotharingica
  • Cotteswoldia mactra
  • Cotteswoldia subcompta
  • Cotteswoldia fluitans
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Hildoceratidae Ammonite.

Harpoceras[104][105]

  • Harpoceras falciferum
  • Harpoceras nitescens
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Hildoceratidae Ammonite.

Harpoceras specimen

Tiltoniceras[103]

  • Tiltoniceras antiquum
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Irlbach
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Hildoceratidae Ammonite.

Tiltoniceras specimen

Hildoceras[106][107][102]

  • Hildoceras levisoni
  • Hildoceras serpentinum[106]
  • Hildoceras semipolitum
  • Hildoceras bifrons
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Irlbach
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Hildoceratidae Ammonite.

Hildoceras specimen

Mercaticeras[106]

  • Mercaticeras forte
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden

Multiple Specimens.

A Hildoceratidae Ammonite.

Mercaticeras specimen

Eleganticeras[106]

  • Eleganticeras exaratum
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Hildoceratidae Ammonite.

Pseudolioceras[106][107]

  • Pseudolioceras compactile
  • Pseudolioceras discoides
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen

Multiple Specimens.

A Hildoceratidae Ammonite.

Dactylioceras[106][108][102]

  • Dactylioceras semiannulatum
  • Dactylioceras vermis
  • Dactylioceras athleticum
  • Dactylioceras annulatum
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Irlbach
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Dactylioceratidae Ammonite.

Dactylioceras commune on Holzmaden

Collina[107][101][102]

  • Collina mucronata
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen

Multiple Specimens.

A Dactylioceratidae Ammonite. Is common on the bituminous marls (incorrectly designated as “Wilder Schiefer”) of the Altdorf High.

Catacoeloceras[107][101][102]

  • Catacoeloceras crassum
  • Catacoeloceras raquinianum
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen

Multiple Specimens.

A Dactylioceratidae Ammonite.

Coeloceras[107][101][102]

  • Coeloceras crassum
  • Coeloceras mucronatum
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

Type Coeloceratidae Ammonite.

Phylloceras[106][109]

  • Phylloceras heterophyllum
  • Phylloceras supraliasicum
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Phylloceratidae Ammonite. The largest ammonite found in the Posidonienschiefer comes from the Ohmden quarry,and belongs to a Phylloceras heterophyllum with a diameter of 87 cm.[106]

Phylloceras restoration

Haugia[106][107]

  • Haugia variabilis
  • Haugia illustris
  • Haugia jugosa
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Irlbach

Multiple Specimens.

A Phymatoceratidae Ammonite.

Denckmannia[106][107]

  • Denckmannia malagma
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau

Multiple Specimens.

A Phymatoceratidae Ammonite.

Phymatoceras[106][107]

  • Phymatoceras lilli
  • Dudelange-Zoufftgen

Multiple Specimens.

A Phymatoceratidae Ammonite.

Loligosepia[110]

  • Loligosepia aalensis
  • Banz
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen

Multiple Specimens.

A Loligosepiidae Loligosepiidan (Vampyromorpha).[111] The Loligosepiidae is believed to be ancestral to the Recent vampire squid, Vampyroteuthis infernalis.[112]

Loligosepia Holzmaden specimen

Jeletzkyteuthis[113][112]

  • Jeletzkyteuthis coriaceus
  • Holzmaden
  • Ohmden
  • Banz
  • Dotternhausen

Multiple Specimens.

A Loligosepiidae Loligosepiidan (Vampyromorpha). Related to the modern Vampyroteuthis infernalis. Gladii of Loligosepia can be distinguished from Jeletzkyteuthis by the transition lateral field/hyperbolar zone.

Geopeltis[114][112]

  • Geopeltis simplex
  • Geopeltis emarginata
  • Holzmaden
  • Ohmden
  • Banz

Multiple Specimens.

A Geopeltidae Loligosepiidan (Vampyromorpha). Related to the modern Vampyroteuthis infernalis. Gladius with weakly arcuated hyperbolar zones.

Geopeltis specimen

Parabelopeltis[115][112]

  • Parabelopeltis flexuosa
  • Holzmaden
  • Ohmden
  • Banz
  • Altdorf

Multiple Specimens.

A Geopeltidae Loligosepiidan (Vampyromorpha). Related to the modern Vampyroteuthis infernalis. It is distinguished from Geoteuthis and Loligosepia by its median rib: this rib forms a narrow ridge between two narrow grooves.

Paraplesioteuthis[115]

  • Paraplesioteuthis sagittata
  • Paraplesioteuthis hastata
  • Holzmaden
  • Ohmden
  • Banz
  • Altdorf

Multiple Specimens.

A Plesioteuthididae Prototeuthidinan (Vampyromorpha). was originally described as "Geoteuthis" sagittata.

Chitinobelus[116]

  • Chitinobelus acifer
  • Holzmaden
  • Ohmden

Multiple Specimens.

A Belemnotheutidae Belemnite. Chitinobelus is an extrange fossil, sice the rostrum was composed of aragonite with organic material, while normal Belemnites had calcite. Has been suggested this rostrum was calcitic.

Chondroteuthis[117]

  • Chondroteuthis wunnenbergi
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Belemnotheutidae Belemnite.

Simpsonibelus[107]

  • Simpsonibelus dorsalis
  • Simpsonibelus lentus[118]
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Belemnotheutidae Belemnite.

Acrocoelites[106][107]

  • Acrocoelites ilminstrensis
  • Acrocoelites pyramidalis
  • Acrocoelites glaber
  • Acrocoelites cf. riegrafi
  • Acrocoelites levidensis
  • Acrocoelites vulgaris
  • Acrocoelites tripartitus
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Megateuthididae Belemnite. Includes some of the Biggest Know Belemnites, with stimated maximum up to 4.5 m long in life, although, most specimens would have been rather smaller.

Dactyloteuthis[106][107]

  • Dactyloteuthis wrighti
  • Dactyloteuthis digitalis[118]
  • Dactyloteuthis semistriata
  • Dactyloteuthis irregularis
  • Dactyloteuthis similis
  • Dactyloteuthis incurvata
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Megateuthididae Belemnite.

Youngibelus[119][120]

  • Youngibelus giganteus
  • Youngibelus ohmdenensis[118]
  • Youngibelus simpsoni
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Megateuthididae Belemnite. Includes really large specimens

Youngibelus Reconstruction

Passaloteuthis[121]

  • Passaloteuthis paxillosa
  • Passaloteuthis bisulcata
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Passaloteuthididae Belemnite.

Passaloteuthis Holzmaden specimen

Salpingoteuthis[118]

  • Salpingoteuthis trisulcata
  • Salpingoteuthis bauhini
  • Salpingoteuthis longisulcata
  • Salpingoteuthis macra
  • Salpingoteuthis tessoniana
  • Salpingoteuthis dorsetiensis
  • Salpingoteuthis blomenhofensis
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple Specimens.

A Salpingoteuthididae Belemnite.

Clarkeiteuthis[122]

  • Clarkeiteuthis conocauda
  • Banz
  • Aichelberg
  • Dotternhausen
  • Holzmaden
  • Ohmden

Multiple Specimens.

A Diplobelidae Coleoidean. It has been found adult individuals of Clarkeiteuthis which caught small teleost fish of the species Leptolepis bronni. Further indirect evidence for the hunting behaviour comes from their body orientation in the water during life.[123]

Clarkeiteuthis Holzmaden specimen

Odontobelus[124]

  • Odontobelus tripartitus
  • Banz
  • Schandelah
  • Dotternhausen
  • Holzmaden
  • Ohmden

Various Specimens.

A Diplobelidae Coleoidean. Has been confused with Acrocoelites tripartitus, hence the species name.

Belotheutis[124]

  • Belotheutis subcostata
  • Schandelah
  • Dotternhausen
  • Holzmaden
  • Ohmden

Various Specimens.

A Diplobelidae Coleoidean. Some specimens belong to Clarkeiteuthis (=Phragmoteuthis) conocauda, but others are clearly different.

Sueviteuthis[125]

  • Sueviteuthis schlierbachensis
  • Sueviteuthis zellensis
  • Banz
  • Dotternhausen
  • Holzmaden
  • Ohmden

Multiple Specimens.

A Sueviteuthididae Coleoidean. Sueviteuthis had at least six arms with rather simple hooks, similar to the present of the genus Phragmoteuthis.

Lioteuthis[126]

  • Lioteuthis problematica
  • Holzmaden
  • Ohmden
  • Dotternhausen

Various Specimens.

Type member of the Lioteuthididae Squid family. The taxonomic position of Lioteuthis is uncertain, although the wings reaching the proximal gladius section and the smooth median field suggest affinity to the Prototeuthididae[126]

Teudopsis[127]

  • Teudopsis bollensis
  • Teudopsis subcostata
  • Teudopsis schubleri
  • Banz
  • Aichelberg
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen

Various Specimens.

A Teudopseina Squid.

Teudopsis Ohmden specimen

Geotheutis[128]

  • Geotheutis bollensis
  • Dotternhausen
  • Holzmaden
  • Ohmden

Various Specimens.

A possible primigenial Cuttlefish. Is one of the most important fossils of Cephalopods on the Posidonia Shale, due to be one of the Earliest examples of Pigments found on any species, also one of the first historically.[129] The pigments are preserved on various specimens with Eumelanin related to its ink sacs and include even phosphatized musculature.[130]

Arthropoda

Cycloidea

Genus Species Location Material Notes Images

Juracyclus[131]

  • Juracyclus posidoniae
  • Tübingen

Partial Specimens.

The First Cycloid Arthropod from the Jurassic, from the family Cycloidae inside Cycloidea.[131] Cycloids are a group of maxillopod arthropods that span between the Paleozoic until the latest Cretaceous, probably related to the crustaceans and probably detritivores.[131]

Ostracoda

Genus Species Location Material Notes Images

Infracytheropteron[132]

  • Infracytheropteron groissi
  • Infracytheropteron rarum
  • Unterstürmig
  • Dotternhausen

Cunchs

A Marine Ostracodan of the family Protostomia. The specimens of this genus are rather fargmentary and of uncertain nature.

Ogmoconcha[6][133]

  • Ogmoconcha rotunda
  • Ogmoconcha ambo
  • Ogmoconcha sp.
  • Unterstürmig

Cunchs

A marine Ostracodan, member of the family Healdiidae inside Podocopida. Rather abundant on the Toarcian profiles on Europe, this genus has a Mussel-like shape, with a very clean and round morphology.

Hermiella[134]

  • Hermiella cincta
  • Hermiella comes
  • Hermiella klingleri
  • Württenberg

Cunchs

A Marine Ostracodan of the family Healdiidae inside Podocopida. This genus is the main reported on the marine facies of the Dobbertin Clay Pit.

Ogmoconchella[134]

  • Ogmoconchella impressa
  • Ogmoconchella propinqua
  • Unterstürmig

Cunchs

A Marine Ostracodan of the family Healdiidae inside Podocopida. This genus is the main reported on the marine facies of the Dobbertin Clay Pit.

Pseudohealdia[135]

  • Pseudohealdia gruendeli
  • Pseudohealdia truncata
  • Württenberg

Cunchs

A Marine Ostracodan of the family Healdiidae inside Podocopida. The genus is rare on the layers.

Kinkelinella[136][132]

  • Kinkelinella procera
  • Kinkelinella costata
  • Württenberg

Cunchs

A Marine Ostracodan of the family Protocytheridae inside Podocopida. A genus related with Fish fossils and anoxic bottoms.

Praeschuleridea[136][132]

  • Praeschuleridea gallemannica
  • Praeschuleridea aspera
  • Württenberg

Cunchs

A Marine Ostracodan of the family Praeschuleridea inside Podocopida.

Eucytherura[132]

  • Eucytherura angulocostata
  • Württenberg

Cunchs

A Marine Ostracodan of the family Cytheruridae inside Podocopida. Is rare and the specimens found are rather incomplete.

Polycope[137]

  • Polycope tenuireticulata
  • Polycope pelta
  • Württenberg

Cunchs

A Marine Ostracodan of the family Polycopidae inside Cladocopina. Scarce but well preserved specimens.

Cytherella[138]

  • Cytherella praecadomensis
  • Württenberg

Cunchs

A Marine Ostracodan of the family Cytherellidae inside Platycopida.

Bairdia[132]

  • Bairdia ohmerti
  • Bairdia thuringica
  • Württenberg

Cunchs

A Marine Ostracodan of the family Bairdiidae inside Bairdioidea. Abundant and diverse, is found associated with Ammonite shells.

Bairdiacypris[132]

  • Bairdiacypris dorisae
  • Württenberg

Cunchs

A Marine Ostracodan of the family Bairdiidae inside Bairdioidea.

Malacostraca

Genus Species Location Material Notes Images

Orhomalus[139]

  • Orhomalus arpi
  • Neumarkt

Partial Specimens.

A hermit crab of the family Paguroidea. Hermit crabs are common of the marine layers of the lower Jurassic Europe, mostly on the Pliensbachian-Toarcian Boundary of France and Germany. Orthomalus is a small sized crab, probably inhabitant of nearshore waters, and even deltaic and beach environments, similar to modern genera.

Palaeopagurus[140]

  • Palaeopagurus sp.
  • Holzmaden

Single Specimen inside an Ammonite Shell.

An hermit crab of the family Paguridae.

Uncina[141][142]

  • Uncina posidoniae
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Banz
  • Mistlegau
  • Altdorf

Nearly complete & Partial Specimens.

An Astacidea Decapodan of the family Uncinidae. Reaching large sizes of almost half a meter (39–47 cm), Uncina Posidoniae is among the largest know Jurassic Crustaceans. Uncina posidoniae is also the largest representative of the genus Uncina.[142] This large crustacean has been found associated with Ammonite and Bivalve filled Bentos, where probably hunted different kinds of prey.[142] Its large claws would have been perfect for hunt small invertebrates and vertebrates.[142]

Uncina posidoniae specimen
The Largest complete Uncina posidoniae specimen, with 44 cm long.

Tonneleryon[143]

  • Tonneleryon schweigerti
  • Holzmaden
  • Ohmden
  • Dotternhausen

Partial Specimens and complete Specimens

A gregarious Polychelidan Lobster.

Proeryon[144][145]

  • Proeryon giganteus
  • Proeryon hauffi[146]
  • Proeryon hartmanni[147]
  • Proeryon laticaudatus[148]
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Banz
  • Mistlegau
  • Altdorf

Partial Specimens.

A Coleiidae Decapodan. The largest Coeloid from the formation, P. giganteus is a species reaching a larger size than most other polychelidans, with up to 15 cm. On the Posidonia Shale there is the most abundant variety of species from the genus, ranging from different sizes and morphologies, that indicate different habitat & feeding adaptations on the genus. Some like P. hartmani show less adaptations to hunt for small nectobenthic preys than other relatives, being abundant on Oyster-filled waters. There is a relatively abundance of the genus in deep-water settings from the Toarcian onward.[149]

Proeryon hartmanni specimen from Banz
Proeryon sp. specimen from Schandelah

Antrimpos[150][151]

  • Antrimpos noricus
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Banz

Partial Specimens.

A Penaeidae Decapodan.

Antrimpos specimen

Palaeastacus[152][153]

  • Palaeastacus sp.
  • Holzmaden
  • Ohmden
  • Dotternhausen

Partial Specimens.

An Erymidae Decapodan.

Palaeastacus specimen

Coleia[154]

  • Coleia theodorii
  • Coleia moorei
  • Coleia sinuata
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Banz

Complete Specimens. Includes specimens inside ammonites shells

An Erymidae Decapodan.

Eryma[155][156][157]

  • Eryma amalthei
  • Eryma sp.
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Banz

Complete Specimens

Type genus of the Erymidae Decapodan family. Originally, was named Glyphea amalthei,informally used by Quenstedt and housed on the Museum Naturkunde in Württemberg. A series of posterior revisions probe it was a different genus.[158]

Eryma specimen

Stenodactylina[159][160]

  • Stenodactylina liasina
  • Holzmaden
  • Ohmden
  • Dotternhausen

Complete Specimens

An Erymidae Decapodan.

Achelata[161]

  • Achelata gen. et sp. indet.
  • Holzmaden

Single complete specimen in late larval stage

The specimen reported represents the oldest fossil record of an achelatan lobster larva, and the first representative of achelatan lobsters in the Posidinia Shale. Shares similarities with the late jurassic genus Cancrinos.

California spiny lobsters are relatives of the Larval specimen from the Posidonia Shale

Stomatopoda[162]

  • Stomatopoda Indeterminate
  • Holzmaden

Single Incomplete Specimen

A Stomatopoda Malacostracan. Its affinities haven't been tested.

A female Odontodactylus Scyllarus mantis shrimp, maybe related to the Posidonia Specimen

Thoracica

Genus Species Location Material Notes Images

Toarcolepas[163]

  • Toarcolepas mutans
  • Aichelberg

Numerous disarticulated individuals, associated with fossil wood.[163]

A phospatic-shelled Cirripede of the family Eolepadidae.[163] Toarcolepas is provisionally interpreted as the oldest epiplanktonic cirripede known, and is thought to have lived attached to floating driftwood.[163]

Modern genus Lepas is the most related taxa to Toarcolepas

Arachnida

Genus Species Location Material Notes Images

Liassoscorpionides[164]

  • Liassoscorpionides schmidti
  • Niedersachsen

Single Incomplete Specimen.

The type genus of the family Liassoscorpionididae, probably related to Mesophonoidea. Being the only Jurassic scorpion known, there is no evidence that L. schmidti was aquatic (which was suggested in the past) and in the absence of further, better preserved material it should be excluded from future considerations of broad patterns of scorpion evolution.[165]

Speculative Reconstruction

Insecta

Insects are a common terrestrial animals that where proabaly drifted to the sea due to Moonsonal conditions present on the Posidonia Shale.[166]

Genus Species Location Material Notes Images

Heterophlebia[167][168]

  • Heterophlebia buckmani
  • Holzmaden
  • Schandelah
  • Braunschweig
  • Hattorf, Fallersleben
  • Bascharage

Specimens

A Dragonfly of the family Heterophlebiidae. A relative abundant genus, present on most of the pits of the Posidonia Shale, even on Holzmaden.

Plagiophlebia[168]

  • Plagiophlebia praecostarea
  • Hattorf, Fallersleben

Specimens

A Dragonfly of the family Heterophlebiidae.

Heterothemis[169]

  • Heterothemis brodiei
  • Holzmaden
  • Schandelah
  • Beienrode
  • Hattorf, Fallersleben
  • Hemmikon[170]
  • Bascharage

Specimens

A Dragonfly of the family Liassogomphidae. Second most abundant genus of Odonatan in the Formation.

Phthitogomphus[168][171]

  • Phthitogomphus angulatus
  • Niedersachsen
  • Luxguard quarry

Specimens

A Dragonfly of the family Liassogomphidae.

Proinogomphus[168]

  • Proinogomphus bodei
  • Proinogomphus kreuzerorum[172]
  • Hattorf, Fallersleben

Specimens

A Dragonfly of the family Liassogomphidae.

Ensphingophlebia[168]

  • Ensphingophlebia undulata
  • Braunschweig

Specimens

A Dragonfly of the family Sphenophlebiidae.

Mesoepiophlebia[171]

  • Mesoepiophlebia veronicae
  • Bascharage

Specimens

A Dragonfly of the family Sphenophlebiidae.

Liassostenophlebia[168]

  • Liassostenophlebia germanica
  • Rhine-Danube canal, Km 112

Specimens

A Dragonfly

Syrrhoe[168]

  • Syrrhoe commissa
  • Niedersachsen

Specimens

A Dragonfly.

Strongylogomphus[168]

  • Strongylogomphus grasselianus
  • Niedersachsen

Specimens

A Dragonfly.

Myopophlebia[168][171]

  • Myopophlebia libera
  • Beienrode
  • Bascharage

Specimens

A Dragonfly of the family Myopophlebiidae.

Paraheterophlebia[168][171]

  • Paraheterophlebia wunnenbergi
  • Paraheterophlebia marcusi
  • Niedersachsen
  • Bascharage

Specimens

A Dragonfly of the family Myopophlebiidae.

Paraplagiophlebia[171]

  • Paraplagiophlebia loneuxi
  • Bascharage

Specimens

A Dragonfly of the family Myopophlebiidae.

Necrogomphus[168][173]

  • Necrogomphus brunswigae
  • Schandelah

Specimens

A Dragonfly of the Family Liassogomphidae.

Campterophlebia[168][173]

  • Campterophlebia elegans
  • Schandelah

Specimens

A Dragonfly of the family Campterophlebiidae.The largest Early Jurassic Insect Know, with a wings size up to 20 cm.[174]

Gallodorsettia[175]

  • Gallodorsettia kronzi
  • South of Foetz

Specimens

A Dragonfly of the family Campterophlebiidae.

Henrotayia[176]

  • Henrotayia marci
  • Bascharage

Specimens

A Dragonfly of the family Henrotayiidae.

Liadoblattina[177]

  • Liadoblattina blakei
  • Holzmaden
  • Niedersachsen

Specimens

A Cockroach of the family Raphidiomimidae.

Ptyctoblattina[168]

  • Ptyctoblattina simplicior
  • Ptyctoblattina dilatata
  • Niedersachsen
  • Beienrode

Specimens

A Cockroach of the family Raphidiomimidae.

Caloblattina[168]

  • Caloblattina mathildae
  • Niedersachsen
  • Beienrode

Specimens

A Cockroach of the family Caloblattinidae.

Blattula[168]

  • Blattula langfeldti
  • Niedersachsen
  • Beienrode

Specimens

A Cockroach of the family Blattulidae.

Schesslitziella[178][179][180]

  • Schesslitziella haupti
  • Schesslitziella integra
  • Feuermühlenberg near Scheßlitz
  • Kerkhofen

Specimens

A Stick insect. One of the described insects found more near the Bohemian Massif, where probably belong most of the terrestrial invertebrate fauna.

Chresmodella

  • Chresmodella fissa
  • Hattorf, Fallersleben

Specimens

A Stick insect of the family Aerophasmidae.

Compactofulgoridium[168]

  • Compactofulgoridium spoliatum
  • Niedersachsen

Specimens

A Grasshopper.

Protogryllus[168]

  • Protogryllus formosus
  • Protogryllus hattorfensis
  • Protogryllus praeacutus
  • Protogryllus symmetricus
  • Protogryllus multoramosus
  • Protogryllus multovenosus
  • Protogryllus laceratus
  • Protogryllus foliolum
  • Protogryllus implicatus
  • Protogryllus fissus
  • Niedersachsen

Specimens

A Grasshopper of the family Protogryllidae.

Panorpidium[168]

  • Panorpidium media
  • Panorpidium geinitzi
  • Panorpidium minima
  • Niedersachsen

Specimens

A Grasshopper of the family Elcanidae.

Acridiopsis[168]

  • Acridiopsis spoliata
  • Hattorf, Fallersleben

Specimens

A Short-horned Grasshopper of the family Acrididae.

Locustopsis[168]

  • Locustopsis procera
  • Locustopsis bernstorffi
  • Locustopsis maculosa
  • Niedersachsen

Specimens

A Grasshopper of the family Locustopsidae.

Liadolocusta[168]

  • Liadolocusta ornata
  • Niedersachsen

Specimens

A Grasshopper of the family Locustopsidae.

Archijassus[168]

  • Archijassus heeri
  • Niedersachsen

Specimens

A Planthopper of the family Archijassidae.

Elasmoscelidium[168]

  • Elasmoscelidium rectemarginatum
  • Elasmoscelidium promotum
  • Elasmoscelidium boreale
  • Niedersachsen
  • Braunschweig

Specimens

A Planthopper.

Fulgoridium[168]

  • Fulgoridium mancomarginatum
  • Fulgoridium semiperspicuum
  • Fulgoridium cubitoramosum
  • Fulgoridium cuneiforme
  • Fulgoridium infuscatum
  • Fulgoridium paulodilatatum
  • Fulgoridium exiguemaculatum
  • Fulgoridium reduncum
  • Fulgoridium fallerslebense
  • Fulgoridium hattorfense
  • Fulgoridium gottingense
  • Fulgoridium tenuimaculatum
  • Fulgoridium incurvatum
  • Fulgoridium praeobtusum
  • Fulgoridium raromaculatum
  • Fulgoridium cubitofurcatum
  • Fulgoridium basilaesum
  • Fulgoridium hondelanum
  • Fulgoridium fabri
  • Fulgoridium hildesheimense
  • Fulgoridium symmetricum
  • Fulgoridium latius
  • Fulgoridium posidonicum
  • Fulgoridium silvaticum
  • Niedersachsen
  • Flechtorf near Fallersleben
  • Aselfingen

Specimens

A Planthopper of the family Fulgoridiidae.

Fulgoridulum[168]

  • Fulgoridulum egens
  • Niedersachsen

Specimens

A Planthopper of the family Fulgoridiidae.

Procerofulgoridium[168]

  • Procerofulgoridium verticillatum
  • Procerofulgoridium praefastigatum
  • Niedersachsen

Specimens

A Planthopper of the family Fulgoridiidae.

Tetrafulgoria[168]

  • Tetrafulgoria parallelogramma
  • Niedersachsen

Specimens

A Planthopper of the family Fulgoridiidae.

Metafulgoridium[168]

  • Metafulgoridium praetruncatum
  • Metafulgoridium spatulaeforme
  • Niedersachsen

Specimens

A Planthopper of the family Fulgoridiidae.

Productofulgoridium[168]

  • Productofulgoridium filiferum
  • Productofulgoridium praeacutum
  • Niedersachsen

Specimens

A Planthopper of the family Fulgoridiidae.

Margaroptilon[168]

  • Margaroptilon formosum
  • Margaroptilon cuneatum
  • Margaroptilon paucisinuatum
  • Margaroptilon detruncatum
  • Margaroptilon procerum
  • Niedersachsen

Specimens

A Planthopper of the family Fulgoridiidae.

Compactofulgoridium[168]

  • Compactofulgoridium fronterotundum
  • Compactofulgoridium obesum
  • Compactofulgoridium aries
  • Compactofulgoridium decapitatum
  • Compactofulgoridium concameratum
  • Compactofulgoridium paenintegrum
  • Niedersachsen

Specimens

A Planthopper of the family Fulgoridiidae.

Procercopis[168]

  • Procercopis lacerata
  • Procercopis abscissa
  • Procercopis wunnenbergi
  • Niedersachsen

Specimens

A Froghopper of the family Procercopidae.

Megalocoris[168]

  • Megalocoris laticlavus
  • Niedersachsen

Specimens

A Shore bug.

Eurynotis[168]

  • Eurynotis incisus
  • Niedersachsen

Specimens

A Shore bug of the family Archegocimicidae.

Somatocoris[168]

  • Somatocoris conservatus
  • Niedersachsen

Specimens

A Shore bug of the family Archegocimicidae.

Corynecoris[168]

  • Corynecoris occultatus
  • Corynecoris semigranulatus
  • Niedersachsen

Specimens

A Shore bug of the family Archegocimicidae.

Entomecoris[168]

  • Entomecoris minor
  • Niedersachsen

Specimens

A Shore bug of the family Archegocimicidae.

Ensphingocoris[168]

  • Ensphingocoris parvulus
  • Ensphingocoris praerotundatus
  • Niedersachsen

Specimens

A Shore bug of the family Archegocimicidae.

Engynabis[168]

  • Engynabis tenuis
  • Niedersachsen

Specimens

A Shore bug of the family Archegocimicidae.

Apicasia[168]

  • Apicasia inolata
  • Niedersachsen

Specimens

A basal Beetle.

Pholipheron[168]

  • Pholipheron articulatus
  • Niedersachsen

Specimens

A basal Beetle.

Grasselites[168]

  • Grasselites pusillus
  • Niedersachsen

Specimens

A basal Beetle.

Omogongylus[168]

  • Omogongylus ovatus
  • Niedersachsen

Specimens

A basal Beetle.

Sideriosemion[168]

  • Sideriosemion punctolineatum
  • Niedersachsen

Specimens

A basal Beetle.

Metanastes[168]

  • Metanastes denudatus
  • Niedersachsen

Specimens

A basal Beetle.

Diatrypamene[168]

  • Diatrypamene angulocollis
  • Niedersachsen

Specimens

A basal Beetle.

Aptilotitus[168]

  • Aptilotitus capitecarens
  • Niedersachsen

Specimens

A basal Beetle.

Rhomaleus[168]

  • Rhomaleus ornatus
  • Niedersachsen

Specimens

A basal Beetle.

Leptosolenophorus[168]

  • Leptosolenophorus brevicollis
  • Niedersachsen

Specimens

A basal Beetle.

Gastroratus[168]

  • Gastroratus dispertitus
  • Niedersachsen

Specimens

A basal Beetle.

Brachylaimon[168]

  • Brachylaimon striatus
  • Niedersachsen

Specimens

A basal Beetle.

Pleuralocista[168]

  • Pleuralocista insculpta
  • Niedersachsen

Specimens

A basal Beetle.

Mesoncus[168]

  • Mesoncus striatulus
  • Niedersachsen

Specimens

A basal Beetle.

Palaeotrachys[168]

  • Palaeotrachys laticollis
  • Niedersachsen

Specimens

A basal Beetle.

Hydroicetes[168]

  • Hydroicetes affictus
  • Schandelah

Specimens

A basal Beetle.

Scalopoides[168]

  • Scalopoides inscissus
  • Niedersachsen

Specimens

A basal Beetle.

Peridosoma[168]

  • Peridosoma praecisum
  • Niedersachsen

Specimens

A basal Beetle.

Syntomopterus[168]

  • Syntomopterus latus
  • Beienrode

Specimens

A basal Beetle.

Sphaerocantharis[168]

  • Sphaerocantharis striata
  • Sphaerocantharis defossa
  • Beienrode

Specimens

A basal Beetle.

Rhysopsalis[168]

  • Rhysopsalis distorta
  • Beienrode

Specimens

A basal Beetle.

Diplocelides[168]

  • Diplocelides minutus
  • Niedersachsen

Specimens

A basal Beetle.

Tripsalis[168]

  • Tripsalis praecisa
  • Niedersachsen

Specimens

A basal Beetle.

Trochiscites[168]

  • Trochiscites capitapertus
  • Niedersachsen

Specimens

A basal Beetle.

Prosynactus[168]

  • Prosynactus gracilis
  • Prosynactus scissus
  • Prosynactus procerus
  • Beienrode
  • Niedersachsen

Specimens

A False Ground Beetle of the family Trachypachidae.

Coreoeicos[168]

  • Coreoeicos dilatatus
  • Beienrode

Specimens

A False Ground Beetle of the family Trachypachidae.

Aposphinctus[168]

  • Aposphinctus conservatus
  • Aposphinctus striatus
  • Beienrode

Specimens

A Water Scavenger Beetle of the family Hydrophilidae.

Zetemenos[168]

  • Zetemenos sexlineatus
  • Niedersachsen

Specimens

A basal Beetle.

Amphoxyne[168]

  • Amphoxyne lineata
  • Amphoxyne minuta
  • Beienrode

Specimens

A basal Beetle.

Tolype[168]

  • Tolype rotundata
  • Beienrode

Specimens

A basal Beetle.

Amblycephalonius[168]

  • Amblycephalonius tenuistriatus
  • Niedersachsen

Specimens

A basal Beetle of the family Coptoclavidae.

Ooperioristus[168]

  • Ooperioristus applanatus
  • Niedersachsen

Specimens

A basal Beetle of the family Coptoclavidae.

Camaricopterus[168]

  • Camaricopterus ovalis
  • Niedersachsen

Specimens

A basal Beetle of the family Schizophoridae.

Megachorites[168]

  • Megachorites brevicollis
  • Niedersachsen
  • Volkmarsdorf, Braunschweig

Specimens

A Giant Beetle. It is among the largest found on all the Jurassic.

Protobittacus[168]

  • Protobittacus desacuminatus
  • Niedersachsen
  • Hattorf, Fallersleben

Specimens

A Hangingfly of the family Bittacidae.

Parabittacus[168]

  • Parabittacus lingula
  • Niedersachsen
  • Hattorf, Fallersleben

Specimens

A Hangingfly of the family Bittacidae.

Haplobittacus[168]

  • Haplobittacus parvus
  • Niedersachsen

Specimens

A Hangingfly of the family Bittacidae.

Mesobittacus[168]

  • Mesobittacus clavaeformis
  • Mesobittacus marginelaesus
  • Beienrode
  • Niedersachsen

Specimens

A Hangingfly of the family Bittacidae.

Orthophlebia[168]

  • Orthophlebia latipennisimilis
  • Orthophlebia fallerslebensis
  • Orthophlebia elongata
  • Niedersachsen
  • Hattorf, Fallersleben

Specimens

A Scorpionfly of the family Orthophlebiidae.

Parorthophlebia[168]

  • Parorthophlebia grasselensis
  • Niedersachsen
  • Hattorf, Fallersleben

Specimens

A Scorpionfly of the family Orthophlebiidae.

Mesopanorpa[168]

  • Mesopanorpa formosa
  • Beienrode

Specimens

A Scorpionfly of the family Orthophlebiidae.

Pseudopolycentropus[168]

  • Pseudopolycentropus obtusus
  • Niedersachsen
  • Hattorf, Fallersleben

Specimens

A Scorpionfly of the family Pseudopolycentropodidae.

Homoeoptychopteris[168]

  • Homoeoptychopteris incerta
  • Niedersachsen

Specimens

A Fly.

Amianta[168]

  • Amianta eurycephala
  • Niedersachsen

Specimens

A Fly.

Culiciscolex[168]

  • Culiciscolex gibberatus
  • Niedersachsen

Specimens

A Fly.

Liassonympha[168]

  • Liassonympha compacta
  • Niedersachsen

Specimens

A Fly.

Bodephora[168]

  • Bodephora arucaeformis
  • Niedersachsen

Specimens

A Fly.

Apistogrypotes[168]

  • Apistogrypotes inflexa
  • Niedersachsen

Specimens

A Fly.

Amphipromeca[168]

  • Amphipromeca acuta
  • Niedersachsen

Specimens

A Fly.

Cyrtomides[168]

  • Cyrtomides maculatus
  • Niedersachsen

Specimens

A Fly.

Sphallonymphites[168]

  • Sphallonymphites decurtatus
  • Niedersachsen

Specimens

A Fly.

Propexis[168]

  • Propexis incerta
  • Niedersachsen

Specimens

A Fly.

Archipleciomima[181]

  • Archipleciomima germanica
  • Niedersachsen

Specimens

A Fly.

Protoplecia[168]

  • Protoplecia hattorfensis
  • Niedersachsen

Specimens

A Fly of the family Protopleciidae.

Mesorhyphus[181]

  • Mesorhyphus ulrichi
  • Niedersachsen

Specimens

A Wood Gnat of the family Anisopodidae.

Metaraphidia[168]

  • Metaraphidia vahldieki
  • Niedersachsen

Specimens

A Snakefly.

Heterorhyphus[168]

  • Heterorhyphus analivarius
  • Heterorhyphus anomalus
  • Niedersachsen

Specimens

A Fly.

Amblylexis[168]

  • Amblylexis gibberata
  • Niedersachsen

Specimens

A Fly.

Ellipibodus[168]

  • Ellipibodus laesa
  • Niedersachsen

Specimens

A Fly.

Homoeoptychopteris[168]

  • Homoeoptychopteris incerta
  • Niedersachsen

Specimens

A Fly.

Protorhyphus[168]

  • Protorhyphus ovisimilis
  • Niedersachsen

Specimens

A Fly of the family Protorhyphidae.

Praemacrochile[168]

  • Praemacrochile decipiens
  • Niedersachsen

Specimens

A primitive Crane fly of the family Tanyderidae.

Nannotanyderus[179][182]

  • Nannotanyderus krzeminskii
  • Kerkhofen

Specimens

A primitive Crane fly of the family Tanyderidae.

Architipula[168]

  • Architipula bodeisimilis
  • Architipula formosa
  • Architipula basiminuta
  • Architipula robusta
  • Architipula clara
  • Architipula bodei
  • Architipula brunsvicensis
  • Architipula analiramosa
  • Architipula aequabilis
  • Architipula fragmentosa
  • Architipula veris
  • Architipula latealata
  • Niedersachsen

Specimens

A Crane fly of the family Limoniidae.

Ozotipula[168]

  • Ozotipula tarda
  • Niedersachsen

Specimens

A Crane fly of the family Limoniidae.

Haplotipula[168]

  • Haplotipula majalis
  • Haplotipula cubitoramosa
  • Niedersachsen

Specimens

A Crane fly of the family Limoniidae.

Leptotipuloides[168]

  • Leptotipuloides fastigata
  • Niedersachsen

Specimens

A Crane fly.

Mikrotipula[168]

  • Mikrotipula dixaeformis
  • Niedersachsen

Specimens

A Crane fly.

Eoptychoptera[179][183]

  • Eoptychoptera simplex
  • Niedersachsen
  • Kerkhofen

Specimens

A Phantom Crane fly of the family Ptychopteridae.

Necrotaulius[179][168][166]

  • Necrotaulius parvulus
  • Holzmaden
  • Niedersachsen
  • Kerkhofen
  • Hattorf, Fallersleben

Specimens

A Caddisfly.

Prohemerobius[168][184]

  • Prohemerobius prodromus
  • Prohemerobius septemvirgatus
  • Prohemerobius mediolatus
  • Niedersachsen

Specimens

A Lacewing of the family Prohemerobiidae.

Parhemerobius[168]

  • Parhemerobius dilatatus
  • Parhemerobius bodei
  • Niedersachsen

Specimens

A Lacewing of the family Prohemerobiidae.

Paractinophlebia[168]

  • Paractinophlebia grasselensis
  • Niedersachsen

Specimens

A Lacewing of the family Prohemerobiidae.

Panfilovia[168]

  • Panfilovia fasciata
  • Niedersachsen
  • Kerkhofen

Specimens

A Lacewing of the family Panfiloviidae. A giant Lacewing, with forewing length more than 5 cm.

Liassopsychops[168][185]

  • Liassopsychops curvata
  • Niedersachsen
  • Kerkhofen

Specimens

A Giant Lacewing, type genus of the subfamily Liassopsychopinae inside Kalligrammatidae. It is one of the oldest known representatives of the Giant pollinator lacewings. The genus Liassopsychops was previously referred to Psychopsidae. Another specimen related, Ma 14504 is regarded here as Kalligrammatidae incertae sedis. This along the occurrence of two distantly-related genera of Kalligrammatidae in the lower Toarcian is unexpected.[185]

Ophtalmogramma[185]

  • Ophtalmogramma klopschari
  • Niedersachsen

Specimens

A Giant Lacewing, genus of the subfamily Kallihemerobiidae inside Kalligrammatidae. Other of the oldest known representatives of the Giant pollinator lacewings. The Toarcian Kalligrammatidae lived in warm and dry conditions [185]

Tetanoptilon[168]

  • Tetanoptilon brunsvicense
  • Niedersachsen

Specimens

A lance Lacewing of the family Osmylidae. The largest non-Kalligrammatidae lacewing of the Jurassic, with a forewing length of 470 mm and a wingspan stimated on 11 cm.

Protoaristenymphes[186]

  • Protoaristenymphes bascharagensis
  • Bascharage

Specimens

A lance Lacewing of the family Mesochrysopidae.

Liassocicada[168][187][179]

  • Liassocicada mueckei
  • Liassocicada antecedens
  • Rhine-Danube canal, Km 112
  • Beienrode

Specimens

A Hairy Cicada of the family Tettigarctidae.

Liassotettigarcta[179]

  • Liassotettigarcta mueckei
  • Kerkhofen

Specimens

A Hairy Cicada of the family Tettigarctidae.

Xulsigia[188]

  • Xulsigia karetsa
  • Bommelscheier industrial area

Specimens

A Sternorrhynchan of the family Pincombeomorpha. It is curious for its peculiar venation on its wings. Has been proposed its own family, Xulsigiidae.

Indutionomarus[189]

  • Indutionomarus treveriorum
  • Bommelscheier industrial area

Specimens

A Coleorrhynchan of the family Progonocimicidae.

Thilopterus[168]

  • Thilopterus lampei
  • Schandelah

Specimens

A Wasp of the family Ephialtitidae.

Symphytopterus[168]

  • Symphytopterus liasinus
  • Niedersachsen

Specimens

A Wasp of the family Ephialtitidae.

Liadobracona[168]

  • Liadobracona raduhna
  • Niedersachsen

Specimens

A Wasp of the family Ephialtitidae.

Pseudoxyelocerus[190]

  • Pseudoxyelocerus bascharagensis
  • Bascharage

Specimens

A Wood Wasp of the family Xyelotomidae.

Agmatozoon[168]

  • Agmatozoon articulatum
  • Niedersachsen

Specimens

A non classified Insect.

Campeulites[168]

  • Campeulites cylindricus
  • Niedersachsen

Specimens

A non classified Insect.

Tomeferusa[168]

  • Tomeferusa abdita
  • Niedersachsen

Specimens

A non classified Insect.

Trimerocephalium[168]

  • Trimerocephalium incisum
  • Niedersachsen

Specimens

A non classified Insect.

Elasmoscolex[168]

  • Elasmoscolex hamatus
  • Niedersachsen

Specimens

A non classified Insect.

Epimetrophora[168]

  • Epimetrophora recta
  • Niedersachsen

Specimens

A non classified Insect.

Oocephalina[168]

  • Oocephalina mutilata
  • Niedersachsen

Specimens

A non classified Insect.

Platycorion[168]

  • Platycorion utroquelaesum
  • Niedersachsen

Specimens

A non classified Insect.

Protomyrmeleon[168][191]

  • Protomyrmeleon grasselensis
  • Protomyrmeleon brunonis[192]
  • Niedersachsen
  • Bascharage

Specimens

A Dicondylian, from the family Protomyrmeleontidae.

Dorniella[168]

  • Dorniella pulchra
  • Beienrode

Specimens

A basal Insect of the family Blattogryllidae.

Geinitzia[168][193]

  • Geinitzia latrunculorum
  • Geinitzia fasciata
  • Geinitzia dorni
  • Niedersachsen

Specimens

A basal Insect of the family Geinitziidae.

Adelocoris[168]

  • Adelocoris ambiguus
  • Niedersachsen

Specimens

A basal Insect of the family Pachymeridiidae.

Stiphroschema[168]

  • Stiphroschema longealatum
  • Niedersachsen

Specimens

A basal Insect of the family Pachymeridiidae.

Echinodermata

Asterozoa

Genus Species Location Material Notes Images

Ophiopholis[194]

  • Ophiopholis trispinosa
  • Holzmaden
  • Ohmden
  • Altforf
  • Banz
  • Mistelau
  • Hemmikon

Few Specimens.

An Ophiuridan of the family Ophiactidae. Very rare on the layers.

Modern Specimen

Mesophiomusium[194]

  • Mesophiomusium geisingense[195]
  • Mesophiomusium scabrum
  • Holzmaden
  • Ohmden
  • Altforf
  • Banz
  • Mistelau
  • Hemmikon

Few Specimens.

An Ophiuridan of the family Ophiolepididae. Very rare on the layers, being Mesophiomusium geisingense the most common of the two species present.

Ophiarachna[194]

  • Ophiarachna liasica
  • Holzmaden
  • Ohmden
  • Altforf
  • Banz
  • Mistelau
  • Hemmikon

Various Specimens.

An Ophiuridan of the family Ophiacanthida. Very Common, related to non anoxic water sedimentation.

Sinosura[194][196]

  • Sinosura brodiei
  • Holzmaden
  • Ohmden

Various Specimens.

An Ophiuridan of the family Aplocomidae. Very Common.

Fossil Specimen

?Ophiura[194]

  • Ophiura astonensis
  • Holzmaden
  • Ohmden

Few Specimens.

An Ophiuridan of the family Ophiuridae. Its relationships haven't been confirmed and it is based on very fragmentary remains.

Appearance

?Ophiocten[194]

  • Ophiocten seeweni
  • Holzmaden
  • Ohmden

Few Specimens.

An Ophiuridan of the family Ophiuridae. Its relationships haven't been confirmed and it is based on very fragmentary remains.

?Pentasteria[194]

  • Pentasteria sp.
  • Banz

Few Specimens.

An Asteroidean of the family Astropectinidae. It is very rare on the layers, and a few fragmentary specimens are known. Only a relatively complete specimen is known from Banz Abbey.

Pentasteria sp. from Banz

Echinoidea

Genus Species Location Material Notes Images

Cidaris[197]

  • Cidaris sp.
  • Holzmaden
  • Ohmden
  • Altforf
  • Banz
  • Irlbach
  • Hemmikon

Few Specimens.

A sea urchin of the family Cidaridae. Common on several layers. Cidaris is genus that still alive today. A bottom dweller, is commonly found associated with Belemnnite fossils, probably due to eating its carcasses.

Modern Specimen

Diademopsis[6]

  • Diademopsis crinifera
  • Diademopsis sp.
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Irlbach
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Docens of Specimens

A sea urchin of the family Pedinidae. It is the most common sea urchin in the formation, present on all the levels with specimens of various sizes. Vinculated to sea bottom sediments, before Gasteropods and Bivalves, Diademospsis was the tird major colonizer of the bottom, in between anoxic changes.

Specimens from Holzmaden

Procidaris[6]

  • Procidaris sp.
  • Holzmaden
  • Banz
  • Irlbach

Few Specimens.

A sea urchin of the family Miocidaridae

Crinoidea

Genus Species Location Material Notes Images

Praetetracrinus[198]

  • Praetetracrinus kutscheri
  • Banz
  • Altdorf
  • Holzmaden
  • Ohmden

Multiple specimens.

A Crinoidean of the family Plicatocrinidae.

Shroshaecrinus[198]

  • ?Shroshaecrinus quedlinburgensis
  • Banz
  • Altdorf
  • Holzmaden
  • Ohmden

Multiple specimens.

A Crinoidean of the family Millericrinidae.

Procomaster[194]

  • Procomaster pentadactylus
  • Banz
  • Altdorf
  • Holzmaden
  • Ohmden

Multiple specimens.

A Crinoidean of the family Isocrinida.

Isocrinus[199]

  • Isocrinus basaltiformis
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach

Multiple specimens.

Type genus of Crinoidean of the family Isocrinida.

Close view of one specimen
Single specimen

Seirocrinus[43]

  • Seirocrinus subangularis
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach

Multiple specimens. Fossilized rafts of +10 m long with complete colonies.

The Largest Know Crinoidean, from the family Pentacrinitidae. Among the tallest animals of its period, Seirocrinus is also one of the most famous fossils from the Posidonia Shale. It consists of fossils of colonies along large wood trunks, with specimens up to 14 m long, with the largest specimen reaching 26 m long,[200] what makes it among the tallest know Mesozoic organisms, one of the largest invertebrates know on the fossil record and one of the tallest know animals. It was an open ocean organism that lived in rafting woods, probably filtering food and serving as a refuge for other animals, such as ammonites.[201] The crinoids had a large colonization process, based on the status of the fossil wood found.[202][203] The large rafts where the home for a high variety of marine organisms, such as Balanoideans, Ammonites and other. It has been stimated that without the presence of modern raft wood predators (that apperared on the Bathonian) those rafts can last up to 5 years, being that the main reason the crinoids where able to attach such huge sizes. The large rafts where also proabaly essential to distribute animals along the Early Jurassic Seas.[204]

Close view of one specimen
Single specimen

Pentacrinites[205]

  • Pentacrinites fossilis
  • Pentacrinites briareus
  • Pentacrinites dichotomus
  • Pentacrinites quenstedti[206]
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Aselfingen
  • Maurach
  • Hemmikon
  • Bascharage

Multiple specimens. Fossilized rafts of +10 m long with complete colonies.

Type genus of Crinoidean from the family Pentacrinitidae. Like Seirocrinus, Pentacrinites formed colonies on rafting wood, getting a different role than bigger crinoid and appearing on the first stages of the descomposition of the rafting wood. Was a smaller genus, with specimens of no more than 1 meter long, usually measuring 40–70 cm.

Close view of one specimen
Reconstruction

Vertebrata

Fishes

Chondrichthyes

Genus Species Location Material Notes Images

Microtoxodus[207]

  • Microtoxodus gulakmani[207]
  • Schömberg

Teeth

A non classified Elasmobranch.

Synechodus[208]

  • Synechodus egertoni
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Schömberg
  • Dudelange-Zoufftgen

Teeth

Thype shark of the family Synechodontiformes.

Rhomphaiodon[207]

  • Rhomphaiodon sp.
  • Schömberg
  • Dudelange-Zoufftgen

Teeth

A shark of the family Synechodontiformes.

Protospinax[207][107]

  • Protospinax sp.
  • Schömberg
  • Dudelange-Zoufftgen

Teeth

A member of the family Protospinacidae.

Palidiplospinax[209]

  • Palidiplospinax smithwoodwardi
  • Holzmaden

Articulated vertebral column, girdles, both fin spines and clasper organ

A member of the family Palaeospinacidae.

Palaeospinax[210]

  • Palaeospinax priscus
  • Holzmaden

Anterior part of body with basicranium, palatoquadrates, Meckel's cartilage, ceratohyals, epihyals, teeth, traces of the branchial arches and the anterior finspine

Type member of the family Palaeospinacidae.

Mesiteia[207]

  • ? Mesiteia sp.
  • Schömberg
  • Dudelange-Zoufftgen

Teeth

A Carpet shark of the family Orectolobiformes.

Chiloscyllium[207]

  • ? Chiloscyllium sp.
  • Schömberg
  • Dudelange-Zoufftgen

Teeth

A Carpet shark of the family Hemiscylliidae.

Palaeobrachaelurus[207][107]

  • Palaeobrachaelurus sp.
  • Schömberg
  • Dudelange-Zoufftgen

Teeth

A Carpet shark of the family Orectolobiformes.

Ornatoscyllium[107]

  • Ornatoscyllium sp.
  • Dudelange-Zoufftgen

Teeth

A Carpet shark of the family Orectolobiformes.

Annea[107][211]

  • Annea sp.
  • Dudelange-Zoufftgen

Teeth

A Carpet shark of the family Orectolobiformes.

Paracestracion[107]

  • Paracestracion sp.
  • Dudelange-Zoufftgen

Teeth

A Bullhead Shark of the family Heterodontidae.

"Heterodontus"[107]

  • "Heterodontus" duffini
  • Dudelange-Zoufftgen

Teeth.

A Bullhead Shark of the family Heterodontidae.

Agaleus[207]

  • ? Agaleus sp.
  • Dudelange-Zoufftgen

Teeth

A shark of the family Agaleidae.

Asteracanthus[207]

  • Asteracanthus sp.
  • Dudelange
  • Holzmaden
  • Ohmden
  • Dotternhausen

Teeth

A shark of the family Acrodontidae.

Lissodus[207]

  • Lissodus guenneguesi
  • Lissodus sp.
  • Dudelange-Zoufftgen

Teeth

A shark of the family Hybodontiformes.

Crassodus[212]

  • Crassodus reifi
  • Dotternhausen

Meckelian Cartilages, Jaws, teeth, Palatoquadrates, placoid scales and desarticualted parts of the labial, hyoid and branchial skeleton.

A shark of the family Hybodontidae. The Type specimen belongs to a large hybodontid, with an estimated total length of up to 3 m.[212] It has a Meckelian Cartilage more robust than Hybodus hauffianus and an acrodontine dentition, as seen on Bdellodus.[212] Probably is related with "Hybodus" delabechei, very likely a junior synonym of Crassodus.[212]

Bdellodus[213]

  • Bdellodus bollensis
  • Holzmaden
  • Ohmden
  • Dotternhausen

Teeth.

A shark of the family Hybodontidae.

Bdellodus

Hybodus[214][215]

  • Hybodus hauffianus
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Altdorf

Complete Specimens and fragments, including Teeth, scales, jaws and Cartilage.

Type shark of the family Hybodontidae. It is the most abundant shark on the layers of the Posidonia Shale, with some of the best preserved specimens of the genus known. It was probably an open ocean hunter, with small horns over the eyes. With a size around 2 m, it was also one of the largest representatives of the Chondrichthyes on the formation.[216][217]

Hybodus Holzmaden specimen, among the best preserved of the genus, with Belemmnites inside.
Hybodus Another Holzmaden specimen

Pseudonotidanus[218]

  • Pseudonotidanus politus
  • Holzmaden
  • Ohmden
  • Dotternhausen

Fragmentary and Complete Specimens

A shark of the family Hexanchiformes.

Bathytheristes[219]

  • Bathytheristes gracilis
  • Ohmden

Partially Complete Specimen, with associated teeth

A member of Callorhynchidae inside Chimaeriformes. Similar to Callorhinchus, among the oldest known of its type.

Callorhinchus milii can be the closest relative of Bathytheristes

Acanthorhina[220]

  • Acanthorhina jaekeli
  • Holzmaden

Head and postcranial remains

A member of Myriacanthidae inside Chimaeriformes. An aberrant Chimaera with an extrange elongated nose and horns over the skull.

Acanthorhina

Metopacanthus[221]

  • Metopacanthus bollensis
  • Holzmaden
  • Ohmden

Head and several postcranial remains

A member of Myriacanthidae inside Chimaeriformes. An aberrant Chimaera with a second jaw-like structure on its head.

Toarcibatis[107][222]

  • Toarcibatis elongata
  • Toarcibatis sp.
  • Dudelange-Zoufftgen
  • Longlaville

Teeth

A Basal member of the Rajiformes of the family Archaeobatidae.

Cristabatis[107][222]

  • Cristabatis crescentiformis
  • Cristabatis sp.
  • Dudelange-Zoufftgen
  • Longlaville

Teeth

A Basal member of the Rajiformes of the family Archaeobatidae.

Doliobatis[222]

  • Doliobatis weisi
  • Dudelange-Zoufftgen

Teeth

A Basal member of the Rajiformes of the family Archaeobatidae.

Jurobatos[107][211]

  • Jurobatos cappettai
  • Jurobatos sp.
  • Dudelange-Zoufftgen

Teeth

A Basal member of the Rajiformes.

Actinopteri

Genus Species Location Material Notes Images

Holzmadenfuro[223]

  • Holzmadenfuro rebmanni
  • Holzmaden

Complete Specimen

First ganoin-scaled Ophiopsiformes (Halecomorphi) from the Posidonienschiefer. The type specimen measures 51 cm, and has elongated and serrated body scales before the dorsal fin and tiny ganoid scales after it.[223]

Ohmdenfuro[223]

  • Ohmdenfuro bodmani
  • Ohmden

Nearly complete specimen with broken skull

First ganoin-scaled Ophiopsiformes (Halecomorphi) from the Posidonienschiefer. Elongated morphology, with a length of ~39 cm, covered by smooth, masive ganoin scales.[223]

Caturus[224]

  • Caturus smithwoodwardi
  • Holzmaden
  • Würtenmberg

Various Complete and nearly complete Specimens

Type Genus of the family Caturidae inside Amiiformes

Saurorhynchus[225]

  • Saurorhynchus hauffi
  • Saurorhynchus brevirostris
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach

Multiple specimens.

The youngest representative of the family Saurichthyidae, known for its large jaws, similar to modern Belonidae.

Ptycholepis[226][227]

  • Ptycholepis bollensis
  • Ptycholepis barrati
  • Dotternhausen
  • Holzmaden
  • Ohmden

Multiple specimens.

Type member of the family Ptycholepididae inside Ptycholepiformes. It is one of the Youngest representatives of its Family.

Pholidophorus

  • Pholidophorus germanicus
  • Pholidophorus hartmanni
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach

Multiple specimens.

A member of the family Pholidophoridae. Is among the most abundant fishes on the late liassic of Europe, present on the sub-mediterranean boreal, with specimens of several sizes.

Luxembourgichthys[228]

  • Luxembourgichthys friedeni
  • Bommelscheier industrial area, Bascharage

Multiple specimens.

A member of the family Pholidophoridae.

Leptolepis[229]

  • Leptolepis jaegeri
  • Leptolepis coryphaenoides
  • Leptolepis bronni
  • Leptolepis normandica
  • Banz
  • Altdorf
  • Oedhof
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach

Multiple specimens.

A member of the family Leptolepididae. The most common member of its family, Leptolepis is commonly associated with Crustaceans and small marine invertebrates, probably main creatures on its diet. One on the most predated vertebrates on the formation, with abundance of larger fishes and reptiles with specimens associated.

Paraleptolepis[230]

  • Paraleptolepis wiedenrothi
  • SW of Braunschweig

MB. f.7612, nearly complete specimen.

A member of the family Leptolepididae. It differs from Leptolepis coryphaenoides in the presence of a few autapomorphics and also in the retention of several primitive features not present on the last one.[230] Small genus, of about 14 cm length.[230]

Euthynotus[231]

  • Euthynotus incognitus
  • Holzmaden
  • Ohmden

Multiple specimens.

A member of the family Pachycormidae.

Saurostomus[232]

  • Saurostomus esocinus
  • Banz
  • Altdorf
  • Dotternhausen
  • Holzmaden
  • Ohmden

Multiple specimens.

A member of the family Pachycormidae. Large representative of the family, reaching sizes up to 2.3 m.

Pachycormus[233][234]

  • Pachycormus macropterus
  • Pachycormus bollensis
  • Banz
  • Altdorf
  • Dotternhausen
  • Holzmaden
  • Ohmden

Multiple specimens.

Type member of the family Pachycormidae. Large representative of its family, with a size up to 1.5 m. One specimen preserved the alimentary canal, with the stomach filled by numerous hooklets that can be referred to the coleoid cephalopod Phragmoteuthis, impliying a diet of cephalopods from this genus.[235]

Sauropsis[236][237]

  • Sauropsis latus
  • Sauropsis veruinalis[224]
  • Holzmaden
  • Ohmden

Multiple specimens.

A large member of the family Pachycormidae.

Ohmdenia[238]

  • Ohmdenia multidentata
  • Ohmden

Partial specimen.

A large member of the family Pachycormidae, with up to 2.5–3 m long and an estimated weight over 200 kg.[238][239] Considered originally a junior synonym of Pachycormus, although the craneal bones suggest a new genus. Among the largest fish found on the formation it is a key fossil on the transition to large filter feeding fishes.[239] Being Coeval with another basal Pachycormiformes show the specialization of the group during the late lower jurassic.[239] Ohmdenia is the sister taxa to group of suspension-feeding Giant Middle-Late Jurassic Fishes (Including the famous Leedsichthys), showing alterations on its dental structure, with jaw indicates a diet based on soft body prey.[239] Its evolutionary significance is equiparable to that of the genus Aetiocetus for the modern Baleen whale.[239]

Dapedium[240]

  • Dapedium pholidotum[241]
  • Dapedium punctatus
  • Dapedium caelatus
  • Dapedium stollorum[242]
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach

Multiple specimens.

A deep-bodied neopterygian, Type member of the family Dapediidae. Unpublished material indicates the presence of one or even two more still undescribed species of Dapedium in the Lower Toarcian.[241]

Lepidotes[243]

  • Lepidotes elvensis
  • Lepidotes gigas
  • Banz
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Niedersachsen
  • Dotternhausen
  • Holzmaden
  • Ohmden

Multiple specimens.

A common member of the Lepisosteiformes.

Tetragonolepis[244]

  • Tetragonolepis drosera
  • Tetragonolepis semicincta
  • Banz
  • Altdorf
  • Mistelgau
  • Aichelberg
  • Schandelah
  • Dotternhausen
  • Holzmaden
  • Ohmden
  • Maurach

Multiple specimens.

A common member of the Semionotidae.

Strongylosteus[245][246]

  • Strongylosteus hindenburgi
  • Holzmaden
  • Ohmden
  • Dotternhausen

Various Specimens

A large member of the Chondrosteidae and the largest non reptilian marine vertebrate of the Posidonia shale, with a size between 3 and 4.5 m, and an estimated weight over 800 kg to 1 tonne.[245] Has been suggested as a junior Synonym of Chrondrosteus, although there haven't been any new revision about the status of the genus. It is related to modern sturgeons, but with a different kind of mouth than common species, made for hunting prey in open waters, with a strong lower jaw, similar to modern Beluga Sturgeons.[247]

Sarcopterygii

Genus Species Location Material Notes Images

Trachymetopon[248][249]

  • Trachymetopon liasicum
  • Holzmaden
  • Ohmden
  • Dotternhausen

Various Specimens with diverse degree of preservation.

A large coelacanth of the family Mawsoniidae, related with the genera Axelrodichthys, Chinlea, Diplurus and the type Mawsonia.[249] The largest specimen know from the Posidonia Shale is GPIT.OS.770 (Holotype), with a length over 1.6 m.[248] The specimen presents an ossified lung inside the abdominal cavity, and most of the body, being also one of the most complete Coelacanths of the Jurassic found.[248][249] Some recent discoveries from the Middle Jurassic show specimens of up to 3.5 m long.[250] Trachymetopon precedes the presence of the family Mawsoniidae in Europe by about 120 Ma and the northernmost occurrence of a member of the group, impliying an extensive geographical range during the Early Jurassic.[249] Due to the specimens being found on pelagic deposits suggest that probably was an open ocean swimmer.[248][249]

Trachymetopon GPIT.OS.770

Reptiles

Ichthyosauria

Inderminate specimens are known.[6]

Genus Species Location Material Notes Images

Temnodontosaurus[251][252]

  • Temnodontosaurus trigonodon
  • Temnodontosaurus burgundiae
  • Temnodontosaurus "sp. A"
  • Temnodontosaurus "sp. B"
  • Banz
  • Altdorf
  • Mistelgau
  • Schlierbach
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Dotternhausen

Multiple specimens with different degree of preservation.

Type genus of the family Temnodontosauridae. A large Macroraptorial Ichthyosaur, apex predator of its environment. It range on sizes between the 9 and the 12 m, being one of the largest know ichthyosaurs, characterized by skulls and jaws over 1 m in length, with the largest being over 1.9 m long. It has been found with fragments of young icthyosaur on his stomach.[253] Of 39 specimens of Temnodontosaurus studied, a 21% (8 Specimens) show pathologies along their body, with several injuries post traumatic, probably done by other marine reptiles.[254]

Temnodontosaurus hunting
Temnodontosaurus trigonodon specimen.

Stenopterygius[255][256][257][258]

  • Banz
  • Altdorf
  • Mistelgau
  • Schlierbach
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Dotternhausen

Multiple specimens. Some of them are among the best preserved Ichthyosaur remains known.[255][256][257][258]

Type genus of the family Stenopterygiidae. A common Toarcian Ichthyosaur, present on multiple layers. The rather exquisite level of preservation has led to know even the coloration, that exposes a clear countershading, with an upper part being more obscure than the lower, similar to modern Killer Whales, the Heaviside's dolphin or the Dall's porpoise. There is also evidence of changes in color with ontogenic chnges, going from dark juveniles to countershaded adults. The skin was flexible & Scaless, as in Dolphins.[259]

Restoration
Fossil mother with embryo

Suevoleviathan[260][261]

  • Suevoleviathan disinteger
  • Suevoleviathan integer[262]
  • Holzmaden
  • Ohmden
  • Banz
  • Dotternhausen

Multiple specimens.

Type genus of the family Suevoleviathanidae. Includes specimens up to 4 m long.

Suevoleviathan integer fossil

Hauffiopteryx[263]

  • Hauffiopteryx typicus
  • Hauffiopteryx altera[264]
  • Holzmaden
  • Ohmden
  • Dotternhausen

Complete Specimen.

Small sized Ichthyosaur, probably a member of Parvipelvia, sister group to Stenopterygius + Ophthalmosauridae.[265] A small- to mid-sized ichthyosaur, 2–3 m in length, with relatively short and slender antorbital rostrum.[264]

Hauffiopteryx typicus fossil

Eurhinosaurus[266][267][268]

  • Eurhinosaurus longirostris
  • Banz
  • Altdorf
  • Holzmaden
  • Ohmden
  • Dotternhausen

Complete and partial Specimens.

A large ichthyosaur of the family Leptonectidae with convergent evolution with modern Swordfish. Like this fishes, Eurhinosaurus is believed to be a fast swimming predator, able to hunt fish schools on same way. Large specimens of up to 6 m are known.

Complete specimen from the Posidonia Shale
Eurhinosaurus restoration

Plesiosauria

Genus Species Location Material Notes Images

Meyerasaurus[269]

  • Meyerasaurus victor
  • Dotternhausen
  • Holzmaden

Nearly complete specimen.

A Rhomaleosauridae Plesiosaur. Its detailed fossils have helped to study plesiosaur movement.[270]

Hauffiosaurus[271]

  • Hauffiosaurus zanoni
  • Dotternhausen
  • Holzmaden

Complete specimen.

A Rhomaleosauridae Plesiosaur. A moderately sized (3.4 m) Rhomaleaosaurid, ecologically adapted to fish hunt, as has been observed due to comparing the long snouted skull with that of Peloneustes, Gharial Crocodiles or Dolphins.[271]

Plesiopterys[272][196]

  • Holzmaden

Complete Specimen.

A basal Plesiosaur that has been linked with Cryptoclididae. It is one of the smallest from the Posidonia, with a complete skeleton measuring less than 2.5 m. It is considered a possible junior synonym of Seeleyosaurus[274]

Seeleyosaurus[275][276]

  • Seeleyosaurus guilelmiimperatoris
  • Holzmaden

Complete Specimens.

A Plesiosaur of the family Microcleididae. It was named originally "Plesiosaurus guilelmiimperatoris". It was a moderate‐sized plesiosauroid, measuring up to 3,5 m in length with a skull length of 170 mm.[276]

Microcleidus[277]

  • Microcleidus brachypterygius
  • Holzmaden

Several Complete Specimens.

Type member of the Plesiosaur family Microcleididae. Small Plesiosaur, with a length of less than 3 m. Possible Junior synonym of Hydrorion

Hydrorion[278][276][279]

  • Hydrorion brachypterygius
  • Holzmaden

Complete Specimen.

A Plesiosaur of the family Plesiosauridae. It is characterised by a really enlongated neck, was probably an ichthyophagous form that occurred rarely in the Posidonienschiefer fauna.

Sphenodontia

Genus Species Location Material Notes Images

Palaeopleurosaurus[280]

  • Palaeopleurosaurus posidoniae
  • Banz
  • Holzmaden
  • Ohmden
  • Dotternhausen
  • Kerkhofen

Complete specimens.

An aquatic sphenodont of the family Pleurosauridae. Palaeopleurosaurus evidences that there was a slightly skeletal specialization for an aquatic lifestyle, achieved trougth the Jurassic gradually on pleurosaurs.[280] It has similarities with other marine reptiles, such with members of Sauropterygia the presence of a defined suture between the centrum and the neural arch, along with reducted sternum.[281] Probably had a semiaquatic style of life, altrought not as adapted as Pleurosaurus, as show limited morphological evidence of adaptation to a complete aquatic lifestyle, defined by no Osteosclerosis and the lack of Pachyostosis, except for a thicker shaft region in the humerus, that is as narrow as in terrestrial rhynchocephalians, such as the terrestrial Clevosaurus.[281] Palaeopleurosaurus probably was still able to walk on land, for example for Oviposition.[281] Recent studies suggest a shorter lifespan than modern Tuatara, based on irregular spacing of growth marks.[282]

Testudinata

Genus Species Location Material Notes Images

Testudinata[283][284][285]

  • Testudinata indeterminate A
  • Testudinata indeterminate B
  • Testudinata indeterminate C
  • Altdorf, Bavaria

Various partial specimens, mostly pleurals.

Unclassified Testudine. Münster (1834) cited: " there were also rare things at the quarries of Altdorf, among other remains there where ones of a turtle on lias limestone"". The remains are not cataloged.[286]

Mesochelydia?[287]

  • Mesochelydia? indeterminate A
  • Mesochelydia? indeterminate B
  • Banz, Bavaria

Various partial specimens, mostly pleurals. Includes a left partial shell.

Assigned to Mesochelydia(?) due to resemblances for the authors to modern turtles. Unclassified and non cataloged.[286]

Perichelydia?[288]

  • Perichelydia? indeterminate A
  • Schandelah

Fragmentary Remains

Can be a more basal taxon or even not a turtle. If it is, can be the first Turtle remains found on the Posidonia Shale since the late 1800s. Now on a private collection.[288]

Crocodrylomorpha

Genus Species Location Material Notes Images

Steneosaurus[289][105]

  • Steneosaurus bollensis
  • Banz
  • Altdorf
  • Mistelgau
  • Schlierbach
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Dotternhausen

Complete specimens.

A marine crocodrylomorph with a diet probably based on fish.[290] It is the best known member of the family Teleosauriade. The Genus was declared Invalid in 2020.

Macrospondylus[291]

  • Macrospondylus bollensis
  • Banz
  • Altdorf
  • Mistelgau
  • Schlierbach
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Dotternhausen

Series of Complete specimens, including skin impressions

A Thalattosuchian of the family Teleosauridae. Was considered synonymous with Steneosaurus until in 2020 this last was recovered as invalid.[292] Includes 5 m specimens.

Macrospondylus bollensis fossil with Skin Impressions

Mystriosaurus[293][294]

  • Mystriosaurus laurillardi
  • Banz
  • Altdorf
  • Holzmaden
  • Ohmden
  • Dotternhausen

Complete specimen.

A Thalattosuchian of the family Teleosauridae. A marine crocodrylomorph with a diet probably based on fish. Was considered synonymous with Steneosaurus until recently.[295]

Platysuchus[296]

  • Platysuchus multiscrobiculatus
  • Banz
  • Altdorf
  • Schandelah
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Dotternhausen

Complete specimen.

A Thalattosuchian of the family Teleosauridae. Platysuchus was slightly more robust than its contemporaneous relatives, being probably adapted to hunt more voluminous fish.

Pelagosaurus[297][298]

  • Pelagosaurus typus
  • Banz
  • Altdorf
  • Ohmden
  • Holzmaden
  • Dotternhausen

Complete specimen.

A Thalattosuchian with a complex assignation, where can be a member of the family Teleosauridae or the basalmost Metriorhynchoidean. With a less heavy built, bigger skull and more visible marine adaptation, its diet probably based completely on fish (one fossil specimen was found with a Leptolepis in its stomach contents). Has been considered a basal Metriorhynchidae[299] or sister taxon to both Teleosauridae and Metriorhynchidae.[300]

Pholidosauridae?[289]

  • Pholidosauridae Indeterminate
  • Holzheim bei Göppingen

Partial specimen.

A possible Pholidosauridae Neosuchian. A marine crocodrylomorph, that can be a mistaken Teleosaurian as has been included on Cf. Platysuchus, although has characters that align it with Anglosuchus, that was also confused with a Teleosaurid.[289]

Goniopholididae?[105]

  • Goniopholididae Indeterminate
  • Altdorf

Frontal Snout

A Possible Goniopholid Neosuchian. Classified as "?Steneosaurus" sp., it was noted initially a morphology more similar to Goniopholis simus and specially Amphicotylus lucasii, being a possible early member of the group.[105]

Protosuchidae?[288]

  • Protosuchidae Indeterminate
  • Schandelah

Fragmentary remains

Possible member of the Protosuchia inside Crocodyliformes. If is confirmed as a member of the family it would be a terrestrial, maybe semi-acuatic crocodrylomorph. Can be a mistaken juvenile Teleosaurian.[288]

Pterosauria

Genus Species Location Material Notes Images

Campylognathoides[301][302][303]

  • Campylognathoides zitteli[303]
  • Campylognathoides liasicus[303]
  • Banz
  • Altdorf
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Dotternhausen

Several specimens with different degrees of preservation.

A Novialoidean Pterosaur, type genus of the family Campylognathoidea. Mark Witton suggests the construction of Campylognathoides' extremely robust forelimbs, with proportionally long wing fingers, could be a specialization for a fast aerial lifestyle comparable to those of Falcons and mastiff bats, being more probably an insect & vertebrate hunter and living on nearshore environments.[304]

Nearly complete Campylognathoides

"Schandelopterus"[305]

  • Novialoidea ("Schandelopterus") Indeterminate
  • Schandelah

Pelvis and several vertebrae.

A Novialoidean Pterosaur, probably a member of the family Campylognathoidea. Has been assigned to the genus Campylognathoides, although it is clearly different than any other pterosaur from the Posidonia Shale.[305] The name "Schandelopterus" is invalid and lacks any study, assigned without species to refer to the specimen on private German Fossil Groups. The pelvis indicates a laterally, slightly upwardly directed orientation of the acetabula which does not support a bird-like bipedal locomotion of this pterosaur as has been suggested on the past.[305]

Parapsicephalus[306]

  • cf. Parapsicephalus purdoni
  • Altdorf

Skull

A Rhamphorhynchinae Pterosaur. Has been assigned to the genus Dorygnathus. It has a really complete skull that can help to explain the estatus of the genus Parapsicephalus.[306]

Dorygnathus[307][308]

  • Dorygnathus banthensis[309]
  • Dorygnathus mistelgauensis?[310]
  • Banz
  • Altdorf
  • Schandelah
  • Niedersachsen
  • Holzmaden
  • Ohmden
  • Dotternhausen

Multiple specimens with different degree of preservation.[311]

A Rhamphorhynchinae Pterosaur. It is one of the best known Early Jurassic Pterosaurs.[307] Unlike Campylognthoides, Dorygnathus was an oceanic hunter, with teeth disposed to catch marine prey, such as Belemnittes and several species of fishes. Dorygnathus mistelgauensis is considered a junior synonym until more data can be recovered from the specimen, held on a private collection.[307]

Dorygnathus on terrestrial pose
Nearly complete Dorygnathus

"Ohmdenodraco"[307]

  • Rhamphorhynchinae ("Ohmdenodraco") indeterminate
  • Ohmden

Femur and a broken tibia-fibula.

A possible Rhamphorhynchinae Pterosaur. Like "Schandelopterus", "Ohmdenodraco" is an invalid name, used on private fossil groups to refer to SMNS 80439, assigned as "?Dorygnathus sp." originally, altrougth characters on the tibia and femur are distinct to any Pterosaur found on the Posidonia.[307]

Dinosauria

Possible teeth from Dinosaurs are known from the Lias Clay pit of Unterstürmig (Referred as "Various Archosaur teeth").[6]

Genus Species Location Material Notes Images

Neotheropoda[312]

  • Neotheropoda? Indeterminate
  • Schandelah
  • Cervical Vertebrae
  • Referred Teeth

An indeterminate possible Theropod dinosaur, possibly Neotheropoda. Hasn't been revised since 1984. The Cervical Vertebrae can be alternatively from a Plesiosaur, although the teeth show a clear theropod morphology.

Sauropodiformes[313]

  • Sauropodiformes Indeterminate
  • Oedhof

3 cm long, tooth-studded fragment of a lower jaw

An indeterminate possible Sauropodomorph dinosaur, possibly a member of Sauropodiformes inside Anchisauria (Resemble Yunnanosaurus teeth).[313] The Fossil was reported from the Lias Epsilon level, that on Oedhof is occupied by the Posidonia Shale.[313] It was found with abundant Plant debris and Belemnite remains.[313] Hasn't been revised since 1956

The Oedhof jaw can belong to a creature similar to Irisosaurus

Ohmdenosaurus[314]

  • Ohmdenosaurus liasicus
  • Ohmden

Tibia and astragalus

A Gravisaurian Sauropod. One of the few formally described from the Toarcian. Has been related with Vulcanodon, although more recent studies placed it as a relative of Rhoetosaurus.[315] It has been claimed to be a small sauropod with a size of 4 m, altrought the tibia measures 405 to 410 mm, leading to a 6.7 m long sauropod. Molina Pérez & Larramendi, stimated a modern size of 6.2 to 6.7 m long, with a weight of 1.3 tonnes.[316]

Synapsida

Genus Species Location Material Notes Images

Cynodontia?[317]

  • Cynodontia? Indeterminate
  • Banz

3 Teeth

The specimens were listed on several notes on the 1800s (As "mammal teeth"), although its existence has not been proved. Can be related to the Trithelodontidae or other late surviving Cynodonts by related characters, such as Irajatherium, but also to the Mammaliformes. If its presence is proved, it would be the first Synapsid found on the Posidonia Shale.

gollark: But I posted mine earlier. Four of them, even.
gollark: Except mine, naturally.
gollark: Everyone knows quicksort is better.
gollark: Unless it is, naturally.
gollark: My strategy this round was just to mess things up subtly so my entry *isn't even there to be guessed*.

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