Coptoclavidae

Coptoclavidae is an extinct family of beetles in the suborder Adephaga.[1][2] The Coptoclavidae lived from the Jurassic to the Early Cretaceous.[3] Coptoclavidae is a clade of Dytiscoidea.[4][5][6] The increase in the teleost fish population and the competition of Gyrinidae and Dytiscidae is believed to have caused their extinction.[7]

Coptoclavidae
Holcoptera giebeli
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Adephaga
Family: †Coptoclavidae
Ponomarenko, 1961
Subfamilies

Charonoscaphinae
Coptoclavinae
Coptoclaviscinae
Hispanoclavininae
Timarchopsinae
Coptoclavidae (incertae sedis)

References
  1. Tree of Life Web Project. 1995. Liadytidae. Liadytes. Version 1 January 1995 (temporary). .01.01 in The Tree of Life Web Project,
  2. Ivan Löbl & Aleš Smetana (May 2003). "1". Catalogue of Palaearctic Coleoptera: Archostemata, Myxophaga, Adephaga. 1. Apollo Books. pp. 6–8. ISBN 978-87-88757-73-6.
  3. Grimaldi, David and Michael S. Engel (16 May 2005). Evolution of the Insects. Cambridge University Press. ISBN 0-521-82149-5.
  4. Ponomarenko, A. G., Arnoldy, L., Jerikin, V. V., & Nikritin, L. M. (1992). Mesozoic Coleoptera. Suborder Adephaga, 1.
  5. Wang, B.; Ponomarenko, A. G.; Zhang, H. -Ch. (15 December 2009). "A new coptoclavid larva (Coleoptera: Adephaga: Dytiscoidea) from the Middle Jurassic of China, and its phylogenetic implication". Paleontological Journal. 43 (6): 652. doi:10.1134/S0031030109060082. ISSN 1555-6174.
  6. Bo, Wang; Ponomarenko, A. G.; Haichun, Zhang (2010). "Middle Jurassic Coptoclavidae (Insecta: Coleoptera: Dytiscoidea) from China: a Good Example of Mosaic Evolution". Acta Geologica Sinica - English Edition. 84 (4): 680–687. doi:10.1111/j.1755-6724.2010.00272.x. ISSN 1755-6724.
  7. Evans, M. E. G. (1982). "Early Evolution of the Adephaga: Some Locomotor Speculations". The Coleopterists Bulletin. 36 (4): 597–607. ISSN 0010-065X. JSTOR 4007996.


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