2018 in arthropod paleontology

This list of fossil arthropods described in 2018 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that were described during the year 2018, as well as other significant discoveries and events related to arthropod paleontology that are scheduled to occur in the year 2018.

List of years in arthropod paleontology
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In paleontology 2015 2016 2017 2018 2019 2020 2021
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General research

• A study on extant arthropods (mainly insects and spiders) living around the resinous tree Hymenaea verrucosa in the lowland coastal forest of Madagascar and trapped by the resin produced by this tree species, and on their implications for inferring whether amber records the true past biodiversity of the entire forest, is published by Solórzano Kraemer et al. (2018).[1]
• A study on the early evolution of stem and crown-arthropods as indicated by Ediacaran and Cambrian body and trace fossils is published by Daley et al. (2018).[2]
• A study on the evolution of ecdysozoan vision, focusing on the evolution of arthropod multi-opsin vision, as indicated by molecular data and data from fossil record, is published by Fleming et al. (2018).[3]
• Grimaldi et al. (2018) report biological inclusions (fungi, plants, arachnids and insects) in amber from the Paleogene Chickaloon Formation of Alaska, representing the northernmost deposit of fossiliferous amber from the Cenozoic.[4]
• A mantis lacewing larva found in association with two spiders, attached to one of the anterior walking legs of one of the spiders, is described from the Cretaceous amber from Myanmar by Haug, Müller & Haug (2018), who interpret this finding as evidence of palaeo-parasitism.[5]

Arachnids

Research

• Review of arachnids known from the Bitterfeld amber (Germany) and a study on their implications for inferring the age of this deposit is published by Dunlop et al. (2018).[6]
• The first fossil representative of the parasitic mite family Pterygosomatidae, assigned to the genus Pimeliaphilus and similar in morphology to the extant species parasitic on cockroaches, is described from the Cretaceous (Albian) Archingeay amber (France) by Sidorchuk & Khaustov (2018).[7]
• Fossil mite "Sejus" bdelloides, originally interpret as a member of Mesostigmata, is reinterpreted as a member of Prostigmata referrable to the group Anystina by Dunlop, Walter & Kontschán (2018).[8]
• A study on the body size of fossil mites is published by Sidorchuk (2018).[9]
• A hard tick wrapped in spider silk is described from the Cretaceous Myanmar amber by Dunlop et al. (2018).[10]
• A male specimen of a tick belonging to the genus Ornithodoros and the subgenus (Alectorobius) is described from the Dominican amber by Estrada-Peña & de la Fuente (2018),[11] who also provide illustrated interpretations intended to support the validity of the identity of the fossils reported by de la Fuente (2003).[12]
• A pseudoscorpion attached to barbules of a contour feather, possibly documenting a phoretic association between pseudoscorpions and Mesozoic birds, is described from the Cretaceous amber from Myanmar by Xing, McKellar & Gao (2018).[13]
• A study on the phylogenetic history of spiders, based on molecular data and re-examination of the global fossil record, as well as on the ancestral predatory strategies among different groups of spiders and some of the possible drivers of spider diversification during the Early Cretaceous greenhouse, is published by Shao & Li (2018).[14]
• A study on the burrows produced by wolf spiders belonging to the genus Pavocosa, aiming to identify ichnological signatures which may facilitate identification of wolf spider burrows in the fossil record, is published by Mendoza Belmontes, Melchor & Piacentini (2018).[15]
• Fossil crevice weaver Misionella didicostae from Dominican amber is transferred to the genus Antilloides by Magalhaes (2018).[16]
• Evidence of a fossilized cobweb, and a possible case of predation of a theridiid spider on a dolichopodid fly, are reported from the Mexican amber by García-Villafuerte (2018).[17]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Alteraraneus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae. The type species is A. gracilipes.
Alteruloborus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Deinopoidea and the new family Alteruloboridae. The type species is A. araneoides.
Burerythrites[19]	Gen. et sp. nov	Valid	Konikiewicz & Mąkol	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A mite belonging to the family Erythraeidae. Genus includes new species B. pankowskii.
Burfessonia[19]	Gen. et sp. nov	Valid	Konikiewicz & Mąkol	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A mite belonging to the family Smarididae. Genus includes new species B. maryae.
Burmaforceps[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae. The type species is B. amputatus.
Burmascutum brevis[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Burmascutidae.
Burmasuccinus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Uloboridae. The type species is B. bulla.
Burmatheridion[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Theridiidae. The type species is B. sinespinae.
Burphanolophus[19]	Gen. et sp. nov	Valid	Konikiewicz & Mąkol	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A mite belonging to the family Erythraeidae. Genus includes new species B. joergwunderlichi.
Chaerilobuthus knodelorum[20]	Sp. nov	Valid	Lourenço	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A scorpion belonging to the family Chaerilobuthidae.
Chimerarachne[21][22]	Gen. et sp. nov	Valid	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	An arachnid closely related to spiders and to the extinct group Uraraneida. Genus includes new species C. yingi.
Converszarqaraneus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae. The type species is C. annulipedes.
Cornicaraneus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae. The type species is C. scutatus.
Crassitibia baculum[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae.
Cretaceoushormiops[23]	Gen. et sp. nov	Valid	Lourenço	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A scorpion belonging to the family Protoischnuridae. Genus includes new species C. knodeli.
Cretamysmena[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Cretamysmenidae. The type species is C. fontana.
Eothrombium fortesambiense[24]	Sp. nov	Valid	Mąkol, Konikiewicz & Klug	Eocene	Baltic amber	Europe (Baltic Sea region)	A mite belonging to the group Prostigmata and to the family Tanaupodidae.
Eotibiaapophysis[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Uloboridae. The type species is E. reliquus.
Frateruloborus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Deinopoidea and the new family Frateruloboridae. The type species is F. bulbosus.
Galianora marcoi[25]	Sp. nov	Valid	García-Villafuerte	Miocene (Aquitanian)	Mexican amber	Mexico	A jumping spider, a species of Galianora
Haemaphysalis (Alloceraea) cretacea[26]	Sp. nov	Valid	Chitimia-Dobler, Pfeffer & Dunlop	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A tick, a species of Haemaphysalis.
Immensmaris[27]	Gen. et sp. nov	Valid	Dunlop, Frahnert & Mąkol	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A mite belonging to the family Smarididae. The type species is I. chewbaccei.	Immensmaris chewbaccei
Kachin serratus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Uloboridae.
Leviunguis altus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis anulus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis anulusoides[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis bruckschoides[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis erectus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis glomulus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis glomus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis graciliembolus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis gradus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis porrigens[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis pseudobruckschi[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Leviunguis quadratus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the new family Leviunguidae.
Microproxiaraneus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae. The type species is M. annulatus.
Miomastigoproctus[28]	Gen. et sp. nov	Valid	Lourenço	Miocene	Mexican amber	Mexico	A member of Thelyphonida belonging to the family Thelyphonidae. Genus includes new species M. knodeli.
Nothrotrombidium myanmarum[19]	Sp. nov	Valid	Konikiewicz & Mąkol	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A mite belonging to the family Trombellidae.
Orchestina sakhalinensis[29]	Sp. nov	Valid	Marusik, Perkovsky & Eskov	Middle Eocene	Sakhalinian amber	Russia	A species of Orchestina.
Palaeoburmesebuthus knodeli[30]	Sp. nov	Valid	Lourenço	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A scorpion belonging to the family Palaeoburmesebuthidae.
Palaeoleptoneta nils[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Leptonetidae.
Palaeoleptoneta thilo[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Leptonetidae.
Paramiagrammopes pusillus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Uloboridae.
Parvibulbus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Pholcochyroceridae. The type species is P. incompletus.
Paurospina[18]	Gen. et 3 sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae. The type species is P. curvata; genus also includes P. fortis and P. paulocurvata.
Pholcochyrocer calidum[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Pholcochyroceridae.
Pholcochyrocer vermiculus[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Pholcochyroceridae.
Planibulbus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Uloboridae. The type species is P. longisoma.
Praeteraneoides[18]	Gen. et 3 sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Leptonetoidea and the new family Protoaraneoididae. The type species is P. bifurcatum; genus also includes P. bipartitum and P. leni.
Proaraneoides[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Leptonetoidea and the new family Protoaraneoididae. The type species is P. cribellatum.
Propolyssenia[24]	Gen. et sp. nov	Valid	Mąkol, Konikiewicz & Klug	Eocene	Baltic amber	Europe (Baltic Sea region)	A mite belonging to the group Prostigmata and to the family Tanaupodidae. Genus includes new species P. wohltmanni.
Protoaraneoides[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Leptonetoidea and the new family Protoaraneoididae. The type species is P. longispina.
Protohylomysobia[31]	Gen. et sp. nov	Valid	Sidorchuk & Bochkov in Sidorchuk et al.	Eocene	Baltic amber	Russia	A mite belonging to the family Myobiidae. Genus includes new species P. erinaceophilus.
Proxiaraneus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae. The type species is P. rarus.
Ramozarqaraneus[18]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae. The type species is R. pauxillus.
Spinicymbium[18]	Gen. et sp. et comb. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and to the family Zarqaraneidae. The type species is S. curvimetatarsus; genus also includes "Hypotheridiosoma" falcata Wunderlich (2015).
Spinipalpitibia hirsuta[18]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the group Leptonetoidea and the new family Protoaraneoididae.
Tuckerella weiterschani[32]	Sp. nov	Valid	Sidorchuk & Khaustov	Eocene	Baltic amber	Europe (Baltic Sea coast)	A peacock mite.
Weygoldtiella[33]	Gen. et sp. nov	Valid	Harvey et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A pseudoscorpion belonging to the family Chthoniidae. Genus includes new species W. plausus.
Weygoldtina[34]	Gen. et comb. nov	Valid	Dunlop	Late Carboniferous	British Middle Coal Measures Mazon Creek fossil beds	United Kingdom  United States	A whip spider belonging to the group Paleoamblypygi and the new family Weygoldtinidae. The type species is "Graeophonus" scudderi Pocock (1911); genus also includes "Graeophonus" anglicus Pocock (1911).

Crustaceans

Research

• Redescription of the Ordovician malacostracan Wuningia multisegmenlata is published by Lin (2018).[35]
• A member of the erymid genus Enoploclytia is described from the Oxfordian deposits of Cricquebœuf (Normandy, France) by Devillez, Charbonnier & Pezy (2018), representing the first Jurassic occurrence of the genus reported so far.[36]
• Redescription of Palaeopalaemon newberryi and a study on the phylogenetic relationships and life habits of this species is published by Jones et al. (2018).[37]
• Description of the morphology of Tetrachela raiblana and a study on its implication for understanding of the homologies of carapace grooves between polychelidans and other decapods is published by Audo, Hyžný & Charbonnier (2018).[38]
• A large-sized (12.9 mm maximum length) right valve of a marine ostracod is described from the Cretaceous amber from Myanmar by Xing et al. (2018).[39]
• A study evaluating how sexual selection related to species extinction in fossil cytheroid ostracods is published by Fernandes Martins et al. (2018), who report that species with more pronounced sexual dimorphism had higher estimated extinction rates.[40]
• An ostracod fauna including four species belonging to the suborder Darwinulocopina is described from the Lower Jurassic (Hettangian) Whitmore Point Member of the Moenave Formation (Arizona and Utah, United States) by Antonietto et al. (2018), potentially representing the last episode of darwinulocopine dominance in nonmarine environments before the Late Jurassic diversification of the cypridocopine/cytherocopine modern ostracods.[41]
• Oxygen-isotope analysis of a whale barnacle specimen collected from early Pleistocene deposits of Apulia (Italy) is published by Collareta et al. (2018), who interpret their findings as indicating that the barnacle lived on a cetacean that seasonally migrated towards high-latitude areas outside the Mediterranean.[42]

New taxa

Malacostracans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Abyssophthalmus adinae[43]	Sp. nov	Valid	Schweitzer et al.	Late Jurassic		Romania	A crab belonging to the family Longodromitidae.
Aethra stalennyii[44]	Sp. nov	Valid	Ossó	Miocene (Serravallian)		Ukraine	A species of Aethra.
Amazighopsis[45]	Gen. et sp. nov	Valid	Garassino & Pasini	Late Cretaceous (Cenomanian-Turonian)	Kem Kem Beds	Morocco	A member of Astacidea of uncertain phylogenetic placement, assigned to the new family Amazighopsidae. The type species is A. cretacica.
Anapagurus muelleri[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A hermit crab belonging to the family Paguridae.
Archeostenoniscus[47]	Gen. et 2 sp. nov	Valid	Broly in Broly, Serrano-Sánchez & Vega	Early Miocene	La Quinta Formation (Mexican amber)	Mexico	A woodlouse belonging to the family Stenoniscidae. The type species is A. robustus; genus also includes A. mexicanus.
Armadilloniscus miocaenicus[47]	Sp. nov	Valid	Broly in Broly, Serrano-Sánchez & Vega	Early Miocene	La Quinta Formation (Mexican amber)	Mexico	A woodlouse, a species of Armadilloniscus.
Asthenognathus sakumotoi[48]	Sp. nov	Valid	Karasawa	Miocene	Korematsu Formation Lower Sandstone Member of the Bihoku Group	Japan	A crab belonging to the family Varunidae.
Axianassa petrea[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Axianassidae.
Balsscallichirus sangermani[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Callianassidae.
Blaculla felthausetsauteri[49]	Sp. nov	Valid	Winkler	Late Jurassic (Tithonian)	Solnhofen Limestone	Germany	A caridean shrimp.
Bryocarpilius stratensis[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab belonging to the family Carpiliidae.
Cabrillia[50]	Gen. et sp. nov	Valid	Nyborg, Pasini & Garassino	Pliocene-Pleistocene	San Diego Formation	United States	A crab belonging to the family Leucosiidae. Genus includes new species C. hendyi.
Calliaxina elegans[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Callianassidae.
Callichirus ornatus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A species of Callichirus.
Carinocalappa[46]	Gen. et sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab, a member of the family Calappidae. The type species is C. lineamenta.
Cenomanocarcinus lenzenwegeri[51]	Sp. nov	Valid	Hyžný et al.	Late Cretaceous (Santonian)		Austria	A crab belonging to the group Palaeocorystoidea.
Ceratiocaris harpago[52]	Sp. nov	Valid	Poschmann, Bergmann & Kühl	Early Devonian	Hunsrück Slate	Germany
Cirolana centinelensis[53]	Sp. nov	Valid	Maguire in Maguire et al.	Miocene (Burdigalian)	Estancia 25 de Mayo Formation	Argentina	An isopod, a species of Cirolana.
Cirolana titanophila[54]	Sp. nov	Valid	Robin et al.	Eocene (Ypresian)	Monte Bolca	Italy	An isopod, a species of Cirolana found in association with electric rays belonging to the genus Titanonarke.
Daciapagurus[55]	Gen. et sp. nov	Valid	Frantescu et al.	Late Jurassic		Romania	A hermit crab belonging to the family Schobertellidae. Genus includes new species D. minusculus.
Daira iugata[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Late Eocene		Italy	A crab belonging to the family Dairidae.
Daira pseudovulgaris[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab belonging to the family Dairidae.
Daranyia tricristata[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab, a member of Grapsoidea of uncertain phylogenetic placement.
Dinochelus radwanskii[56]	Sp. nov	Valid	Fraaije et al.	Late Cretaceous (Maastrichtian)		Poland	A species of Dinochelus.
Diogenes denticulatus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A hermit crab belonging to the family Diogenidae, a species of Diogenes.
Discapseudes (Miodiscapseudes)[57]	Subgen. et sp. nov	Valid	Heard in Heard, de Lourdes Serrano-Sánchez & Vega	Early Miocene	Mexican amber	Mexico	A member of Tanaidacea belonging to the group Apseudomorpha. The subgenus includes new species D. chiapasensis.
Eopetrolisthes levistriatus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A porcelain crab.
Eucalliax multisetae[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Callianassidae.
Eucalliax rugata[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Callianassidae.
Exucarcinus[58]	Gen. et sp. nov	Valid	Prado & Luque in Prado et al.	Early Cretaceous (late Aptian–early Albian)	Romualdo Formation	Brazil	A crab belonging to the group Raninoida and the family Orithopsidae. The type species is E. gonzagai.
Galathea hexacristata[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A species of Galathea.
Gemmellarocarcinus disalvoi[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab, a member of Dromiacea of uncertain phylogenetic placement.
Guinotosia ornata[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab, a member of the family Etyiidae.
Homarus hungaricus[59]	Sp. nov	Valid	Tshudy et al.	Oligocene (Chattian)	Mány Formation	Hungary	A species of Homarus.
Hoploparia nasilowensis[56]	Sp. nov	Valid	Fraaije et al.	Paleocene (Danian)		Poland
Hyastenus antiquus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab belonging to the family Epialtidae, a species of Hyastenus.
Hypothalassia campolongensis[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab, a species of Hypothalassia.
Icriobranchiocarcinus[60]	Gen. et sp. nov	Valid	Vega in Vega et al.	Late Cretaceous (Maastrichtian)		Mexico	A member crab related to Icriocarcinus. Genus includes new species I. tzutzu.
Inachus eocenicus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab belonging to the family Inachidae, a species of Inachus.
Kromtitis tergospinosus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab belonging to the family Dynomenidae.
Laevitealliocaris[61]	Gen. et sp. nov	Valid	Yang et al.	Late Carboniferous	Tupo Formation	China	A relative of Tealliocaris. The type species is L. xiaheyanensis.
Lessiniamathia[62]	Gen. et sp. nov	Valid	Ceccon & De Angeli	Eocene (Ypresian)		Italy	A crab belonging to the family Epialtidae. Genus includes new species L. bolcense.
Lessinipagurus bericus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A hermit crab belonging to the family Xylopaguridae.
Longipaguristes regularis[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A hermit crab belonging to the family Diogenidae.
Longodromites aubreyae[43]	Sp. nov	Valid	Schweitzer et al.	Late Jurassic		Romania	A crab belonging to the family Longodromitidae.
Maeandricampus starri[63]	Sp. nov	Valid	Feldmann, Schweitzer & Goedert	Late Oligocene to earliest Miocene	Lincoln Creek Formation	United States	A crab belonging to the family Carcinidae.
Mesolambrus bragai[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab belonging to the family Parthenopidae.
Occultocaris feckei[49]	Sp. nov	Valid	Winkler	Late Jurassic (Tithonian)	Solnhofen Limestone	Germany	A caridean shrimp.
Pachycheles cristatus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A porcelain crab, a species of Pachycheles.
Pachycheles multituberculatus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A porcelain crab, a species of Pachycheles.
Pachycheles semiornatus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A porcelain crab, a species of Pachycheles.
Paguristes michikoae[64]	Sp. nov	Valid	Karasawa & Fudouji	Early Oligocene	Sari Sandstone Member of the Kishima Group	Japan	A species of Paguristes.
Paguristes teruakii[64]	Sp. nov	Valid	Karasawa & Fudouji	Early Oligocene	Sari Sandstone Member of the Kishima Group	Japan	A species of Paguristes.
Paguristes vallionensis[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A species of Paguristes.
Palaemon monsdamarum[65]	Sp. nov	Valid	Pasini & Garassino	Miocene (Messinian)		Italy	A species of Palaemon.
Palaeoarmadillo[66]	Gen. et sp. nov	Valid	Poinar	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	An isopod belonging to the family Armadillidae. Genus includes new species P. microsoma.
Palaeolibrinus[47]	Gen. et sp. nov	Valid	Broly in Broly, Serrano-Sánchez & Vega	Early Miocene	La Quinta Formation (Mexican amber)	Mexico	A woodlouse belonging to the family Olibrinidae. The type species is P. spinicornis.
Palaeospherarmadillo[47]	Gen. et 2 sp. nov	Valid	Broly in Broly, Serrano-Sánchez & Vega	Early Miocene	Mazantic Shale (Mexican amber)	Mexico	A woodlouse. The type species is P. mazanticus; genus also includes P. rotundus.
Palicoides faggioni[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab belonging to the family Palicidae.
Panopeus paucicarinatus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab belonging to the family Panopeidae, a species of Panopeus.
Paraclythia cernavoda[55]	Sp. nov	Valid	Frantescu et al.	Early Cretaceous (Aptian)	Ramadan Formation	Romania	A lobster.
Petrolisthes tuberculatus[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A porcelain crab, a species of Petrolisthes.
Pilidromia[43]	Gen. et comb. nov	Valid	Schweitzer et al.	Late Jurassic		Germany  Romania	A crab belonging to the family Longodromitidae. The type species is "Planoprosopon" thiedeae Schweigert & Koppka (2011).
Pithonoton lluismariaorum[67]	Sp. nov	Valid	Ossó et al.	Early Cretaceous (Aptian)	Margas de Forcall Formation	Spain	A crab.
Politohepatiscus[46]	Gen. et sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab, a member of the family Aethridae. The type species is P. zorzini.
Porcellana elegans[46]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A porcelain crab, a species of Porcellana.
Prebranchioplax[68]	Gen. et sp. nov	Valid	Vega et al.	Late Cretaceous (Campanian)	Cerro del Pueblo Formation Parras Shale	Mexico	A crab belonging to the family Mathildellidae. Genus includes new species P. cretacica.
Priorhyncha[69]	Gen. et sp. nov	Valid	Alencar et al.	Early Cretaceous (Aptian-Albian)		Brazil	A member of Dendrobranchiata belonging to the family Solenoceridae. Genus includes new species P. feitosai.
Proeryon charbonnieri[70]	Sp. nov	Valid	Audo & Schweigert	Middle Jurassic (Callovian)	La Voulte-sur-Rhône Lagerstätte	France	A member of the family Coleiidae.
Prosopon barbulescuae[43]	Sp. nov	Disputed	Schweitzer et al.	Late Jurassic		Romania	A crab belonging to the family Prosopidae. Klompmaker et al. (2020) considered it to be a junior synonym of Europrosopon aculeatum (von Meyer, 1857).[71]
Pseudodromilites arzignanensis[72]	Sp. nov	Valid	De Angeli & Alberti	Eocene (Lutetian)		Italy	A crab belonging to the family Dromiidae.
Pseudopancolus[57]	Gen. et sp. nov	Valid	Heard in Heard, de Lourdes Serrano-Sánchez & Vega	Early Miocene	Mexican amber	Mexico	A member of Tanaidacea belonging to the group Tanaidomorpha. Genus includes new species P. minutus.
Rathbunopon tarraconensis[67]	Sp. nov	Valid	Ossó et al.	Early Cretaceous (Aptian)	Margas de Forcall Formation	Spain	A crab.
Romaleon franciscae[73]	Sp. nov	Valid	Robin et al.	Miocene		French Southern and Antarctic Lands (Kerguelen Islands)	A species of Romaleon.
Romaniacheiros[55]	Gen. et sp. nov	Valid	Frantescu et al.	Late Jurassic (Kimmeridgian-Tithonian)		Romania	A member of Glypheoidea belonging to the family Mecochiridae. Genus includes new species R. lophia.
Romualdocarcinus[58]	Gen. et sp. nov	Valid	Prado & Luque in Prado et al.	Early Cretaceous (late Aptian–early Albian)	Romualdo Formation	Brazil	A crab belonging to the group Heterotremata, possibly a member of the family Eogeryonidae. The type species is R. salesi.
Sakakurapsus[48]	Gen. et sp. nov	Valid	Karasawa	Miocene	Lower Sandstone Member of the Bihoku Group	Japan	A crab belonging to the family Varunidae. The type species is S. kogisorum.
Schramocaris clarksoni[74]	Sp. nov	Valid	Clark, Miller & Ross	Carboniferous (Viséan)		United Kingdom	A eumalacostracan crustacean similar to Tealliocaris and Pseudogalathea.
Schramocaris matthewi[74]	Sp. nov	Valid	Clark, Miller & Ross	Carboniferous (Viséan)		Canada	A eumalacostracan crustacean similar to Tealliocaris and Pseudogalathea.
Sume[75]	Gen. et sp. nov	Valid	Saraiva, Pinheiro & Santana	Early Cretaceous (Aptian/Albian)	Romualdo Member	Brazil	A member of the family Luciferidae. Genus includes new species S. marcosi.
Tanidromites starzykae[43]	Sp. nov	Valid	Schweitzer et al.	Late Jurassic		Romania	A crab belonging to the family Tanidromitidae.
Tethyranina[76]	Gen. et comb. nov	Valid	Pasini & Garassino	Early Miocene – early Pleistocene		Italy	A crab belonging to the family Raninidae. The type species is "Ranina" propinqua Ristori (1891).
Vegaranina rivasi[77]	Sp. nov	Valid	Arano-Ruiz et al.	Late Cretaceous		Cuba	A member of the family Raninidae.
Zannatoius[46]	Gen. et sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A crab, a member of the family Leucosiidae. The type species is Z. vicetinus.

Ostracods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acratia caeracella[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Acratia maugerii[79]	Sp. nov	Valid	Crasquin, Sciuto & Reitano	Late Triassic (Carnian)	Mufara Formation	Italy
Advenocypris similis[80]	Sp. nov	Valid	Teterina	Pliocene	Beken Formation	Russia	A member of the family Candonidae.
Aechmina akumame[81]	Sp. nov	Valid	Stocker et al.	Carboniferous (Moscovian)	Ichinotani Formation	Japan	A member of Podocopa belonging to the order Beyrichiocopida and the family Aechminidae.
Aechmina? meidlai[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Aechminidae.
Alciella irizukii[83]	Sp. nov	Valid	Yasuhara et al.	Eocene		Madagascar
Alicenula longiformis[84]	Sp. nov	Valid	Leite et al.	Early Cretaceous	Quiricó Formation	Brazil
Alicenula sousaensis[85]	Sp. nov	Valid	Sousa, Carvalho & Ferreira	Early Cretaceous	Sousa Formation	Brazil
Amplocypris qaidamensis[86]	Sp. nov	Valid	Yang et al.	Quaternary	Qaidam Basin	China	A member of the family Cyprididae.
“Anchistrocheles” gemmellaroi[79]	Sp. nov	Valid	Crasquin, Sciuto & Reitano	Late Triassic (Carnian)	Mufara Formation	Italy
Aphelocythere dilgeri[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian and Bajocian)	Upper Opalinuston Formation	Germany	A member of the family Cytheruridae.
Bairdia asgarda[88]	Sp. nov	Valid	Ayress & Gould	Early Cretaceous	Åsgard Formation	North Sea Norwegian Sea	A member of Podocopida belonging to the group Bairdiocopina and the family Bairdiidae.
Bairdia liviae[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Bairdia mandruella[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Bairdia monostorii[78]	Nom. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Bairdia scaliae[79]	Sp. nov	Valid	Crasquin, Sciuto & Reitano	Late Triassic (Carnian)	Mufara Formation	Italy
Bairdia tagaensis[89]	Sp. nov	Valid	Tanaka in Tanaka et a.	Permian (Cisuralian)	Ryozensan Limestone Formation	Japan
Bairdiacypris ikeyanoriyukii[89]	Sp. nov	Valid	Tanaka in Tanaka et a.	Permian (Cisuralian)	Ryozensan Limestone Formation	Japan
Bairdiacypris sorgunensis[90]	Sp. nov	Valid	Forel in Forel, Ozsvárt & Moix	Late Triassic (Carnian)		Turkey	A member of the family Bairdiidae.
Baltonotella? admirabilis[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Aparchitidae.
Bektasia yawella[91]	Sp. nov	Valid	Forel in Ketmuangmoon et al.	Middle Triassic (Anisian)	Pha Kan Formation	Thailand
Bolbinella limbata[92]	Sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Mangazeya Formation	Russia	A member of Leperditellocopida belonging to the family Bolliidae.
Bythoceratina spinosa[93]	Sp. nov	Valid	Barros & Piovesan in Barros, Piovesan & Agostinho	Late Cretaceous (Maastrichtian)	Gramame Formation	Brazil	A member of Podocopida belonging to the family Bythocytheridae.
Cardobairdia tesakovae[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian)		Germany	A member of the family Healdiidae.
Clithrocytheridea ljubimovae[94]	Sp. nov	Valid	Dykan in Dykan et al.	Paleogene	Boltysh crater	Ukraine	A member of the family Cytherideidae.
Clivosolobella[92]	Gen. et sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Ust’-Stolbovaya Formation	Russia	A member of Hollinocopida belonging to the family Egorovellidae. The type species is C. rugula.
Collibolbina habeotubercula[92]	Sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Mangazeya Formation	Russia	A member of Hollinocopida belonging to the family Tetradellidae.
Crescentilla laciniata[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Aechminidae.
Cypridea paraibensis[85]	Sp. nov	Valid	Sousa, Carvalho & Ferreira	Early Cretaceous	Sousa Formation	Brazil
Cypridea samesi[95]	Sp. nov	Valid	Choi, Wang & Huh	Early Cretaceous	Jinju Formation	South Korea
Cypridea vianai[85]	Sp. nov	Valid	Sousa, Carvalho & Ferreira	Early Cretaceous	Sousa Formation	Brazil
Cypridea zalanyii[96]	Sp. nov	Valid	Cséfán & Tóth	Early Cretaceous (Albian)		Hungary
Cyprideis civisi[97]	Sp. nov		Ruiz, González-Regalado & Abad	Pliocene		Spain	A member of the family Cytherideidae.
Cytherella boltyshevae[94]	Sp. nov	Valid	Dykan in Dykan et al.	Paleogene	Boltysh crater	Ukraine	A member of the family Cytherellidae.
Cytherella centrocompressa[93]	Sp. nov	Valid	Barros & Piovesan in Barros, Piovesan & Agostinho	Late Cretaceous (late Maastrichtian)	Gramame Formation	Brazil	A member of the family Cytherellidae.
Cytherella paraibensis[93]	Sp. nov	Valid	Barros & Piovesan in Barros, Piovesan & Agostinho	Late Cretaceous (late Maastrichtian)	Gramame Formation	Brazil	A member of the family Cytherellidae.
Cytherelloidea baghinensis[98]	Sp. nov	Valid	Khosravi, Vaziri & Arab	Early Cretaceous (Aptian)		Iran
Cytherelloidea mazarensis[98]	Sp. nov	Valid	Khosravi, Vaziri & Arab	Early Cretaceous (Aptian)		Iran
Cytherelloidea sirizensis[98]	Sp. nov	Valid	Khosravi, Vaziri & Arab	Early Cretaceous (Aptian)		Iran
Cytheropterina alacostata[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian)	Achdorf Formation	Germany	A member of the family Cytheruridae.
Darwinula oryzalis[99]	Nom. nov	Valid	Naumcheva	Early Triassic	Vokhma Formation	Russia	A replacement name for Darwinula acuta Mishina (1966).
Darwinula vetusta[100]	Nom. nov	Valid	Naumcheva	Permian		Russia	A replacement name for Darwinula mera Starozhilova (1967).
Disulcina (?) complanata[92]	Sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Dolborsa Formation	Russia	A member of Hollinocopida belonging to the family Sylthiidae.
Dorsogibbella[92]	Gen. et 3 sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Mangazeya Formation	Russia	A member of Hollinocopida belonging to the family Piretellidae. The type species is D. hippocrepicus; genus also includes new species D. claviformis and D. costaventralis.
Easchmidtella gibbosa[92]	Sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Dolborsa Formation	Russia	A member of Leperditellocopida belonging to the family Schmidtellidae.
Eucyprinotus qinghaiensis[86]	Sp. nov	Valid	Yang et al.	Quaternary	Qaidam Basin	China	A member of the family Cyprididae.
Eucypris lobatoi[101]	Sp. nov	Valid	Bergue, Ramos & Maranhão	Oligocene	Tremembé Formation	Brazil	A member of the family Cyprididae.
Eucytherura ventrotuberculata[93]	Sp. nov	Valid	Barros & Piovesan in Barros, Piovesan & Agostinho	Late Cretaceous (Maastrichtian) and Paleocene (Danian)	Gramame Formation	Argentina  Brazil	A member of Podocopida belonging to the family Cytheruridae.
Euprimites (Euprimites) kralodvorensis[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Tvaerenellidae.
Gaindella[92]	Gen. et sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Dolborsa Formation	Russia	A member of Hollinocopida belonging to the family Egorovellidae. The type species is G. signata.
Healdia mizuboradanensis[81]	Sp. nov	Valid	Stocker et al.	Carboniferous (Moscovian)	Ichinotani Formation	Japan	A member of Podocopa belonging to the order Podocopida and the family Healdiidae.
Hemicypris lomastroi[102]	Sp. nov	Valid	Spadi, Gliozzi & Medici	Piacenzian–Gelasian	Dunarobba Fossil Forest	Italy
Heterocypris subtriangularis[86]	Sp. nov	Valid	Yang et al.	Quaternary	Qaidam Basin	China	A member of the family Cyprididae.
Hungarella mahmudiaensis[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Hungarella poli[91]	Sp. nov	Valid	Chitnarin in Ketmuangmoon et al.	Middle Triassic (Anisian)	Pha Kan Formation	Thailand
Hungaroleberis[103]	Gen. et sp. et comb. nov	Valid	Tóth & Cséfán	Late Permian to Middle Jurassic (Bajocian)		China  Greece  Hungary	A member of Myodocopa. The type species is H. retiferus; genus also includes "Cypridina" gussevaae Crasquin-Soleau & Baud (1998).
?Infracytheropteron bisulcatum[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian)		Germany	A member of the family Cytheruridae.
Judahella? montanarii[79]	Sp. nov	Valid	Crasquin, Sciuto & Reitano	Late Triassic (Carnian)	Mufara Formation	Italy
Kellettina noriyukii[89]	Sp. nov	Valid	Tanaka in Tanaka et a.	Permian (Cisuralian)	Ryozensan Limestone Formation	Japan
Kempfina tineriella[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Kinkelinella (Kinkelinella) geisingensis[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian)		Germany	A member of the family Progonocytheridae.
Langiella fauthi[93]	Sp. nov	Valid	Barros & Piovesan in Barros, Piovesan & Agostinho	Late Cretaceous (Maastrichtian)	Gramame Formation	Brazil	A member of Podocopida belonging to the family Trachyleberididae.
Langtonia abdomorostrata[94]	Sp. nov	Valid	Dykan in Dykan et al.	Paleogene	Boltysh crater	Ukraine
Lenatella arcuata[92]	Sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Mangazeya Formation	Russia	A member of the family Piretellidae.
Leviella lampangensis[91]	Sp. nov	Valid	Forel in Ketmuangmoon et al.	Middle Triassic (Anisian)	Pha Kan Formation	Thailand
Liuzhinia antalyaformis[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Liuzhinia bithynica[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Liuzhinia edvardgriegi[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Microcheilinella? contrariella[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Microcheilinella shigensis[89]	Sp. nov	Valid	Tanaka in Tanaka et a.	Permian (Cisuralian)	Ryozensan Limestone Formation	Japan
Milleratia tungusica[92]	Sp. nov	Valid	Gonta & Kanygin	Ordovician		Russia	A member of Eridocopida belonging to the family Cryptophillidae.
Muellerina eocenica[83]	Sp. nov	Valid	Yasuhara et al.	Eocene		Madagascar
Neocyprideis polygonoreticulata[83]	Sp. nov	Valid	Yasuhara et al.	Eocene		Madagascar
Neomonoceratina afroangulosa[83]	Sp. nov	Valid	Yasuhara et al.	Eocene		Madagascar
Neomonoceratina iwasakii[104]	Sp. nov	Valid	Tanaka in Tanaka et al.	Eocene	Akasaki Formation	Japan	A member of Podocopida belonging to the family Schizocytheridae.
Neonesidea potyensis[93]	Sp. nov	Valid	Barros & Piovesan in Barros, Piovesan & Agostinho	Late Cretaceous (Maastrichtian)	Gramame Formation	Brazil	A member of Podocopida belonging to the family Bairdiidae.
Neorichterina[103]	Gen. et sp. nov	Valid	Tóth & Cséfán	Early Cretaceous (Albian)		Brazil  Gabon  Hungary  Israel  Morocco   Switzerland  United Kingdom Pacific Ocean (Mid Pacific Mountains)	A member of Myodocopa belonging to the family Entomozoidae. The type species is N. striata.
Neothlipsura khiska[105]	Sp. nov	Valid	Salas et al.	Late Silurian	Tambolar Formation	Argentina
Oliganisus ryozensannensis[89]	Sp. nov	Valid	Tanaka in Tanaka et a.	Permian (Cisuralian)	Ryozensan Limestone Formation	Japan
Paijenborchella amakusensis[104]	Sp. nov	Valid	Tanaka et al.	Eocene	Akasaki Formation	Japan	A member of Podocopida belonging to the family Schizocytheridae.
Paijenborchellina madagascarensis[83]	Sp. nov	Valid	Yasuhara et al.	Eocene		Madagascar
Palaeocypridina goeroegae[103]	Sp. nov	Valid	Tóth & Cséfán	Middle Triassic (Ladinian)	Buchenstein Formation	Hungary	A member of Myodocopa.
Paracypris oanaella[78]	Sp. nov	Valid	Forel & Grădinaru	Middle Triassic (Anisian)		Romania
Paralimnocythere turgida[102]	Sp. nov	Valid	Spadi, Gliozzi & Medici	Piacenzian–Gelasian	Dunarobba Fossil Forest	Italy
Parenthatia hanamiloslavensis[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Bolliidae.
Penthesilenula pintoi[84]	Sp. nov	Valid	Leite et al.	Early Cretaceous	Quiricó Formation	Brazil
Pleurocythere ohmerti[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian and Bajocian)		Germany	A member of the family Protocytheridae.
Pokornyopsis csurgokutensis[103]	Sp. nov	Valid	Tóth & Cséfán	Early Jurassic (Toarcian)		Hungary	A member of Myodocopa.
Polycope circulosa[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian)		Germany	A member of the family Polycopidae.
Pontocyprella valhalla[88]	Sp. nov	Valid	Ayress & Gould	Early Cretaceous	Tuxen Formation Valhall Formation	North Sea Norwegian Sea
Potamocypris taubatensis[101]	Sp. nov	Valid	Bergue, Ramos & Maranhão	Oligocene	Tremembé Formation	Brazil	A member of the family Cyprididae.
Praeschuleridea concentrica[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian)		Germany	A member of the family Schulerideidae.
Procytheropteron catena[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian)		Germany	A member of the family Cytheruridae.
Progonocythere scutula[87]	Sp. nov	Valid	Franz, Ebert & Stulpinaite	Middle Jurassic (Aalenian)		Germany	A member of the family Progonocytheridae.
Protobuntonia punctatum[93]	Sp. nov	Valid	Barros & Piovesan in Barros, Piovesan & Agostinho	Late Cretaceous (late Maastrichtian) and Paleocene (early Danian)	Gramame Formation	Brazil	A member of Podocopida belonging to the family Trachyleberididae.
Pseudulrichia lucberkensis[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Aechminidae.
Pseudulrichia risumata[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Aechminidae.
Psychrodromus mangnaiensis[86]	Sp. nov	Valid	Yang et al.	Quaternary	Qaidam Basin	China	A member of the family Cyprididae.
Pustulobairdia ohmiensis[89]	Sp. nov	Valid	Tanaka in Tanaka et a.	Permian (Cisuralian)	Ryozensan Limestone Formation	Japan
Rectella bublei[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Rectellidae.
Reigiopsis conicus[92]	Sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Mangazeya Formation	Russia	A member of Hollinocopida belonging to the family Tetradellidae.
Reigiopsis tumulus[92]	Sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Mangazeya Formation	Russia	A member of Hollinocopida belonging to the family Tetradellidae.
Scutumella gruminosa[92]	Sp. nov	Valid	Gonta & Kanygin	Late Ordovician	Mangazeya Formation	Russia	A member of Leperditellocopida belonging to the family Bolliidae.
Spinigerites ostrovites[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Spinigeritidae.
Spiricopia[106]	Gen. et sp. nov	Valid	Siveter et al.	Silurian	Wenlock Series Lagerstätte	United Kingdom	A member of Myodocopida belonging to the family Cylindroleberididae. The type species is S. aurita.
Stigmatobradleya[83]	Gen. et sp. nov	Valid	Yasuhara et al.	Eocene		Madagascar	Genus includes new species S. hunti.
Timiriasevia sanfranciscanensis[84]	Sp. nov	Valid	Leite et al.	Early Cretaceous	Quiricó Formation	Brazil
Trajancypris nanlingqiuensis[86]	Sp. nov	Valid	Yang et al.	Quaternary	Qaidam Basin	China	A member of the family Cyprididae.
Triassocypris phakanella[91]	Sp. nov	Valid	Forel in Ketmuangmoon et al.	Middle Triassic (Anisian)	Pha Kan Formation	Thailand
Tyrrhenocythere pulcherrima[107]	Sp. nov	Valid	Sciuto et al.	Miocene (Messinian) to early Pleistocene)		Italy	A member of Podocopida belonging to the family Hemicytheridae.
Uroleberis paranuda[83]	Sp. nov	Valid	Yasuhara et al.	Eocene		Madagascar
Wangshangkia[108]	Gen. et 2 sp. nov	Valid	Song & Gong	Late Devonian	Gelaohe Formation	China	A member of Podocopida belonging to the superfamily Bairdiocypridoidea and the family Bairdiocyprididae. The type species is W. dushaniensis; genus also includes W. bailouiensis.
Warthinia trinoda[82]	Sp. nov	Valid	Lajblová & Kraft	Ordovician (Katian)	Králův Dvůr Formation	Czech Republic	A member of the family Bolliidae.
Xestoleberis renemai[83]	Sp. nov	Valid	Yasuhara et al.	Eocene		Madagascar

Other crustaceans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Diestheria deshenglingensis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Diestheria latovata[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Diestheria ovaliformis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Diestheria striaris[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Diestheria xiaoqiaomugouensis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Diestheria yangshugouensis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Eosestheria gracilis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Eosestheria jiuwuensis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Eosestheria lata[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Eosestheria magna[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Eosestheria obliqua[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Eosestheria paucilinearis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Eosestheria shanwanensis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Eosestheria shibapanensiss[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Eosestheria yangshugouensiss[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Euestheria kozuri[110]	Sp. nov	Valid	Geyer & Kelber	Late Triassic (Carnian)	Hassberge Formation	Germany	A clam shrimp.
Gregoriusella striatula[110]	Sp. nov	Valid	Geyer & Kelber	Late Triassic (Carnian)	Hassberge Formation	Germany	A clam shrimp.
Hemicycloleaia ejinaqiensis[111]	Sp. nov		Niu in Lu et al.	Permian		China	A diplostracan belonging to the family Leaiidae.
Hexaminius venerai[112]	Sp. nov	Valid	Kočí et al.	Middle Eocene	La Meseta Formation	Antarctica (Seymour Island)	A barnacle.
Platyestheria[113]	Gen. et comb. nov	Valid	Bittencourt et al.	Early Cretaceous	Quiricó Formation	Brazil	A clam shrimp; a new genus for "Pseudestheria" abaetensis Cardoso (1971).
Polygrapta neimengguensis[111]	Sp. nov		Niu in Lu et al.	Permian		China	A clam shrimp belonging to the family Euestheriidae.
Skuinocyclus[114]	Gen. et sp. nov	Valid	Mychko & Alekseev	Permian (Cisuralian)		Russia	A member of Cyclida. Genus includes new species S. juliae.
Undulatesta[115]	Gen. et sp. nov	Valid	Gueriau, Rabet & Du Tien Hat	Devonian (late Famennian)		Belgium	A clam shrimp belonging to the group Vertexioidea. The type species is U. bounonensis.
Uralocyclus[114]	Gen. et comb. nov	Valid	Mychko & Alekseev	Permian (Asselian)		Russia	A member of Cyclida. Genus includes "Cyclus" miloradovitchi Kramarenko (1961).
Yanjiestheria anchungouensis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Yanjiestheria deshenglingensis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Yanjiestheria rotunda[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Yanjiestheria shanwanensis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.
Yanjiestheria xiaoqiaomugouensis[109]	Sp. nov		Niu in Niu & Xin	Cretaceous		China	A clam shrimp.

Insects

Main article: 2018 in insect paleontology

Trilobites

Research

• Frontal auxiliary impressions (muscle scars on the glabella) are described in Mesolenellus hyperboreus from Cambrian Stage 4 strata of North Greenland by Lerosey‐Aubril & Peel (2018), providing new information on the evolution of a pouch‐like digestive organ with powerful extrinsic muscles (i.e. a crop) in trilobites.[116]
• A study on complete moulted exoskeletons of Estaingia bilobata and Redlichia takooensis from the Cambrian Emu Bay Shale (Australia), and their implications for inferring the course of moulting in trilobites, is published by Drage et al. (2018).[117]
• A study assessing the morphological differences within and among three populations of Oryctocephalus indicus from the United States, Siberia and China, using a new method of analysing small morphological features, is published by Esteve et al. (2018).[118]
• A study on early post-embryonic developmental stages of the Cambrian trilobite Ellipsostrenua granulosa is published by Laibl, Cederström & Ahlberg (2018).[119]
• A study on the ontogeny of the co-occurring shumardiid trilobites Akoldinioidia latus and Koldinioidia choii from the middle Furongian Hwajeol Formation (South Korea) is published by Park (2018).[120]
• Remains of the digestive system of a specimen of the bathycheilid species Prionocheilus vokovicensis from the Ordovician Šárka Formation (Czech Republic) are described by Fatka & Budil (2018).[121]
• A study on the postembryonic development of Dalmanitina, based on a large sample of specimens from the Upper Ordovician Prague Basin, is published by Drage, Laibl & Budil (2018).[122]
• A large and almost complete dorsal exoskeleton of a trilobite referred to the species Dipleura dekayi is described from the Devonian Floresta Formation (Colombia) by Carvalho (2018).[123]
• A study on the phylogenetic relationships of species belonging to the genus Metacryphaeus is published by Carbonaro et al. (2018), who also perform a palaeobiogeographic analysis for the distribution of the genus.[124]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Adelogonus prichardi[125]	Sp. nov	Valid	Smith, Paterson & Brock	Cambrian	Goyder Formation	Australia
Amphiroaspis[126]	Gen. et comb. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the family Phillipsiidae belonging to the subfamily Weaniinae. The type species is "Carbonocoryphe (Winterbergia)" amphiro Hahn, Hahn & Müller (1996); genus also includes "Carbonocoryphe (Winterbergia)" sporada Hahn, Hahn & Müller (2007).
Asaphellus albae[127]	Sp. nov	Valid	Meroi Arcerito et al.	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Austerops salamandaroides[128]	Sp. nov	Valid	Crônier et al.	Devonian	Chefar el Ahmar Formation	Algeria
Bailiella sepulcra[129]	Sp. nov	Valid	Álvaro & Vizcaïno	Cambrian (Furongian)	Bailiella Formation	Morocco	A member of the family Conocoryphidae.
Belgibole kalypso[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Bradocryphaeus echinatus[130]	Sp. nov	Valid	Van Viersen & Vanherle	Devonian (Frasnian)	Moulin Liénaux Formation	Belgium	A member of the family Acastidae.
Bradocryphaeus laomedeia[130]	Sp. nov	Valid	Van Viersen & Vanherle	Devonian (Frasnian)	Grands Breux Formation	Belgium	A member of the family Acastidae.
Bradocryphaeus neptuni hottonensis[130]	Subsp. nov	Valid	Van Viersen & Vanherle	Devonian (Frasnian)	Grands Breux Formation	Belgium	A member of the family Acastidae.
Brevibole archinalae[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Brevibole halszkae[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Buchiproetus? exconcha[131]	Sp. nov	Valid	Flick	Devonian		Germany	A member of Proetida belonging to the family Tropidocoryphidae.
Buenellus chilhoweensis[132]	Sp. nov	Valid	Webster & Hageman	Cambrian Series 2 (Montezuman)	Murray Shale	United States
Chlupacula (Chiides) pandora[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Cornuproetus pusulosus[133]	Sp. nov	Valid	Feist & Belka	Devonian (Emsian)		Morocco
Cornuproetus? runzheimeri[131]	Sp. nov	Valid	Flick	Devonian		Germany	A member of Proetida belonging to the family Tropidocoryphidae.
Crassibole ate[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Crassibole? walliseri[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Cyphaspis koimeterionensis[130]	Sp. nov	Valid	Van Viersen & Vanherle	Devonian (Frasnian)	Grands Breux Formation	Belgium	A member of the family Aulacopleuridae.
Cyrtoproetus eremus[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Cyrtoproetus galapagos[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Cyrtoproetus pegasus[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Deltapliomera[134]	Gen. et 3 sp. et comb. nov	Valid	McAdams, Adrain & Karim	Early Ordovician		United States	The type species is D. humphriesi; genus also includes new species D. eppersoni and D. inglei, as well as D. heimbergi.
Diacoryphe (Diacoryphe) phoenix[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Ditomopyge (Carniphillipsia) mosquensis[135]	Sp. nov	Valid	Mychko & Alekseev	Carboniferous (Gzhelian)		Russia
Duyunaspis jianheensis[136]	Sp. nov	Valid	Chen et al.	Cambrian Stage 4		China	An oryctocephalid trilobite.
Dysplanus acutigenia putilovoensis[137]	Subsp. nov	Valid	Krylov	Ordovician (Dapingian)		Russia
Erdbachaspis[126]	Gen. et comb. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the family Phillipsiidae belonging to the subfamily Archegoninae. The type species is "Phillipsia" granulifera Holzapfel (1889); genus also includes E. aithyia (Hahn, Hahn & Müller, 2001), E. bathapoikos (Hahn, Hahn & Müller, 1998) and possibly "Proliobole" krambergensis Hahn, Hahn & Müller (1996).
Harpes hamarlaghdadensis[138]	Sp. nov	Valid	Crônier et al.	Devonian (Emsian)		Morocco
Hebeia stewarti[125]	Sp. nov	Valid	Smith, Paterson & Brock	Cambrian	Goyder Formation	Australia
Ibexaspis coadyi[134]	Sp. nov	Valid	McAdams, Adrain & Karim	Early Ordovician		United States
Ibexaspis leuppi[134]	Sp. nov	Valid	McAdams, Adrain & Karim	Early Ordovician		United States
Ibexaspis rupauli[134]	Sp. nov	Valid	McAdams, Adrain & Karim	Early Ordovician		United States
Latibole gamma[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Latibole parapaprothae[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Latibole pleon[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Leonaspis bassei[139]	Sp. nov	Valid	Alberti	Devonian (Emsian)		Germany
Liebsteinella[126]	Gen. et comb. et 2 sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the subfamily Cystispininae. The type species is "Tawstockia" martini Müller (2004); genus also includes new species L. sauerlandica and L. weberi.
Liobole (Liobole) meyeri[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Liobole (Liobole) io[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Liobole (Panibole) emrichterae[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Liobole (Panibole) groeningae[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Liobole (Panibole) isis[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Liobole (Panibole) paraglabroides[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Liobole (Panibole) subaequalis hschmidti[126]	Subsp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Lioharpes wendti[133]	Sp. nov	Valid	Feist & Belka	Devonian (Emsian)		Morocco
Liostracina joyceae[125]	Sp. nov	Valid	Smith, Paterson & Brock	Cambrian	Goyder Formation	Australia
Longaspis[140]	Gen. et sp. nov	Valid	Peng et al.	Cambrian Stage 4	Balang Formation	China	A member of the family Oryctocephalidae. Genus includes new species L. paiwuensis.
Luciaspis[141]	Gen. et sp. nov	Valid	Álvaro et al.	Cambrian	Huérmeda Formation	Spain	A saukiandid trilobite. Genus includes new species L. matiasi.
Magreanops[130]	Gen. et 2 sp. nov	Valid	Van Viersen & Vanherle	Devonian (Frasnian)	Bovesse Formation Moulin Liénaux Formation	Belgium	A member of the family Phacopidae. The type species is M. renateae; genus also includes M. monachus.
Maladioidella? destombesi[129]	Sp. nov	Valid	Álvaro & Vizcaïno	Cambrian (Furongian)	Bailiella Formation	Morocco	A member of the family Idahoiidae.
Millardaspis[134]	Gen. et 2 sp. nov	Valid	McAdams, Adrain & Karim	Early Ordovician		United States	The type species is M. milsteadi; genus also includes M. knoxi.
Morocops davidbrutoni[138]	Sp. nov	Valid	Crônier et al.	Devonian (Emsian)		Morocco
Neoillaenus[137]	Gen. et comb. nov	Valid	Krylov	Ordovician (Darriwilian—Katian)		Estonia  Russia  Sweden	The type species is "Illaenus" excellens Holm (1886); genus also includes Neoillaenus atavus (Eichwald, 1857), N. oblongatus (Angelin, 1854), N. kukersianus (Holm, 1886), N. itferensis (Holm, 1886) and N. praecurriens (Jaanusson, 1957).
Omegops tilabadensis[142]	Sp. nov	Valid	Ghobadi Pour et al.	Devonian (Famennian)	Khoshyeilaq Formation	Iran	A member of the family Phacopidae.
Onaraspis garciae[141]	Sp. nov	Valid	Álvaro et al.	Cambrian	Huérmeda Formation	Spain	A saukiandid trilobite.
Panderia balashovae[137]	Sp. nov	Valid	Krylov	Early Ordovician		Russia
Panderia koshkarovi[137]	Sp. nov	Valid	Krylov	Ordovician (Darriwilian)		Russia
Paulaspis[141]	Gen. et sp. nov	Valid	Álvaro et al.	Cambrian	Huérmeda Formation	Spain	A saukiandid trilobite. Genus includes new species P. tiergaensis.
Phacops ouarouroutensis[128]	Sp. nov	Valid	Crônier et al.	Devonian	Chefar el Ahmar Formation	Algeria
Phaetonellus dillensis[143]	Sp. nov	Valid	Basse & Müller	Devonian		Germany
Phaetonellus greifensteinensis[143]	Sp. nov	Valid	Basse & Müller	Devonian		Germany
Phaetonellus heckelmanni[143]	Sp. nov	Valid	Basse & Müller	Devonian		Germany
Phaetonellus kimi[143]	Sp. nov	Valid	Basse & Müller	Devonian		Germany
Phaetonellus martinius[143]	Sp. nov	Valid	Basse & Müller	Devonian	Leun Limestone	Germany
Perliproetus africanus[144]	Sp. nov	Valid	Feist & Weyer	Devonian (Famennian)		Morocco
Perliproetus kallenhardtensis[144]	Sp. nov	Valid	Feist & Weyer	Devonian (Famennian)		Germany
Pterocoryphe platymarginata[130]	Sp. nov	Valid	Van Viersen & Vanherle	Devonian (Frasnian)	Champ Broquet Formation	Belgium	A member of the family Proetidae.
Quadratispina excelsa[130]	Sp. nov	Valid	Van Viersen & Vanherle	Devonian (Frasnian)	Bovesse Formation	Belgium	A member of the family Acastidae.
Redlichia isuelaensis[141]	Sp. nov	Valid	Álvaro et al.	Cambrian	Huérmeda Formation	Spain
Spasskyproetus[143]	Gen. et comb. nov	Valid	Basse & Müller	Devonian (Emsian)		Russia	A new genus for "Phaetonellus" bondarevi Spassky (1989).
Spatulata (Spatulata?) pachycraspedata[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Spatulata (Spatulata) tilsleyi[126]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Stenopareia kossovae[137]	Sp. nov	Valid	Krylov	Ordovician (Katian)		Russia
Syspacephalus mccollumorum[145]	Sp. nov	Valid	Sundberg	Cambrian	Emigrant Formation	United States	A member of Ptychopariida belonging to the group Ptychopariacea.
Tawstockia longispina germanica[126]	Subsp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany
Toxophacops (Atopophacops) fujiwara[146]	Sp. nov	Valid	Stocker et al.	Devonian (probably Givetian)	Naidaijin Formation	Japan
Trephina[125]	Gen. et sp. nov	Valid	Smith, Paterson & Brock	Cambrian	Goyder Formation	Australia	Genus includes new species T. ranfordi.
Tuleaspis[134]	Gen. et sp. nov	Valid	McAdams, Adrain & Karim	Early Ordovician		United States	The type species is T. jeneki.
Xylomelon[126]	Gen. et comb. et sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the subfamily Cystispininae. The type species is "Xenadoche" kalliroe Hahn, Hahn & Müller (1996); genus also includes X. agenor (Hahn, Hahn & Müller, 2000), X. elissa (Hahn, Hahn & Müller, 2000) and X. medusa (Hahn, Hahn & Müller, 1999), as well as new species X. xenios.

Other arthropods

Research

• A study on the external and internal anatomy and possible lifestyle of Waptia fieldensis, based on revision of all available specimens available from the repositories in the National Museum of Natural History and the Royal Ontario Museum, is published by Vannier et al. (2018).[147]
• A study on the lateral tail flexibility in the eurypterid Slimonia acuminata published by Persons & Acorn (2017)[148] is criticized by Lamsdell, Marshall & Briggs (2018);[149] Persons (2018) defends his original conclusions on the basis of a study of additional eurypterid specimens.[150]
• A study on the microstructure of gnathobasic spines on the coxa of extant Atlantic horseshoe crab, Silurian eurypterid Eurypterus tetragonophthalmus and Cambrian artiopodan Sidneyia inexpectans is published by Bicknell et al. (2018).[151]
• A study on the mechanical performance of the feeding appendages of Sidneyia inexpectans, based on comparisons with extant Atlantic horseshoe crab, is published by Bicknell et al. (2018).[152]
• Bicknell, Pates & Botton (2018) report abnormalities in specimens of extant (Limulus polyphemus, Tachypleus tridentatus) and fossil (Euproops danae, Mesolimulus walchi) horseshoe crabs, and compare injuries to horseshoe crab cephalothoraces to those affecting Cambrian trilobites to explore the possible timing of the injuries.[153]
• Sánchez-García et al. (2018) report the presence of probable male clasping organs for courtship and securing the female during copulation in the springtail Pseudosminthurides stoechus and an aggregation of up to 45 specimens of the springtail Proisotoma communis from the Cretaceous amber from Spain.[154]
• Closely associated arthropleurid trackways, interpreted as evidence of arthropleurid mating behaviour, are described from the Lower Carboniferous of Fife (Scotland, United Kingdom) by Whyte (2018).[155]
• Two immature specimens of polyxenidan millipedes are described from the Eocene Bitterfeld amber (Germany) by Haug et al. (2018).[156]
• First known millipede fossil material from Central America (disarticulated trunk segments of members of the genus Nyssodesmus from the late Pleistocene of Costa Rica) is described by Laurito & Valerio (2018).[157]
• Insect resting traces and arthropod trackways are described from the Eocene Mount Wawel Formation (King George Island, Antarctica) by Uchman, Gaździcki & Błażejowski (2018), who name a new ichnofamily Protichnidae and a new ichnospecies Glaciichnium australis.[158]
• Revision of blood-feeding arthropods from the Paleogene Baltic amber is published by Pielowska, Sontag & Szadziewski (2018).[159]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Adelophthalmus khakassicus[160]	Sp. nov	Valid	Shpinev & Filimonov	Devonian (Givetian)		Russia	An eurypterid.
Alutella siku[161]	Sp. nov	Valid	Peel & Willman	Cambrian Series 2	Buen Formation	Greenland	A member of Bradoriida.
Concavicaris viktoryni[162]	Sp. nov	Valid	Rak, Broda & Kumpan	Carboniferous		Czech Republic	A member of Thylacocephala (a group of arthropods of uncertain phylogenetic placement, possibly crustaceans). Originally described as a species of Concavicaris; Broda, Rak & Hegna (2020) made it the type species of a separate genus Paraconcavicaris.[163]
Eodiscus coloholcus[164]	Sp. nov	Valid	Westrop, Broda & Dengler	Cambrian	Shallow Bay Formation	Canada	A member of Agnostida belonging to the family Eodiscidae.
Eodiscus confossus[164]	Sp. nov	Valid	Westrop, Broda & Dengler	Cambrian	Manuels River Formation	Canada	A member of Agnostida belonging to the family Eodiscidae.
Eodiscus tuberculus[164]	Sp. nov	Valid	Westrop, Broda & Dengler	Cambrian	Manuels River Formation	Canada	A member of Agnostida belonging to the family Eodiscidae.
Gogglops[165]	Gen. et sp. nov	Valid	Siveter et al.	Ordovician		China	A member of Aglaspidida. Genus includes new species G. ensifer.
Isoxys globulus[166]	Sp. nov	Valid	Liu et al.	Cambrian Stage 4	Balang Formation	China
Isoxys jianheensis[166]	Sp. nov	Valid	Liu et al.	Cambrian Stage 4	Balang Formation	China
Messorocaris[167]	Gen. et sp. nov	Valid	Lerosey-Aubril & Skabelund	Cambrian (Drumian)	Wheeler Formation	United States	A possible relative of Sanctacaris. The type species is M. magna.
Misszhouia canadensis[168]	Sp. nov	Valid	Mayers, Aria & Caron	Wuliuan	Burgess Shale	Canada	A member of the family Naraoiidae.
Naraoia arcana[168]	Sp. nov	Valid	Mayers, Aria & Caron	Wuliuan	Burgess Shale	Canada	A member of the family Naraoiidae.
Naraoia magna[168]	Sp. nov	Valid	Mayers, Aria & Caron	Wuliuan	Burgess Shale	Canada	A member of the family Naraoiidae.
Pagetia aspinosa[145]	Sp. nov	Valid	Sundberg	Cambrian	Emigrant Formation	United States
Pagetia claytonensis[145]	Sp. nov	Valid	Sundberg	Cambrian	Emigrant Formation	United States
Permocrassacus[169]	Gen. et sp. nov	Valid	Khramov et al.	Permian (Kungurian)	Koshelevka Formation	Russia	A member of Scolopendromorpha of uncertain phylogenetic placement. The type species is P. novokshonovi.
Permocryptops[169]	Gen. et sp. nov	Valid	Khramov et al.	Late Permian	Poldarsa Formation	Russia	A member of Scolopendromorpha of uncertain phylogenetic placement. The type species is P. shelleyi.
Sinosoma[170]	Gen. et sp. nov	Valid	Huang et al.	Middle Triassic		China	A helminthomorph millipede sharing a number of characters with nematophorans. Genus includes new species S. luopingense.
Soligorskopterus[171]	Gen. et sp. nov	Valid	Plax et al.	Devonian (Famennian)	Soligorsk Formation	Belarus	A eurypterid. Genus includes new species S. tchepeliensis.
Symphylella patrickmuelleri[172]	Sp. nov	Valid	Moritz & Wesener	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Symphyla belonging to the family Scolopendrellidae.
Woodesmus[173]	Gen. et sp. nov	Valid	Ross, Edgecombe & Clark in Ross et al.	Carboniferous (Tournaisian)	Ballagan Formation	United Kingdom	A millipede belonging to the group Archipolypoda, assigned to the new family Woodesmidae. Genus includes new species W. sheari.
Xiphosuroides[174]	Gen. et sp. nov	Valid	Shpinev & Vasilenko	Carboniferous	Sarskaya Formation	Russia	Probably a xiphosuran belonging to the order Xiphosurida and the suborder Bellinurina. Genus includes new species X. khakassicus.

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84. Amanda Moreira Leite; Dermeval Aparecido Do Carmo; Caio Bussaglia Ress; Murilo Pessoa; Guilherme Miranda Caixeta; Matheus Denezine; Rodrigo Rodrigues Adorno; Lucas Silveira Antonietto (2018). "Taxonomy of limnic Ostracoda (Crustacea) from the Quiricó Formation, Lower Cretaceous, São Francisco basin, Minas Gerais State, Southeast Brazil". Journal of Paleontology. 92 (4): 661–680. doi:10.1017/jpa.2018.1.
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