List of human evolution fossils
The following tables give an overview of notable finds of hominin fossils and remains relating to human evolution, beginning with the formation of the tribe Hominini (the divergence of the human and chimpanzee lineages) in the late Miocene, roughly 7 to 8 million years ago.
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As there are thousands of fossils, mostly fragmentary, often consisting of single bones or isolated teeth with complete skulls and skeletons rare, this overview is not complete, but show some of the most important findings. The fossils are arranged by approximate age as determined by radiometric dating and/or incremental dating and the species name represents current consensus; if there is no clear scientific consensus the other possible classifications are indicated.
The early fossils shown are not considered direct ancestors to Homo sapiens but are closely related to direct ancestors and are therefore important to the study of the lineage. After 1.5 million years ago (extinction of Paranthropus), all fossils shown are human (genus Homo). After 11,500 years ago (11.5 ka, beginning of the Holocene), all fossils shown are Homo sapiens (anatomically modern humans), illustrating recent divergence in the formation of modern human sub-populations.
Late Miocene (7.2–5.5 million years old)
The chimpanzee–human divergence likely took place during about 10 to 7 million years ago.[1] The list of fossils begins with Graecopithecus, dated some 7.2 million years ago, which may or may not still be ancestral to both the human and the chimpanzee lineage. For the earlier history of the human lineage, see Timeline of human evolution#Hominidae, Hominidae#Phylogeny.
Image | Name | Age (Ma) | Species | Year discovered |
Country | Discovered by | Now located at |
---|---|---|---|---|---|---|---|
El Graeco | 7.20 | Graecopithecus freybergi | 2017 (1944) | Greece, Bulgaria Site:Pyrgos Vassilissis, Azmaka |
Böhme (Tübingen), Spassov (BAS) | Met, Athens; Tübingen, Germany | |
TM 266 (Toumai) | 7.00-6.00[2] | Sahelanthropus tchadensis | 2001 | Chad Site:Djurab Desert |
Michel Brunet, Alain Beauvilain, Fanone Gongdibe, Mahamat Adoum and Ahounta Djimdoumalbaye | N'Djamena (Chad), BEAC | |
BAR 1000'00 | 6.1 - 5.7[3] | Orrorin tugenensis | 2000 | Kenya Site:Lukeino |
Martin Pickford, Kiptalam Cheboi, Dominique Gommery, Pierre Mein, Brigitte Senut, | ||
Trachilos footprints | 5.7 | hominin or hominin-like primate | 2002 | Greece | Gerard D. Gierliński | ||
ALA-VP 1/20[4] | 5.65±0.150 | Ardipithecus kadabba | 1997 | Ethiopia Site:Middle Awash |
Yohannes Haile-Selassie | ||
Pliocene (5.3–2.58 million years old)
Pleistocene
Lower Paleolithic: 2.58–0.3 million years old
Name | Age (Ma) | Species | Date discovered |
Country | Discovered by | Now located at | |
---|---|---|---|---|---|---|---|
KNM-WT 17000 (The Black Skull) |
2.50 | Paranthropus aethiopicus | 1985 | Kenya | Alan Walker | ||
BOU-VP-12/130[21] | 2.50 | Australopithecus garhi | 1997 | Ethiopia | Yohannes Haile-Selassie | ||
STS 71[22] | 2.61 -2.07 | Australopithecus africanus | 1947 | Sterkfontein, South Africa | Robert Broom and John T. Robinson | Ditsong National Museum of Natural History | |
STS 52 | 2.61 - 2.07 | Australopithecus africanus | 1947 | Sterkfontein, South Africa | Robert Broom | Ditsong National Museum of Natural History | |
UR 501 (Uraha jawbone) | 2.40±0.10 | Homo rudolfensis[23] | 1991 | Malawi | Tyson Msiska, Timothy Bromage, Friedemann Schrenk | ||
STS 5 (Mrs. Ples) (STS 14)[24] |
2.07[25] | Australopithecus africanus | 1947 | Sterkfontein, South Africa | Robert Broom | Ditsong National Museum of Natural History | |
DNH 134 (Simon)[26] | 2.04[26] | Homo erectus | 2015 | Drimolen ,Drimolen Main Quarry, South Africa | Andy Herries' team (excavated by Richard Curtis, Andy Herries, Angeline Leece; reconstructed by Jesse Martin) | University of the Witwatersrand | |
Wushan Man | 2.04 - 2[27] | Homo erectus | 1985 | Longgupo, Zhenlongping Village, Miaoyu Town of Wushan County[28] | Three Gorges Museum | ||
DNH 152 (Khethi) | 2.04-1.95 | Paranthropus robustus | 2018 | Drimolen ,Drimolen Main Quarry, South Africa | Andy Herries' team (first part found by Khethi Nkosi. later parts by Amber Jaeger, Eunice Lalunio; reconstructed by Jesse Martin & Angeline Leece) | University of the Witwatersrand | |
DNH 7 (Eurydice)[29] |
2.04 - 1.95 | Paranthropus robustus | 1994 | Drimolen, Drimolen Main Quarry, South Africa | R. Smith and André Keyser | University of the Witwatersrand | |
TM 1517[30] | unknown | Paranthropus robustus | 1938 | South Africa | Gert Terblanche | Ditsong National Museum of Natural History | |
MH1 (Karabo)[31][32] | 1.98[33] | Australopithecus sediba | 2008 | Malapa, South Africa | Lee R. Berger | University of the Witwatersrand | |
KNM-ER 1813 | 1.90 | Homo habilis | 1973 | Kenya | Kamoya Kimeu | ||
KNM-ER 1470 | 1.90 | Homo rudolfensis | 1972 | Kenya | Bernard Ngeneo[34] | ||
SK 48 | 2.25 - 1.80 | Paranthropus robustus | 1948 | Swartkrans, South Africa | Robert Broom | Ditsong National Museum of Natural History | |
SK 46[35] | 2.25 - 1.80 | Paranthropus robustus | 1949 | Swartkrans, South Africa | Robert Broom | Ditsong National Museum of Natural History | |
SK 847[36] | 2.25 -1.80 | Homo habilis | 1949 | Swartkrans, South Africa | Ditsong National Museum of Natural History | ||
OH 24 (Twiggy)[37] |
1.80 | Homo habilis | 1968 | Tanzania | Peter Nzube | ||
OH 8[38] | 1.80 | Homo habilis | 1960 | Olduvai, Tanzania | |||
D2700 (Dmanisi Skull 3) | 1.81±0.40[39] | Homo erectus | 2001 | Dmanisi, Georgia | David Lordkipanidze and Abesalom Vekua | ||
D3444 (Dmanisi Skull 4) | 1.81±0.40 | Homo erectus | 2003 | Dmanisi, Georgia | David Lordkipanidze | ||
D4500 (Dmanisi Skull 5) | 1.81±0.40 | Homo erectus | 2005 (published in 2013) | Dmanisi, Georgia | David Lordkipanidze | ||
KNM-ER 62000–62003[40] | 1.84±0.60 | Homo rudolfensis | 2012 | Koobi Fora, Kenya | Meave Leakey's team | ||
OH 5 (Zinj or nutcracker man) |
1.75 | Paranthropus boisei | 1959 | Tanzania | Mary Leakey | ||
OH 7 | 1.75 | Homo habilis | 1960 | Tanzania | Jonathan Leakey | ||
StW 53 | 1.8 - 1.6[25] | variously A. africanus, H. habilis, H. gautengensis | 1976 | Sterkfontein, South Africa | A. R. Hughes | University of the Witwatersrand | |
KNM-ER 1805 | 1.74 | Homo habilis | 1973/4 | Kenya | Paul Abell | ||
Yuanmou Man | 1.70 or 0.60–0.50 (disputed)[41] |
Homo erectus | 1965 | China | Fang Qian | ||
KNM-ER 406 | 1.70 | Paranthropus boisei | 1969 | Kenya | Richard Leakey | ||
KNM-ER 732[42] | 1.70 | Paranthropus boisei | 1970 | Kenya | Richard Leakey | ||
KNM-ER 23000[43] | 1.70 | Paranthropus boisei | 1990 | Koobi Fora, Kenya | Benson Kyongo | ||
KNM-WT 17400[44][45] | 1.70 | Paranthropus boisei | Not known[46] | Lake Turkana (West Lake Turkana) Kenya | unknown[46] | National Museums of Kenya, Nairobi (Kenia) | |
KNM-ER 3733 | 1.63±0.15[47] | Homo ergaster (a.k.a. African Homo erectus) | 1975 | Kenya | |||
KNM-WT 15000 (Turkana Boy) |
1.60 | Homo ergaster (a.k.a. African Homo erectus) | 1984 | Lake Turkana (West Lake Turkana), Kenya | Kamoya Kimeu | Kenya National Museum | |
Peninj Mandible | 1.50 | Paranthropus boisei | 1964 | Tanzania | Richard Leakey | ||
KNM-ER 992 | 1.50 | Homo ergaster (a.k.a. African Homo erectus) | 1971 | Kenya | Richard Leakey | ||
KNM-ER 3883 | 1.57±0.08 | Homo erectus | 1976 | Kenya | Richard Leakey | ||
Mojokerto 1 (Mojokerto child) |
1.43±0.10 | Homo erectus | 1936 | Indonesia | Andojo, G.H.R. von Koenigswald | ||
KGA 10-525[48][49] | 1.40 | Paranthropus boisei | 1993 | Konso-Gardula, Ethiopia | A. Amzaye | ||
OH 9 (Chellean Man)[50] |
1.40 | Homo erectus | 1960 | Olduvai, Tanzania | Louis Leakey | ||
Atapuerca Jawbone[51] | 1.20 | Homo sp. | 2008 | Spain | Eudald Carbonell | Museo de la Evolución Humana, Burgos (Spain) | |
Kocabas | 1.10[52] | Homo erectus [53] | 2002 | Turkey | M. Cihat Alçiçek | ||
Daka | 1.00 | Homo erectus | 1997 | Ethiopia | Henry Gilbert | ||
Sangiran 4 | 1.00 | Homo erectus | 1939 | Indonesia | G.H.R. von Koenigswald | ||
Sangiran 2 | 1.15±0.45 | Homo erectus | 1937 | Indonesia | G.H.R. von Koenigswald | ||
Madam Buya[54] | 1.00 | Homo heidelbergensis or Homo erectus |
1997 | Eritrea | Ernesto Abbate | National Museum of Eritrea | |
ATD6-15 and ATD6-69
(Niño de la Gran Dolina 342) |
0.900[55] | Homo antecessor or Homo erectus |
1994 | Spain | Bermúdez & Arsuaga | Museo de la Evolución Humana, Burgos (Spain) | |
Trinil 2 Pithecanthropus-1 or Java Man[56] |
0.850±0.150 | Homo erectus | 1891 | Indonesia | Eugène Dubois | Naturalis Biodiversity Center, Leiden | |
Ternifine 2-3 now Tighennif[57] | 0.70 | Homo erectus | 1954 | Algeria | C. Arambourg & B. Hoffstetter | ||
Sangiran 17[58] | 0.70 | Homo erectus | 1969 | Indonesia | S. Sartono | ||
Peking Man | 0.73±0.50[59] | Homo erectus | 1921 | China | Davidson Black | Lost/stolen | |
Bodo[60] | 0.600 | Homo heidelbergensis or Homo erectus |
1976 | Ethiopia | A. Asfaw | ||
Mauer 1 (Heidelberg Man) |
0.50 | Homo heidelbergensis | 1907 | Germany | Daniel Hartmann | Heidelberg University | |
Saldanha man[61] | 0.50 | Homo rhodesiensis | 1953 | South Africa | |||
Boxgrove Man | 0.50[62] | Homo heidelbergensis | 1994 | UK | Natural History Museum | ||
Arago 21 (Tautavel Man) |
0.45 | Homo erectus | 1971 | France | Henry de Lumley | ||
Argil Ceprano Man[63][64] |
0.450±0.050 | Homo cepranensis /Homo heidelbergensis |
1994 | Ceprano, Italy | Italo Biddittu | Servizio di antropologia, Soprintendenza ai beni culturali, Regione Lazio, Italy | |
Gawis cranium | 0.350±0.015 | Homo erectus/Homo sapiens | 2006 | Ethiopia | Asahmed Humet | ||
Skull 5 (Miguelón) | 0.400 | Homo heidelbergensis | 1992 | Spain | Bermúdez, Arsuaga & Carbonell | Museo de la Evolución Humana, Burgos (Spain) | |
Aroeira 3 | 0.40 | Homo heidelbergensis | 2014 | Portugal | João_Zilhão | Museu Nacional de Arqueologia, Lisbon | |
Salé[65][66][67] | 0.40 | Homo rhodesiensis | 1971 | Morocco | A quarry worker | ||
Swanscombe Man[68] | 0.40 | Homo Neanderthalensis | 1935, 1936, 1955 | Swanscombe | Alvan T Marston, John J Wymer and Adrian Gibson | Natural History Museum | |
Ndutu[69] | 0.40 | Homo rhodesiensis,[70][71] late Homo erectus,[72] or early Homo sapiens[73] | 1973 | Tanzania | A.A. Mturi | ||
[74] | Hexian cranial vault (PA 830)[75][76] | 0.412±0.025 | Homo erectus | 1980 | Hexian, China | ||
Steinheim Skull | 0.35 | Homo heidelbergensis | 1933 | Germany | |||
Dinaledi Chamber hominins | 0.325±0.090[77] | Homo naledi | 2013 | South Africa | Rick Hunter and Steven Tucker | University of the Witwatersrand (South Africa) | |
Middle Paleolithic: 300,000–50,000 years old
Upper Paleolithic: 50,000–11,500 years old
Holocene (11,500–5,000 years old)
Name | Age (ka) | Culture / association |
Year discovered |
Country | |
---|---|---|---|---|---|
Luzia | 11.5[139] | Paleo-Indian | 1975 | Brazil | |
[140] | Cerro Sota 2[141] | 11 | 1936 | Chile | |
"Satsurblia" | 10 | Caucasian Epipaleolithic (CHG)[136] | Georgia | ||
Kow Swamp 1 | 13–9 | 1968 | Australia | ||
Talgai Skull[142] | 10±1 | 1886 | Australia | ||
La Brea Woman | 10 | Paleo-Indian | 1914 | United States | |
Combe Capelle | 9.6 (7600 BC)[143] | European Mesolithic | 1909 | France | |
Cheddar Man | 9 (7000 BC) | British Mesolithic | 1903 | UK | |
Kennewick Man | 9 (7000 BC) | Archaic period (North America) | 1996 | United States | |
Tepexpan man | 8±3 | Paleo-Indian | 1947 | Mexico | |
Loschbour man[144] | 8 (6000 BC) | European Mesolithic (WHG) | 1935 | Luxembourg | |
Minnesota Woman | 7.9±0.1 | Paleo-Indian | 1931 | Minnesota, United States | |
Lothagam 4b (Lo 4b)[145] | 7.5±1.5[146] | 1965–1975 | Kenya | ||
Ötzi | 5.3 (3300 BC) | European Neolithic | 1991 | Ötztal Alps, Italy |
Abbreviations used in fossil catalog name
- AL – Afar Locality, Ethiopia
- ARA-VP – Aramis Vertebrate Paleontology, Ethiopia
- BAR – (Lukeino, Tugen Hills) Baringo District, Kenya
- BOU-VP – Bouri Vertebrate Paleontology, Ethiopia
- D – Dmanisi, Georgia
- ER – East (Lake) Rudolf, Kenya
- KGA – Konso-Gardula, Ethiopia
- KNM – Kenya National Museum
- KP – Kanapoi, Kenya
- LB – Liang Bua, Indonesia
- LH – Laetoli Hominid 4, Tanzania
- MH – Malapa Hominin, South Africa
- NG – Ngandong, Indonesia
- OH – Olduvai Hominid, Tanzania
- SK – Swartkrans, South Africa
- Sts, Stw – Sterkfontein, South Africa
- TM – Transvaal Museum, South Africa
- TM – Toros-Menalla, Chad
- WT – West (Lake) Turkana, Kenya
See also
Further reading
- Gibbons, Ann. The First Human: The Race to Discover our Earliest Ancestor. Anchor Books (2007). ISBN 978-1-4000-7696-3
- Hartwig, Walter Carl (2004) [2002]. Hartwig, Walter (ed.). The Primate Fossil Record. Cambridge University Press. Bibcode:2002prfr.book.....H. ISBN 978-0-521-08141-2..
- Johanson, Donald & Wong, Kate. Lucy's Legacy: The Quest for Human Origins. Three Rivers Press (2009). ISBN 978-0-307-39640-2
- Jones, Steve; Martin, Robert D.; Pilbeam, David R, eds. (1994). The Cambridge Encyclopedia of Human evolution. Cambridge University Press. ISBN 978-0-521-46786-5. (Note: this book contains very useful, information dense chapters on primate evolution in general, and human evolution in particular, including fossil history).
- Leakey, Richard & Lewin, Roger. Origins Reconsidered: In Search of What Makes us Human. Little, Brown and Company (1992). ISBN 0-316-90298-5
- Lewin, Roger. Bones of Contention: Controversies in the Search for Human Origins. Penguin Books (1987). ISBN 0-14-022638-9
- Morwood, Mike & van Oosterzee, Penny. A New Human: The Startling Discovery and Strange Story of the 'Hobbits' of Flores, Indonesia. Smithsonian Books (2007). ISBN 978-0-06-089908-0
- Oppenheimer, Stephen. Out of Eden: The Peopling of the World. Constable (2003). ISBN 1-84119-697-5
- Roberts, Alice. The Incredible Human Journey: The Story of how we Colonised the Planet. Bloomsbury (2009). ISBN 978-0-7475-9839-8
- Shreeve, James. The Neanderthal Enigma: Solving the Mystery of Modern Human Origins. Viking (1996). ISBN 0-670-86638-5
- Stringer, Chris. The Origin of Our Species. Allen Lane (2011). ISBN 978-1-84614-140-9
- Stringer, Chris & Andrews, Peter. The Complete World of Human Evolution. Thames & Hudson (2005). ISBN 0-500-05132-1
- Stringer, Chris & McKie, Robin. African Exodus: The Origins of Modern Humanity. Jonathan Cape (1996). ISBN 0-224-03771-4
- van Oosterzee, Penny. The Story of Peking Man. Allen & Unwin (1999). ISBN 1-86508-632-0
- Walker, Allan & Shipman, Pat. The Wisdom of the Bones: In Search of Human Origins. Weidenfeld & Nicolson (1996). ISBN 0-297-81670-5
- Wade, Nicholas. Before the Dawn: Recovering the Lost History of our Ancestors. Penguin Press (2006). ISBN 978-0-7156-3658-9
- Weiss, M.L.; Mann, A.E. (1985). 'Human Biology and Behaviour: An anthropological perspective (4th ed.). Boston: Little Brown. ISBN 978-0-673-39013-4. (Note: this book contains very accessible descriptions of human and non-human primates, their evolution, and fossil history).
- Wells, Spencer (2004). The Journey of Man : A Genetic Odyssey. New York: Random House Trade Paperbacks. ISBN 978-0-8129-7146-0.
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External links
- Interactive map of primate fossil finds around the world
- Informative lecture on Australopithecines
- The Age of Homo sapiens – Interactive Map of Human Evolution Fossils
- Human Timeline (Interactive) – Smithsonian, National Museum of Natural History (August 2016).