2012 in arthropod paleontology

This list of fossil arthropods described in 2012 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2012. The list only includes taxa at the level of genus or species.

List of years in arthropod paleontology
In paleontology
2009
2010
2011
2012
2013
2014
2015

Arachnids

Name Novelty Status Authors Age Unit Location Notes Images

Ampezzoa[1]

Gen. et sp. nov.

Valid

Lindquist & Grimaldi in Schmidt et al.

Carnian

 Italy

A mite. The type species is Ampezzoa triassica.

Archaelagonops[2]

Gen. et sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Lagonomegopidae. Type species is A. salticoides.

Baltleucauge propinqua[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

A long-jawed orb weaver

Craspedisia yapchoontecki[4]

Sp. nov.

Valid

Penney et al.

Miocene

Dominican amber

 Dominican Republic

A theridiid

Eodoter lonimammillae[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

A sac spider

Eohalinobius calefactus[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

A succinomid

Eohalinobius hiddenseeensis[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

A succinomid

Eohalinobius patina[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

A succinomid

?Eophantes seorsum[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

A linyphiid

Ephalmator tredecim[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

An ephalmatorid

Ero (Succinero) clunis[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

A pirate spider

Ero (Succinero) gracilitibialis[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

A pirate spider

Ero (Succinero) veta[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

A pirate spider

Geratonephila[5]

Gen. et sp. nov.

jr synonym

Poinar in Poinar & Buckley

Early Cretaceous

Burmese amber

 Myanmar

A nephiline. The type species is G. burmanica.
Moved to Nephila in 2015.

Nephila burmanica

Graea magnocoli[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

Europe

An orb-weaver spider

Hypotheridiosoma[2]

Gen. et sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

Originally classified as a ray spider, but subsequently transferred to the araneoid family Zarqaraneidae.[6] The type species is H. paracymbium.

Korearachne[7]

gen et sp nov

Valid

Selden et al..

Early Cretaceous

Jinju Formation

 South Korea

A member of Araneomorphae, possibly a lycosoid. The type species is K. jinju.

Lagonoburmops[2]

Gen. et sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Lagonomegopidae. The type species is L. plumosus.

Leclercera longissipes[2]

Sp. nov

Jr synonym

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Ochyroceratidae. Originally described as a species of Leclercera, but subsequently transferred to the genus Priscaleclercera.[8]

Leclercera spicula[2]

Sp. nov

Jr synonym

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Ochyroceratidae. Originally described as a species of Leclercera, but subsequently transferred to the genus Priscaleclercera.[8]

Leviunguis[2]

Gen. et sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

Originally classified as a ray spider, subsequently transferred to the separate araneoid family Leviunguidae.[6] The type species is L. bruckschi.

Mesobunus dunlopi[9]

Sp. nov.

Valid

Giribet et al.

Middle Jurassic

Daohugou Beds

 China

A sclerosomatid harvestman.

Myanlagonops[2]

Gen. et sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Lagonomegopidae. The type species is M. gracilipes.

Ocululoborus[2]

Gen. et sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Uloboridae. The type species is O. curvatus.

Orchestina (Baltorchestina) angulata[10]

Nom. nov

Valid

Wunderlich

Eocene

Bitterfeld amber

 Germany

An oonopid; a replacement name for Orchestina (Baltorchestina) rectangulata Wunderlich (2011).

Orchestina gappi[11]

sp nov

Valid

Saupe et al..

Latest Albian or earliest Cenomanian

Chartense amber

 France

An oonopid

Orchestina rabagensis[11]

Species

Valid

Saupe et al.

Early Albian

 Spain

An oonopid, a species of Orchestina.

Palaeogrosphus jacquesi[12]

Sp. nov

Valid

Lourenço & Henderickx

Holocene

Malagasy copal

 Madagascar

A buthid scorpion.

Palaeoleptoneta[2]

Gen. et sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Leptonetidae. The type species is P. calcar.

Palaeolychas weitschati[13]

Sp. nov.

Valid

Lourenço

Eocene

 Russia (Kaliningrad Oblast)

A buthid scorpion, a species of Palaeolychas.

Palaeotrilineatus[14]

Gen. et sp. nov.

Valid

Lourenço

Early Cretaceous

Burmese amber

 Myanmar

A scorpion. The type species is P. ellenbergeri.

?Pholcochyrocer baculum[2]

Sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Pholcochyroceridae, possibly a species of Pholcochyrocer.

Pholcochyrocer pecten[2]

Sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Pholcochyroceridae, a species of Pholcochyrocer.

Piankhi[15]

Gen. et sp. nov.

Valid

Dunlop, Bartel & Mitov

Eocene

Baltic amber

Europe

A harvestman found in Baltic amber. The type species is P. steineri.

Piankhi steineri

?Poliochera cretacea[16]

Sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Ricinulei, possibly a species of Poliochera.

Praoppiella[17]

Gen. et sp. nov.

Valid

Miko & Mourek in Miko et al.

Early Pleistocene

 Slovenia

A mite. The type species is P. oanae.

Pseudogarypus synchrotron[18]

Sp. nov

Valid

Henderickx in Henderickx, Tafforeau & Soriano

Late Eocene

Baltic Amber

 Europe

A pseudoscorpion

?Psiloderces filiformis[2]

Sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

First described as possibly species of Psiloderces; transferred to the genus Eopsiloderces.[8]

Rhinoppioides[17]

Gen. et sp. nov.

Valid

Miko in Miko et al.

Early Pleistocene

 Slovenia

A mite. The type species is R. quadrituberculatus.

Saetosoma[2]

Gen. et sp. nov

Valid

Wunderlich

Cretaceous

Burmese amber

 Myanmar

A member of Tetrablemmidae. The type species is S. filiembolus.

Samlandicmeta[3]

Gen. et sp. nov

Valid

Wunderlich

Eocene

Baltic amber

 Europe

A long-jawed orb weaver. The type species is S. mutila.

?Scytodes hani[2]

Sp. nov

Valid

Wunderlich

Cretaceous

Lebanese amber

 Jordan

A spitting spider, possibly a species of Scytodes.

Succiniropsis runcinata[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

 Europe

A zoropsid.

?Succinomus gibbosus[3]

Sp. nov

Valid

Wunderlich

Eocene

Baltic amber

 Europe

A succinomid, possibly a species of Succinomus.

Trhypochthonius lopezvallei[19]

Sp. nov.

Valid

Arillo, Subías & Shtanchaeva

Early Cretaceous (Albian)

 Spain

A trhypochthoniid mite.

Triasacarus[1]

Gen. et sp. nov.

Valid

Lindquist & Grimaldi in Schmidt et al.

Carnian

 Italy

A mite. The type species is T. fedelei.

Crustaceans

Name Novelty Status Authors Age Unit Location Notes Images

Acanthocythereis inouei[20]

Sp. nov

Valid

Yamaguchi, Mashiba & Kamiya

Miocene

Osaki Formation

 Japan

An ostracod, a species of Acanthocythereis.

Acanthophoenicides[21]

Gen. et sp. nov

Valid

Audo & Charbonnier

Late Cretaceous (Cenomanian)

Hadjoula Lagerstätte

 Lebanon

A slipper lobster belonging to the subfamily Arctidinae. The type species is A. peterpani.

Acratia candyae[22]

Sp. nov

Valid

Forel

Late Permian (Changhsingian)

Wujiaping Formation

 China

An acratiid ostracod, a species of Acratia.

Agnocarcinus[23]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Pilumnidae. The type species is A. zannatoi.

Alberticarcinus[23]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Atelecyclidae. The type species is A. eocaenus.

Ammopylocheles robertboreki[24]

Sp. nov

Valid

Fraaije et al.

Middle Oxfordian

 Poland

A pylochelid, a species of Ammopylocheles.

Amphicytherura grandicribra[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A schizocytherid ostracod, a species of Amphicytherura.

Amphicytherura occulta[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

An ostracod. Originally described as a species of Amphicytherura; Antonietto et al. (2013) transferred this species to the genus Dinglecythere.[26]

Amphissites gifuensis[27]

Sp. nov

Valid

Tanaka & Yuan in Tanaka et al.

Early Permian

 Japan

An ostracod, a species of Amphissites.

Antonioranina[28]

Gen. et comb. nov

Valid

Van Bakel et al.

Eocene

 Italy
 Spain
 United States

A raninid crab. The type species is "Cyrtorhina" globosa Beschin, Busulini, De Angeli & Tessier (1988); genus also contains A. fusseli (Blow & Manning, 1996) and A. ripacurtae (Artal & Castillo, 2005). Van Bakel et al. (2012) tentatively assigned "Cyrtorhina" oblonga Beschin, Busulini, De Angeli & Tessier (1988) to the genus Antonioranina as well;[28] however, Karasawa et al. (2014) made it the type species of a separate genus Claudioranina.[29]

Archaeoniscus coreaensis[30]

Sp. nov

Valid

Park et al.

Early Cretaceous

Jinju Formation

 South Korea

An isopod crustacean, a species of Archaeoniscus.

Asciocythere cabbagehillensis[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A cytherideid ostracod, a species of Asciocythere.

Asymmetricythere semoupathei[31]

Sp. nov

Valid

Sarr

Ypresian to Lutetian

 Mauritania
 Senegal

A trachyleberidid ostracod, a species of Asymmetricythere.

Aurikirkbya miyakei[27]

Sp. nov

Valid

Tanaka & Ono in Tanaka et al.

Early Permian

 Japan

An ostracod, a species of Aurikirkbya.

Aysegulina chapeltonensis[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A hemicytherid ostracod, a species of Aysegulina.

Aysegulina riominhoensis[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A hemicytherid ostracod, a species of Aysegulina.

Aysegulina sagitta[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A hemicytherid ostracod, a species of Aysegulina.

Aysegulina ventrocurva[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A hemicytherid ostracod, a species of Aysegulina.

Bairdia adelineae[22]

Sp. nov

Valid

Forel

Early Triassic (Griesbachian)

Daye Formation

 China

A bairdiid ostracod, a species of Bairdia.

Bairdia fujisan[27]

Sp. nov

Valid

Tanaka & Maeda in Tanaka et al.

Early Permian

 Japan

An ostracod, a species of Bairdia.

Bairdia? huberti[22]

Sp. nov

Valid

Forel

Late Permian (Changhsingian)

Wujiaping Formation

 China

A bairdiid ostracod, a possible species of Bairdia.

Bairdia ikeyai[27]

Sp. nov

Valid

Tanaka in Tanaka et al.

Early Permian

 Japan

An ostracod, a species of Bairdia.

Bairdia jeromei[22]

Sp. nov

Valid

Forel

Late Permian (Changhsingian) and Early Triassic (Griesbachian)

Daye Formation
Wujiaping Formation

 China

A bairdiid ostracod, a species of Bairdia.

Bairdia minoensis[27]

Sp. nov

Valid

Tanaka & Ono in Tanaka et al.

Early Permian

 Japan

An ostracod, a species of Bairdia.

Bairdia spindlica[27]

Sp. nov

Valid

Tanaka & Yuan in Tanaka et al.

Early Permian

 Japan

An ostracod, a species of Bairdia.

Balowella crassicostata[32]

Sp. nov

Valid

Luppold

Middle Jurassic

 Germany

An ostracod, a species of Balowella.

Bathypluma pliocenica[33]

Sp. nov

Valid

Garassino, Pasini & Marini

Early Pliocene

 Italy

A retroplumid crab, a species of Bathypluma.

Bournelyreidus[28]

Gen. et comb. et sp. nov

Valid

Van Bakel et al.

Late Cretaceous to Paleocene

 Denmark
 France
 Germany
 Greenland
 United States

A lyreidid crab. The type species is "Hemioon" eysunesensis Collins & Wienberg Rasmussen (1992); according to the authors of its description genus also contains "Raninella" carlilensis Feldmann & Maxey (1980), "Palaeocorystes" laevis Schlüter in von der Marck & Schlüter (1868), "Raninella" manningi Bishop & Williams (2000), "Raninella" oaheensis Bishop (1978) and "Raninella" tridens Roberts (1962), as well as the new species Bournelyreidus teodorii.[28] On the other hand, Karasawa et al. (2014) transferred B. teodorii to the genus Lyreidus, "R." tridens to the genus Macroacaena and "R." manningi to the genus Heus.[29]

Brachycythere jamaicaensis[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A trachyleberidid ostracod, a species of Brachycythere.

Bretonia[34]

gen et comb nov

Valid

Schweitzer et al.

Cenomanian

Sables du Perche Formation

 France

crab, New genus for "Xanthosia" danielae (Collins and Breton, 2009).

Buntonia bodergati[31]

Sp. nov

Valid

Sarr

Thanetian to Ypresian

 Mali
 Senegal

A trachyleberidid ostracod, a species of Buntonia.

Buntonia nana[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A trachyleberidid ostracod, a species of Buntonia.

Buntonia vulgaris[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A trachyleberidid ostracod, a species of Buntonia.

Bythocypris antoniettae[35]

Sp. nov

Valid

Sciuto

Early Pleistocene

 Italy

An ostracod, a species of Bythocypris.

Bythocythere solisdeus[36]

Sp. nov

Valid

Sciuto

Early Pleistocene

 Italy

A bythocytherid ostracod, a species of Bythocythere.

Calliaxina altavillensis[37]

Sp. nov

Valid

Beschin & De Angeli

Eocene

 Italy

A member of Callianassidae.

Carcinoplax parsensis[38]

Sp. nov

Valid

Vega et al.

Miocene (Burdigalian or Langhian)

Mishan Formation

 Iran

A goneplacid decapod crustacean, a species of Carcinoplax.

Carpopenaeus garassinoi[39]

Sp. nov

Valid

Charbonnier in Petit & Charbonnier

Late Cretaceous

Sahel Alma Lagerstätte

 Lebanon

Cenocorystes bretoni[28]

Sp. nov

Valid

Van Bakel et al.

Late Cretaceous (late Cenomanian)

 France

A palaeocorystid crab, a species of Cenocorystes.

Cenomanocarcinus cantabricus[28]

Sp. nov

Valid

Van Bakel et al.

Early Cretaceous (Albian)

 Spain

A cenomanocarcinid crab, a species of Cenomanocarcinus.

Coleia martinlutheri[40]

sp nov

Valid

Schweigert & Ernst

Sinemurian

 Germany

A coleiid polychelid, a species of Coleia.

Cretacolana[41]

Gen. et comb. nov

Valid

Schweitzer & Feldmann

Cenomanian

 France

A porcelain crab, a new genus for "Porcellana" antiqua A. Milne-Edwards, 1862.

Cretatrizocheles[42]

gen et sp nov

Valid

Fraaije, Klompmaker, & Artal

Albian or Cenomanian

 Spain

A hermit crab. The type species is Cretatrizocheles olazagutiensis.

Cromimaia[23]

Gen. et comb. nov

Valid

Beschin et al.

Eocene

 Italy

A crab belonging to the family Majidae. The type species is "Micromaia" meneguzzoi Beschin, Busulini, De Angeli & Tessier (1985); genus also includes "Micromaia" elegans Beschin, Busulini, De Angeli & Tessier (1985).

Cushmanidea tessieri[31]

Sp. nov

Valid

Sarr

Ypresian to Lutetian

 Senegal

A cushmanideid ostracod, a species of Buntonia.

Cytherelloidea guineacornensis[25]

Sp. nov

Valid

Puckett in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

Guinea Corn Formation

 Jamaica

A cytherellid ostracod, a species of Cytherelloidea.

Cytheropteron eleonorae[36]

Sp. nov

Valid

Sciuto

Early Pleistocene

 Italy

A cytherurid ostracod, a species of Cytheropteron.

Cytheropteron italoi[35]

Sp. nov

Valid

Sciuto

Early Pleistocene

 Italy

An ostracod, a species of Cytheropteron.

Cytheropteron rossanae[35]

Sp. nov

Valid

Sciuto

Early Pleistocene

 Italy

An ostracod, a species of Cytheropteron.

Daira coccoi[23]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Dairidae; a species of Daira.

Distefania renefraaijei[43]

Sp. nov

Valid

Klompmaker, Feldmann & Schweitzer

Cretaceous (Albian or Cenomanian)

 Spain

A goniodromitid crab, a species of Distefania.

Dolocythere tuberculata[32]

Sp. nov

Valid

Luppold

Middle Jurassic

 Germany

An ostracod. Originally described as a species of Dolocythere; Lord, Cabral & Danielopol (2020) transferred this species to the genus Minyocythere.[44]

Ebullitiocaris elatus[45]

Sp. nov

Valid

Womack et al.

Carboniferous

 United Kingdom

A member of Cladocera, a species of Ebullitiocaris.

Eocytheropteron hazeli[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A cytherurid ostracod, a species of Eocytheropteron.

Eocytheropteron jamaicaensis[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A cytherurid ostracod, a species of Eocytheropteron.

Eouroptychus[46]

Gen. et sp. nov

Valid

De Angeli & Ceccon

Eocene (Ypresian)

 Italy

A member of Chirostylidae. The type species is E. montemagrensis.

Eucytherura mitchelli[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A cytherurid ostracod, a species of Eucytherura.

Eomunidopsis aldoirarensis[47]

Sp. nov

Valid

Klompmaker et al.

Cretaceous (Albian or Cenomanian)

 Spain

A galatheoid, a species of Eomunidopsis.

Eopaguropsis nidiaquilae[48]

Sp. nov

Valid

Fraaije et al.

Oxfordian

 Poland

A diogenid, a species of Eopaguropsis.

Eosymethis[28]

Gen. et sp. nov

Valid

Van Bakel et al.

Eocene (Ypresian)

Roda Formation

 Spain

A raninid crab. The type species is Eosymethis aragonensis.

Eohexapus orthogonius[23]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Hexapodidae.

Eucorystes iserbyti[28]

Sp. nov

Valid

Van Bakel et al.

Early Cretaceous (Albian)

 France

A palaeocorystid crab, a species of Eucorystes.

Eucorystes navarrensis[28]

Sp. nov

Valid

Van Bakel et al.

Early Cretaceous (Albian)

 Spain

A palaeocorystid crab, a species of Eucorystes.

Faksecarcinus[34]

gen et comb nov

Valid

Schweitzer et al.

Albian or Cenomanian to Danian

 Denmark  Spain

A crab, a new genus for "Xanthosia" gracilis.

Ferroranina[28]

Gen. et comb. et sp. nov

Valid

Van Bakel et al.

Late Cretaceous (late Cenomanian to late Campanian)

 Czech Republic[49]
 India
 Madagascar
 Mexico
 United States

A palaeocorystid crab. The type species is "Notopocorystes" dichrous Stenzel (1945); genus also contains "Notopocorystes" australis Secretan (1964) and "Notopocorystes" fritschi Glaessner (1929),[49] as well as the new species Ferroranina tamilnadu.

Floricythereis[25]

Gen. et sp. et comb. nov

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica
 United States

A trachyleberidid ostracod. Genus includes new species Floricythereis exquisita, as well as F. lixula (Crane, 1965) (the type species was not designated).

Galathea mainensis[50]

Sp. nov

Valid

Ceccon & De Angeli

Middle Eocene

 Italy

A species of Galathea.

Gebiacantha tuscia[51]

Sp. nov

Valid

Garassino, Pasini, De Angeli & Charbonnier in Garassino et al.

Early Pliocene

 Italy

An upogebiid decapod, a species of Gebiacantha.

Geffenina mariebeatriceae[52]

Sp. nov

Valid

Chitnarin in Chitnarin et al.

Late Early Permian

Tak Fa Formation

 Thailand

A knoxitid palaeocopidan ostracod, a species of Geffenina.

Geffenina mariebeatriceae[52]

Sp. nov

Valid

Chitnarin in Chitnarin et al.

Early Permian

Pha Nok Khao Formation

 Thailand

A knoxitid palaeocopidan ostracod, a species of Geffenina.

Glyphea wiffenae[53]

Species

Valid

Feldmann and Stilwell

Late Cretaceous

Maungataniwa Sandstone

 New Zealand

A glypheid lobster, a species of Glyphea.

Glyptopleura mugiensis[27]

Sp. nov

Valid

Tanaka & Yuan in Tanaka et al.

Early Permian

 Japan

An ostracod, a species of Glyptopleura.

Goniodromites kubai[54]

sp nov

Valid

Starzyk, Krzemiska, & Krzemiski

Oxfordian

 Poland

A crab, a species of Goniodromites.

Gonioplacoides[55]

Gen. et sp. nov

Valid

Quayle & Collins

Eocene

Headon Hill Formation

 United Kingdom

A goneplacid. The type species is Gonioplacoides minuta.

Graptocarcinus urbasaensis[56]

Sp. nov

Valid

Van Bakel et al.

Late Cretaceous

 Spain

A dynomenid crab, a species of Graptocarcinus.

Grinioneis pachycosta[57]

Sp. nov

Valid

Guernet et al.

Eocene (Lutetian)

Paris Basin

 France

A trachyleberidid ostracod, a species of Grinioneis.

Grolamaia[23]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the subfamily Pisinae. The type species is G. vicariottoi.

Hadrocarcinus[58]

gen et comb et sp nov

Valid

Schweitzer, Feldmann & Lamanna

Late Cretaceous

Antarctica

A necrocarcinid crab, a new genus for "Necrocarcinus" wrighti. The genus also contains "Necrocarcinus" carinatus and a new species Hadrocarcinus tectilacus.

Harbinia alta[59]

Sp. nov

Valid

Antonietto et al.

Aptian or early Albian

Santana Formation

 Brazil

An ostracod crustacean, a species of Harbinia.

Headonipus[55]

Gen. et sp. nov

Valid

Quayle & Collins

Late Eocene

Headon Beds

 United Kingdom

A hexapodid. The type species is Headonipus tuberculatus.

Hemiparacytheridea exquisita[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A paracytherideid ostracod, a species of Hemiparacytheridea.

Hispanigalathea[47]

Gen. et 2 sp. nov

Valid

Klompmaker et al.

Cretaceous (Albian or Cenomanian)

 Spain

A galatheoid. Genus contains two species: Hispanigalathea pseudolaevis and H. tuberosa.

Huanghestheria linxiensis[60]

Sp. nov

Valid

Niu in Zhang et al.

Late Permian

Linxi Formation

 China

A vertexiine lioestheriid spinicaudatan clam shrimp, a species of Huanghestheria.

Joeranina[28]

Gen. et comb. et sp. nov

Valid

Van Bakel et al.

Cretaceous (Albian to early Campanian)

 Canada
 France
 Greenland
 Japan
 Madagascar
 Spain
  Switzerland
 Syria
 United Kingdom
 United States

A palaeocorystid crab. The type species is "Corystes" broderipii Mantell (1844). According to the authors of its description genus also contains "Palaeocorystes" harveyi Woodward (1896), "Eucorystes" japonicus Jimbô (1894), "Notopocorystes" paututensis Collins & Wienberg Rasmussen (1992), "Eucorystes" platys Schweitzer & Feldmann (2002) and "Notopocorystes" syriacus Withers (1928), as well as the new species Joeranina gaspari.[28] On the other hand, Karasawa et al. (2014) transferred "P." harveyi, E. platys and "N." paututensis to the genus Eucorystes; the authors also transferred the species "Notopocorystes" xizangensis Wang (1981) to the genus Joeranina.[29]

Juracrista[61]

Gen. et 2 sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidid. The type species is Juracrista perculta; genus also contains Juracrista costaspinosa.

Jurapylocheles iwonae[24]

Sp. nov

Valid

Fraaije et al.

Middle Oxfordian

 Poland

A pylochelid, a species of Jurapylocheles.

Kinkelinella circumcostata[32]

Sp. nov

Valid

Luppold

Middle Jurassic

 Germany

An ostracod, a species of Kinkelinella.

Koryncheiros[62]

Gen. et sp. nov

Valid

Feldmann et al.

Middle or late Anisian

Guanling Formation

 China

A decapod crustacean; a clytiopsid erymoid. The type species is Koryncheiros luopingensis.

Kroemmelbeincypris[63]

Gen. et comb. nov

Valid

Poropat & Colin

Aptian

An ostracod crustacean, a new genus for the species "Hourcqia" symmetrica Krömmelbein & Weber and "Hourcqia" angulata Krömmelbein & Weber.

Langdaia meesooki[52]

Sp. nov

Valid

Chitnarin in Chitnarin et al.

Late Early to Middle Permian

Tak Fa Formation

 Thailand

A knoxitid palaeocopidan ostracod, a species of Langdaia.

Leiocythereis[25]

Gen. et sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A trachyleberidid ostracod. The type species is Leiocythereis polita.

Lepas sattmanni[64]

Sp. nov

Valid

Harzhauser & Schlögl

Early Miocene

Ebelsberg Formation

 Austria

A lepadid barnacle, a species of Lepas.

Lessinicarcinus[65]

Gen. et comb. nov

Valid

De Angeli

Eocene

 Croatia[66]
 Italy

A crab belonging to the family Pilumnidae. The type species is "Titanocarcinus" euglyphos Bittner (1875).

Lessinipagurus[23]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A hermit crab. Originally assigned to the family Paguridae,[23] subsequently transferred to the separate family Xylopaguridae by Gašparič et al. (2016).[67] The type species is L. granulatus.

Lienenklausicythere[68]

Gen. et comb. nov

Valid

Uffenorde, Monostori & Tóth

Eocene to Oligocene

Europe

An ostracod, a new genus for "Hermanites" acuticosta and "Hermanites" camela; genus also contains a new, yet-unnamed species.

Ljubimovella debilicostatus[32]

Sp. nov

Valid

Luppold

Middle Jurassic

 Germany

An ostracod, a species of Ljubimovella.

Lobonotus beschini[69]

Sp. nov

Valid

De Angeli & Checchi

Eocene (Lutetian)

 Italy

A crab belonging to the family Tumidocarcinidae.

Loxoconcha pindarsensis[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A loxoconchid ostracod, a species of Loxoconcha.

Marylyreidus[28]

Gen. et comb. nov

Valid

Van Bakel et al.

Cretaceous (late Albian–early Cenomanian)

Pawpaw Formation

 United States

A lyreidid crab. The type species is "Notopocorystes" punctatus Rathbun (1935).

Megokkos patagoniensis[70]

Sp. nov

Valid

Schweitzer et al.

Eocene

Rio Turbio Formation

 Argentina

A crab, a species of Megokkos.

Mesoparapylocheles[42]

gen et sp nov

Valid

Fraaije, Klompmaker, & Artal

Albian or Cenomanian

 Spain

A hermit crab. The type species is Mesoparapylocheles michaeljacksoni.

Microcheilinella subovata[27]

Sp. nov

Valid

Tanaka & Maeda in Tanaka et al.

Early Permian

 Japan

An ostracod, a species of Microcheilinella.

Microcheilinella tsubogawensis[27]

Sp. nov

Valid

Tanaka & Ichida in Tanaka et al.

Early Permian

 Japan

An ostracod, a species of Microcheilinella.

Microcoelonella takfaensis[52]

Sp. nov

Valid

Chitnarin in Chitnarin et al.

Middle Permian

Tak Fa Formation

 Thailand

A coelonellid palaeocopidan ostracod, a species of Microcoelonella.

Microcoelonella takliensis[52]

Sp. nov

Valid

Chitnarin in Chitnarin et al.

Middle Permian

Tak Fa Formation

 Thailand

A coelonellid palaeocopidan ostracod, a species of Microcoelonella.

Munida deangelii[71]

Sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Oligocene to early Miocene

Río Foyel Formation

 Argentina

A munidid, a species of Munida.

Myra gurii[38]

Sp. nov

Valid

Vega et al.

Miocene (Burdigalian or Langhian)

Mishan Formation

 Iran

A leucosiid decapod crustacean, a species of Myra.

Navarradromites[43]

Gen. et sp. nov

Valid

Klompmaker, Feldmann & Schweitzer

Cretaceous (Albian or Cenomanian)

 Spain

A goniodromitid crab. The type species is Navarradromites pedroartali.

Navarrahomola[72]

Gen. et sp. nov.

Valid

Artal et al.

Cretaceous (Albian or Cenomanian)

 Spain

A homolid crab. The type species is Navarrahomola hispanica.

Neocallichirus bacatus[73]

Sp. nov

Valid

Busulini, Beschin & Tessier

Eocene (Priabonian)

 Italy

A member of Callianassidae.

Neocytherideis labyrinthoidea[57]

Sp. nov

Valid

Guernet et al.

Eocene (Lutetian)

Paris Basin

 France

A neocytherideid ostracod, a species of Neocytherideis.

Neocytheromorpha priscipacifica[20]

Sp. nov

Valid

Yamaguchi, Mashiba & Kamiya

Miocene

Osaki Formation

 Japan

An ostracod, a species of Neocytheromorpha.

Neurocythere jakovlevae[74]

Sp. nov

Valid

Kolpenskaya in Tesakova et al.

Late Jurassic (Early Kimmeridgian)

 Russia

An ostracod, a species of Neurocythere.

Nitotacarcinus antipodes[70]

Sp. nov

Valid

Schweitzer et al.

Eocene

Rio Turbio Formation

 Argentina

A crab, a species of Nitotacarcinus.

Nodosculda[75]

Gen. et sp. nov

Valid

Franţescu

Early Cretaceous (late Albian)

 United States

A mantis shrimp. The type species is Nodosculda fisherorum.

Noetlingocarcinus messinai[23]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the superfamily Dromioidea and the family Basinotopidae, a species of Noetlingocarcinus.

Nogarhomola[76]

Gen. et sp. nov

Valid

De Angeli & Alberti

Eocene (Lutetian)

 Italy

A homolid crab. The type species is Nogarhomola aurorae.

Notopocorystes kerri[77]

Sp. nov

Valid

Luque et al.

Early Cretaceous (late Aptian)

Paja Formation

 Colombia

A palaeocorystid raninoid crab. Initially described as a species of Notopocorystes,[77] but subsequently reassigned to the genus Joeranina.[78]

Nykteripteryx[47]

Gen. et sp. nov

Valid

Klompmaker et al.

Cretaceous (Albian or Cenomanian)

 Spain

A galatheoid. The type species is Nykteripteryx rostrata.

Ochtholambrus gambillarai[73]

Sp. nov

Valid

Busulini, Beschin & Tessier

Eocene (Priabonian)

 Italy

A crab belonging to the family Parthenopidae.

Odontodactylus italicus[23]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A mantis shrimp belonging to the family Odontodactylidae; a species of Odontodactylus.

Ommaciria[23]

Gen. et comb. nov

Valid

Beschin et al.

Eocene

 Italy

A crab belonging to the family Majidae. The type species is "Micromaia" mainensis Beschin, Busulini, De Angeli & Tessier (1985).

Orthobairdia jeanlouisi[22]

Sp. nov

Valid

Forel

Early Triassic (Griesbachian)

Daye Formation

 China

A bairdiid ostracod, a species of Orthobairdia.

Ovocytheridea rotunda[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A cytherideid ostracod, a species of Ovocytheridea.

Pagurus valdagnensis[23]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A hermit crab belonging to the family Paguridae, a species of Pagurus.

Palaega picena[79]

Sp. nov

Valid

Pasini & Garassino

Miocene

 Italy

A cirolanid isopod, a species of Palaega.

Palaega pisana[80]

Sp. nov

Junior synonym

Pasini & Garassino

Pliocene

 Italy

A cirolanid isopod. Originally described as a species of Palaega; considered to be a junior synonym of Bathynomus sismondai (Ristori, 1891) by Hyžný, Pasini & Garassino (2019).[81]

Palaega steatopigia[82]

Sp. nov

Nomen dubium

Pasini & Garassino

Pliocene

 Italy

A cirolanid isopod. Originally described as a species of Palaega; Hyžný, Pasini & Garassino (2019) considered this species to be a nomen dubium and reassigned it to the genus Bathynomus.[81]

Palaeohomarus pacificus[83]

Sp. nov

Valid

Aguirre-Urreta, Lazo & Rawson

Late Hauterivian or early Barremian

Agrio Formation

 Argentina

A decapod crustacean, a species of Palaeohomarus.

Paracypris keiji[57]

Sp. nov

Valid

Guernet et al.

Eocene (Lutetian)

Paris Basin

 France

A candonid ostracod, a species of Paracypris.

Paranecrocarcinus balla[28]

Sp. nov

Valid

Van Bakel et al.

Late Cretaceous (Cenomanian)

 France

A necrocarcinid crab, a species of Paranecrocarcinus.

Periosocypris[84]

Gen. et sp. nov

Valid

Whatley, Khosla & Rathore

Late Cretaceous (Maastrichtian)

Lameta Formation

 India

A cypridid ostracod. The type species is Periosocypris megistus.

Petrochirus savii[23]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A hermit crab belonging to the family Diogenidae, a species of Petrochirus.

Phrixocythere[25]

Gen. et sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A hemicytherid ostracod. The type species is Phrixocythere unionensis.

Planocarcinus[77]

Gen. et comb. nov

Valid

Luque et al.

Early Cretaceous (late Aptian)

Paja Formation

 Colombia

A necrocarcinid raninoid crab, a new genus for "Dakoticancer" olssoni Rathbun (1937).

Podocallichirus laepaensis[85]

Sp. nov

Valid

Hyžný & Muñiz

Late Miocene

 Spain

A callianassid ghost shrimp. Originally described as a species of Podocallichirus; subsequently transferred to the genus Balsscallichirus by Hyžný (2016).[86]

Protobuntonia warorei[31]

Sp. nov

Valid

Sarr

Paleocene to Ypresian

 Senegal
 Togo

A trachyleberidid ostracod, a species of Buntonia.

Pterygocythereis babinoti[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A trachyleberidid ostracod, a species of Pterygocythereis.

Pylochelitergites[87]

Gen. et 2 sp. nov

Valid

Fraaije et al.

Oxfordian to Kimmeridgian

 Germany
 Poland

A pylochelid hermit crab. Genus contains two species: P. westerbergensis (the type species) and P. gelasinus.

Quasilaeviranina minuscula[23]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Raninidae, a species of Quasilaeviranina.

Raninoides rioturbiensis[70]

Sp. nov

Valid

Schweitzer et al.

Eocene

Rio Turbio Formation

 Argentina

A crab, a species of Raninoides.

Rhinolambrus elongatus[23]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Parthenopidae; a species of Rhinolambrus.

Ristoria[51]

Gen. et comb. nov

Valid

Garassino, Pasini, De Angeli & Charbonnier in Garassino et al.

Early Pliocene

 Italy

A leucosiid crab, a new genus for "Ilia" pliocaenica Ristori, 1891.

Samarella sonei[52]

Sp. nov

Valid

Chitnarin in Chitnarin et al.

Middle Permian

Tak Fa Formation

 Thailand

A paraparchitid palaeocopidan ostracod, a species of Samarella.

Samarella viscusforma[52]

Sp. nov

Valid

Chitnarin in Chitnarin et al.

Early to Middle Permian

Pha Nok Khao Formation
Tak Fa Formation

 Thailand

A paraparchitid palaeocopidan ostracod, a species of Samarella.

Sargentina chantarameei[52]

Sp. nov

Valid

Chitnarin in Chitnarin et al.

Early to Middle Permian

Pha Nok Khao Formation
Tak Fa Formation

 Thailand

A knoxitid palaeocopidan ostracod, a species of Sargentina.

Scalpellum paratethyianum[64]

Sp. nov

Valid

Harzhauser & Schlögl

Early Miocene

Lakšárska Nová Ves Formation

 Slovakia

A scalpellid barnacle, a species of Scalpellum.

Schizoptocythere nana[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A trachyleberidid ostracod, a species of Schizoptocythere.

Schizoptocythere pinna[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A trachyleberidid ostracod, a species of Schizoptocythere.

Schuleridea exigua[25]

Sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica

A cytherideid ostracod, a species of Schuleridea.

Semicytherura carioui[31]

Sp. nov

Valid

Sarr

Ypresian to Lutetian

 Senegal

A cytherurid ostracod, a species of Semicytherura.

Sineruga[88]

gen et sp nov

Valid

  • Perrier

Silurian (uppermost Ludlow to lower Pridoli)

Le Val Formation

 France

myodocope ostracod. type species is Sineruga insolita.

Southcavea (Saxosouthcavea)[32]

Subgen. et 3 sp. nov

Valid

Luppold

Middle Jurassic

 Germany

An ostracod, a subgenus of Southcavea containing three new species: Southcavea (Saxosouthcavea) extendata, Southcavea (Saxosouthcavea) bartenbrandi and Southcavea (Saxosouthcavea) malzi (with two subspecies: S. (S.) malzi malzi and S. (S.) malzi reticulata).

Spinicytheridea[25]

Gen. et sp. nov

Valid

Puckett & Colin in Puckett, Colin & Mitchell

Late Cretaceous (Maastrichtian)

 Jamaica
 Mexico

A cytherideid ostracod. The type species is Spinicytheridea compta from Jamaica; genus also includes a second, unnamed species from the Maastrichtian Ocozocoautla Formation of Chiapas, Mexico.

Spinirostrimaia[23]

Gen. et comb. nov

Valid

Beschin et al.

Eocene

 Italy
 Spain

A crab belonging to the family Majidae. The type species is "Micromaia" margaritata Fabiani (1910).

Stagmacaris adielklompmakeri[87]

Sp. nov

Valid

Fraaije et al.

Early late Kimmeridgian

 Germany

A pylochelid hermit crab, a species of Stagmacaris.

Stagmacaris biarcus[87]

Sp. nov

Valid

Fraaije et al.

Middle Oxfordian

 Poland

A pylochelid hermit crab, a species of Stagmacaris.

Steorrosia[34]

gen et comb nov

Valid

Schweitzer et al.

Late Albian to Cenomanian

 United States

Crab, New genus for "Xanthosia" aspera. Other species S. pawpawensis and S. reidi.

Styracocarcinus[41]

Gen. et comb. nov

Valid

Schweitzer & Feldmann

Late Cretaceous (Senonian)

 Morocco

A tumidocarcinid decapod, a new genus for "Titanocarcinus" meridionalis Secretan, 1961.

Superpendicythere[32]

Gen. et sp. nov

Valid

Luppold

Middle Jurassic

 Germany

An ostracod. The type species is S. binodosa.

Symethoides[28]

Gen. et sp. nov

Valid

Van Bakel et al.

Paleocene (early Danian)

Tinton Formation

 United States

A raninid crab. The type species is Symethoides monmouthorum.

Szaboa deppermanni[89]

Sp. nov

Valid

Fraaije et al.

Oligocene

Ratzeburg Formation

 Germany

A matutid crab, a species of Szaboa.

Thalamita italica[51]

Sp. nov

Valid

Garassino, Pasini, De Angeli & Charbonnier in Garassino et al.

Early Pliocene

 Italy

A portunid crab, a species of Thalamita.

Tridactylastacus[62]

Gen. et sp. nov

Valid

Feldmann et al.

Middle or late Anisian

Guanling Formation

 China

A decapod crustacean; a glypheoid belonging to the family Litogastridae.[90] The type species is Tridactylastacus sinensis.

Turbiocheir[70]

Gen. et sp. nov

Valid

Schweitzer et al.

Eocene

Rio Turbio Formation

 Argentina

A thalassinidean. The type species is Turbiocheir minutospinata.

Vecticallichirus[55]

Gen. et sp. nov

Valid

Quayle & Collins

Late Eocene and Oligocene[91]

Headon Beds

 United Kingdom

A callichirid[91] ghost shrimp. The type species is Vecticallichirus abditus. Hyžný (2020) considered V. abditus to be a junior synonym of "Callianassa" batei Woodward (1869), resulting in a new combination Vecticallichirus batei.[91]

Vegaranina[28]

Gen. et comb. nov

Valid

Van Bakel et al.

Late Cretaceous (Campanian–Maastrichtian)

 Cuba
 France
 Mexico
 United States ( Puerto Rico)

A raninid crab. The type species is "Lophoranina" precocious Feldmann, Vega, Tucker, García-Barrera & Avendaño (1996).

Vernoniella tenuipora[32]

Sp. nov

Valid

Luppold

Middle Jurassic

 Germany

An ostracod, a species of Vernoniella.

Viaia[72]

Gen. et sp. nov.

Valid

Artal et al.

Cretaceous (Albian or Cenomanian)

 Spain

A viaiid glaessneropsoid crab. The type species is Viaia robusta.

Yunnanopalinura[62]

Gen. et sp. nov

Valid

Feldmann et al.

Middle or late Anisian

Guanling Formation

 China

A decapod crustacean; a spiny lobster. The type species is Yunnanopalinura schrami.

Insects

Blattodea

Name Novelty Status Authors Age Unit Location Notes Images

Archimesoblatta shiva[92]

Sp. nov

Valid

Engel & Pérez-de La Fuente

Middle Jurassic

Kota Formation

 India

A mesoblattinid cockroach

Arcofuzia[93]

Gen. et sp. nov

Valid

Wei, Shih & Ren

Middle Jurassic

Jiulongshan Formation

 China

A fuziid cockroach. The type species is Arcofuzia cana.

Cariblattoides labandeirai[94]

Sp. nov

Valid

Vršanský et al.

Eocene

Green River Formation

 United States

A blattellid cockroach

Divocina[95]

Gen. et sp. nov.

Valid

Liang, Vršanský & Ren

Middle Jurassic (Bathonian)

Jiulongshan Formation

 China

A raphidiomimid cockroach. The type species is Divocina noci.

Entropia[96]

Gen. et sp. nov

Valid

Vršanský, Liang & Ren

Middle Jurassic (Bathonian)

 China

A liberiblattinid cockroach. The type species is Entropia initialis.

Graciliblatta[97]

Gen. et sp. nov

Valid

Liang, Huang & Ren

Middle Jurassic

Jiulongshan Formation

 China

A raphidiomimid cockroach. The type species is Graciliblatta bella.

Mutovia[98]

Gen. et sp. nov

Valid

Vršanský & Aristov

Late Permian

 Russia

A mutoviid cockroach. The type species is Mutovia intercalaria.

Parvifuzia peregrina[99]

Sp. nov

Valid

Wei, Liang & Ren

Middle Jurassic

Jiulongshan Formation

 China

A fuziid cockroach, a species of Parvifuzia.

Qilianiblatta[100]

Sp. nov

Valid[101]

Zhang, Schneider & Hong

Carboniferous (earliest Pennsylvanian)

 China

A cockroach. The type species is Qilianiblatta namurensis.

Stegoblatta[102]

Gen. et sp. nov

Valid

Anisyutkin & Gröhn

Eocene

Baltic amber

Europe

A blaberid cockroach. The type species is Stegoblatta irmgardgroehni.

Coleoptera

Name Novelty Status Authors Age Unit Location Notes Images
Abrocar concavus[103]

Sp. nov

Valid[104]

Davis et al.

Early Cretaceous

Yixian Formation

 China

A weevil, a species of Abrocar.

Abrocar relicinus[103]

Sp. nov

Valid[104]

Davis et al.

Early Cretaceous

Yixian Formation

 China

A weevil, a species of Abrocar.

Abrohadeocoleodes[105]

Gen. et 4 sp. nov

Valid[106]

Tan et al.

Middle Jurassic

Jiulongshan Formation

 China

A schizophorid archostematan beetle. Genus contains four species: A. eurycladus, A. ooideus, A. nii and A. patefactus.

Alloioscarabaeus[107]

Gen. et sp. nov

Valid

Bai, Ren & Yang in Bai et al.

Middle Jurassic

Jiulongshan Formation

 China

An alloioscarabaeid scarabaeoid beetle. The type species is Alloioscarabaeus cheni.

Alphitopsis[108]

Gen. et sp. nov

Valid

Kirejtshuk, Nabozhenko & Nel

Early Cretaceous

Yixian Formation

 China

A tenebrionine darkling beetle. The type species is Alphitopsis initialis.

Anotylus archaicus[109]

sp nov

Valid

Yue, Makranczy & Ren

Early Cretaceous

Yixian Formation

 China

An oxyteline rove beetle, a species of Anotylus.

Aphebodactyla[110]

Gen. et sp. nov

Valid

Chatzimanolis et al.

Cretaceous

Burmese amber

 Myanmar

A ptilodactylid byrrhoid beetle. The type species is Aphebodactyla rhetine.

Aploceble (Chalcoaploceble) viridiaeneus[111]

Subgen. et sp. nov

Valid

Tshernyshev

Eocene

Baltic amber

 Russia (Kaliningrad Oblast)

A beetle belonging to the family Dasytidae, a subgenus and species of Aploceble.

Apticax[112]

Gen. et 2 sp. nov

Valid

Schomann & Solodovnikov

Early Cretaceous (Aptian–Albian)

Crato Formation

 Brazil

A rove beetle of uncertain phylogenetic placement. The type species is A. volans; genus also includes A. solidus.

Archaeosciaphilus[113]

Gen. et sp. nov

Valid

Legalov

Eocene

Europe

A curculionid beetle found in Baltic amber. The type species is Archaeosciaphilus marshalli.

Attagenus yantarnyi[114]

Sp. nov

Valid

Háva & Bukejs

Late Eocene

Prussian Formation

 Russia (Kaliningrad Oblast)

A dermestid beetle found in Baltic amber, a species of Attagenus.

Baltocar groehni[115]

Sp. nov

Valid

Riedel in Riedel et al.

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A weevil belonging to the family Attelabidae.

Baltocar hoffeinsorum[115]

Sp. nov

Valid

Riedel in Riedel et al.

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A weevil belonging to the family Attelabidae.

Baltocar subnudus[115]

Sp. nov

Valid

Riedel in Riedel et al.

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A weevil belonging to the family Attelabidae.

Cartodere (Aridius) succinobaltica[116]

Sp. nov

Valid

Bukejs, Rücker & Kirejtshuk

Late Eocene

Europe

A latridiid beetle found in Baltic amber, a species of Cartodere.

Catops perkovskyi[117]

Sp. nov

Valid

Perreau

Eocene

 Russia (Kaliningrad Oblast)

A leiodid beetle found in Baltic amber, a species of Catops.

Conapium (Palaeoconapion) alleni[118]

Subgen. et sp. nov

Valid

Legalov

Eocene

Baltic amber

 Russia (Kaliningrad Oblast)

A beetle belonging to the family Brentidae, a subgenus and species of Conapium.

Crassisorus[119]

Gen. et sp. nov

Valid

Nikolajev, Wang & Zhang

Early Cretaceous

Yixian Formation

 China

A member of the family Hybosoridae. The type species is C. fractus.

Cretohypna[120]

Gen. et sp. nov

Valid

Yan, Nikolajev & Ren

Early Cretaceous

Yixian Formation

 China

A glaphyrid beetle. The type species is Cretohypna cristata.

Cretotanaos[121]

Gen. et sp. nov

Valid[122]

Legalov

Early Cretaceous (Aptian)

 Mongolia

An apionine brentid weevil. Type species is Cretotanaos bontsaganensis.

Cryptophagus harenus[123]

Sp. nov

Valid

Lyubarsky & Perkovsky

Late Eocene

 Ukraine

A cryptophagid beetle found in Rovno amber, a species of Cryptophagus.

Electotreta[124]

Gen. et sp.

Valid

Kazantsev

Eocene

 Lithuania

A firefly. The type species is Electotreta rasnitsyni.

Electromethes[124]

Gen. et sp. nov

Valid

Kazantsev

Eocene

 Lithuania

An omethid beetle. The type species is Electromethes alleni.

Electropteron[125]

Gen. et sp. nov

Valid

Kazantsev

Oligocene

 Dominican Republic

A lycid. The type species is E. avus.

Elodes beigeli[126]

Sp. nov

Valid

Klausnitzer

Eocene

Europe

A scirtid beetle found in Baltic amber, a species of Elodes.

Elodes mysticopalpalis[126]

Sp. nov

Valid

Klausnitzer

Eocene

Europe

A scirtid beetle found in Baltic amber, a species of Elodes.

Enicmus adrianae[127]

Sp. nov

Valid

Bukejs, Reike & Rücker

Late Eocene

 Russia (Kaliningrad Oblast)

A member of Latridiidae found in Baltic amber, a species of Enicmus.

Eocenorhynchites[113]

Gen. et sp. nov

Valid

Legalov

Eocene

Europe

A rhynchitid beetle found in Baltic amber. The type species is Eocenorhynchites vossi.

Eodromeus daohugouensis[128]

sp nov

Valid

Wang, Zhang and Ponomarenko

Middle Jurassic

Daohugou Beds

 China

A trachypachid, a species of Eodromeus.

Eodromeus robustus[128]

sp nov

Valid

Wang, Zhang and Ponomarenko

Middle Jurassic

Daohugou Beds

 China

A trachypachid, a species of Eodromeus.

Eoluciola[129]

Gen. et sp. nov

Valid

Kazantsev

Eocene

Europe (Baltic Sea coast)

A lucioline firefly found in Baltic amber. The type species is Eoluciola varang.

Euroleptochromus[130]

Gen. et sp. nov

Valid

Jałoszyński

Eocene

 Lithuania

A rove beetle found in Lithuanian Eocene Baltic amber. The type species is Euroleptochromus sabathi.

Helophorus (Mesosperchus) inceptivus[131]

Sp. nov

Valid

Fikáček et al.

Late Jurassic

 Mongolia

A hydrophiloid beetle, a species of Helophorus.

Helophorus (Mesosperchus) yixianus[131]

Sp. nov

Valid

Fikáček et al.

Jurassic-Cretaceous boundary

Yixian Formation

 China

A hydrophiloid beetle, a species of Helophorus.

Juroglypholoma[132]

Gen. et sp. nov

Valid

Cai et al.

Middle Jurassic

Daohugou Beds

 China

A glypholomatine rove beetle. The type species is Juroglypholoma antiquum.

Kararhynchus gratshevi[133]

Species

Valid

Legalov

Middle-Late Jurassic

Karabastau Formation

 Kazakhstan

An obrieniid beetle, a species of Kararhynchus.

Kararhynchus jurassicus[133]

Species

Valid

Legalov

Middle-Late Jurassic

Karabastau Formation

 Kazakhstan

An obrieniid beetle, a species of Kararhynchus.

Laetopsia[131]

Gen. et comb. nov

Valid

Fikáček et al.

Early Cretaceous

 China
 Mongolia
 Russia

A hydrophiloid beetle; a new genus for "Hydrophilopsia" baissensis Ponomarenko (1987), "H." bontsaganica Prokin (2009), "H." hydraenoides Prokin, Ren & Fikáček (2010), "H." mongolica Ponomarenko (1987) and "H." shatrovskiyi Prokin, Ren & Fikáček (2010).

Lasiosyne cataphracta[134]

Sp. nov

Valid

Yan

Early Cretaceous

 Mongolia

A lasiosynid beetle, a species of Lasiosyne.

Lasiosyne decora[134]

Sp. nov

Valid

Yan

Early Cretaceous

 Russia

A lasiosynid beetle, a species of Lasiosyne.

Lasiosyne insculpta[134]

Sp. nov

Valid

Yan

Late Jurassic

 Mongolia

A lasiosynid beetle, a species of Lasiosyne.

Lasiosyne longitarsa[134]

Sp. nov

Valid

Yan

Late Jurassic

 Mongolia

A lasiosynid beetle, a species of Lasiosyne.

Lasiosyne punctata[134]

Sp. nov

Valid

Yan

Early Cretaceous

 Mongolia

A lasiosynid beetle, a species of Lasiosyne.

Lasiosyne shartegiensis[134]

Sp. nov

Valid

Yan

Late Jurassic

 Mongolia

A lasiosynid beetle, a species of Lasiosyne.

Leehermania[135]

Gen. et sp. nov

Valid

Chatzimanolis, Grimaldi & Engel in Chatzimanolis et al.

Late Triassic (late Carnian or early Norian)

Cow Branch Formation

 United States

A beetle of uncertain phylogenetic placement, might be the earliest known rove beetle or a member of Myxophaga related to the family Hydroscaphidae.[136] The type species is Leehermania prorova.

Lithohypna lepticephala[137]

Sp. nov

Valid

Nikolajev & Ren

Early Cretaceous

Yixian Formation

 China

A beetle belonging to the family Glaphyridae, a species of Lithohypna.

Lithohypna longula[137]

Sp. nov

Valid

Nikolajev & Ren

Early Cretaceous

Yixian Formation

 China

A beetle belonging to the family Glaphyridae, a species of Lithohypna.

Lithohypna tuberculata[137]

Sp. nov

Valid

Nikolajev & Ren

Early Cretaceous

Yixian Formation

 China

A beetle belonging to the family Glaphyridae, a species of Lithohypna.

Lithohypna yuxiana[137]

Sp. nov

Valid

Nikolajev & Ren

Early Cretaceous

Yixian Formation

 China

A beetle belonging to the family Glaphyridae, a species of Lithohypna.

Macratria alleni[138]

Sp. nov

Valid

Telnov

Late Eocene

 Latvia or  Lithuania

A macratriine anthicid beetle found in Baltic amber, a species of Macratria.

Melanapion (Melanapionoides) wanati[118]

Subgen. et sp. nov

Valid

Legalov

Eocene

Baltic amber

 Russia (Kaliningrad Oblast)

A beetle belonging to the family Brentidae, a subgenus and species of Melanapion.

Menopraesagus oryziformis[105]

Sp. nov

Valid[139]

Tan et al.

Middle Jurassic

Jiulongshan Formation

 China

A schizophorid archostematan beetle, a species of Menopraesagus.

Nephus subcircularis[140]

Sp. nov

Valid

Kirejtshuk & Nel

Earliest Eocene

 France

A scymnine coccinellid beetle found in Oise amber, as species of Nephus

Notocupes lini[141]

Sp. nov

Valid

Ponomarenko et al.

Early Jurassic

 China

Originally described as a species of Notocupes; Kirejtshuk (2020) transferred this species to the genus Zygadenia.[142]

Olemehlia[143]

Gen. et sp. nov

Junior homonym

Batelka

Eocene (Lutetian)

Baltic amber

 Lithuania

A member of the family Ripiphoridae belonging to the subfamily Ripidiinae. The type species is O. krali. The generic name is preoccupied by Olemehlia Holzschuh (2011); Batelka (2017) coined a replacement name Olemehliella.[144]

Omma delicata[145]

Sp. nov

Tan et al.

Middle Jurassic

Jiulongshan Formation

 China

A species of Omma

Orchesia (Orchestera) turkini[146]

Sp. nov

Valid

Alekseev & Bukejs

Late Eocene

Prussian Formation

 Russia (Kaliningrad Oblast)

A melandryid beetle found in Baltic amber, a species of Orchesia.

Palaeoerirhinus[121]

Gen. et 4 sp. nov

Valid[122]

Legalov

Early Cretaceous (Aptian)

 Mongolia

A curculionid weevil. Type species is Palaeoerirhinus ponomarenkoi; genus also contains new species P. latus, P. thompsoni and P. longirostris.

Palaeometrioxena[113]

Gen. et sp. nov

Junior synonym

Legalov

Eocene

Baltic amber

Europe (Baltic Sea region)

A belid. The type species is Palaeometrioxena zherikhini. Legalov (2019) considered the genus Palaeometrioxena to be a junior synonym of the genus Archimetrioxena, and transferred the species P zherikhini to the latter genus.[147]

Pareuryomma[145]

Nom. et 2 sp. nov

Tan et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Ommatidae; a replacement name for Euryomma Tan et al. (2006). The type species is "Euryomma" tylodes Tan et al. (2006); genus also includes new species P. cardiobasis. Additional new species described by Tan et al., P. ancistrodonta from the Middle Jurassic Jiulongshan Formation, was subsequently transferred to the genus Omma by Kirejtshuk (2020).[142]

Pleocoma dolichophylla[148]

Sp. nov

Valid

Nikolajev & Ren

Early Cretaceous

Yixian Formation

 China

A rain beetle, a species of Pleocoma.

Prosinodendron[149]

Gen. et sp. nov

Valid

Bai, Ren, & Yang

Barremian to early Aptian

Yixian Formation

 China

A stag beetle. The type species is Prosinodendron krelli.

Prosolierius[150]

Gen. et 3 sp. nov

Valid

Thayer, Newton, & Chatzimanolis

Barremian to Albian

 Lebanon
 Myanmar

A rove beetle.

Pseudobrienia[133]

Gen. et sp. nov

Valid

Legalov

Middle-Late Jurassic

Karabastau Formation

 Kazakhstan

An obrieniid beetle. Its type species is Pseudobrienia rasnitsyni.

Pseudoglaesotropis[113]

Gen. et sp. nov

Valid

Legalov

Eocene

Baltic amber

Europe (Baltic Sea region)

An anthribid beetle. The type species is Pseudoglaesotropis martynovi. Legalov (2019) subsequently changed the rank of the genus Pseudoglaesotropis to the subgenus of the genus Glaesotropis, and transferred the species P. martynovi to the latter genus.[147]

Pulcherhybosorus[151]

Gen. et sp. nov

Valid

Yan, Bai & Ren

Early Cretaceous

Yixian Formation

 China

A hybosorid beetle. The type species is Pulcherhybosorus tridentatus.

Rhyzobius antiquus[140]

Sp. nov

Valid

Kirejtshuk & Nel

Earliest Eocene

Oise amber

 France

A cocciduline coccinellid beetle, a species of Rhyzobius

Rhyzobius gratiosus[140]

Sp. nov

Valid

Kirejtshuk & Nel

Earliest Eocene

Oise amber

 France

A cocciduline coccinellid beetle, a species of Rhyzobius

Septiventer[152]

Gen. et sp. nov

Valid[153]

Bai et al.

Late Jurassic or Early Cretaceous

Yixian Formation

 China

A septiventerid scarabaeoid beetle. The type species is Septiventer quadridentatus.

Shartegopsis[154]

Gen. et sp. nov

Valid

Kirejtshuk

Late Jurassic

 Mongolia

A clerid beetle. The type species is Shartegopsis miranda.

Sinodromeus[128]

Gen. et sp. nov

Valid

Wang, Zhang & Ponomarenko

Early Cretaceous

Yixian Formation

 China

A trachypachid. The type species is Sinodromeus liutiaogouensis.

Sinopeltis[155]

Gen. et 2 sp. nov

Valid

Yu et al.

Middle Jurassic

Jiulongshan Formation

 China

A peltine trogossitid beetle. Genus contains two species: S. jurrasica (type species) and S. amoena. Kirejtshuk (2017) considered the genus Sinopeltis to be a junior synonym of the genus Cervicatinius Tan & Ren (2007).[156]

Sinoschizala[157]

Gen. et sp. nov

Valid

Jarzembowski et al.

Middle Jurassic

Daohugou Beds

 China

A schizophorid beetle. The type species is Sinoschizala darani.

Succinometrioxena[158]

Gen. et sp. nov

Valid

Legalov

Eocene

Europe

A belid weevil found in Baltic amber. The type species is Succinometrioxena poinari.

Sucinoptinus bukejsi[159]

Sp. nov

Valid

Alekseev

Eocene (Priabonian)

Prussian Formation

 Russia (Kaliningrad Oblast)

A spider beetle found in Baltic amber, a species of Sucinoptinus.

Tafforeus[117]

Gen. et sp. nov

Valid

Perreau

Eocene

 Russia (Kaliningrad Oblast)

A leiodid beetle found in Baltic amber. The type species is Tafforeus cainosternus.

Talbragarus[160]

Gen. et sp. nov

Valid

Oberprieler & Oberprieler

Late Jurassic

Talbragar Fish Bed

 Australia

A nemonychid weevil. The type species is Talbragarus averyi.

Tetraphalerus decorosus[145]

Sp. nov

Tan et al.

Middle Jurassic

Jiulongshan Formation

 China

Originally described as a species of Tetraphalerus; Kirejtshuk (2020) transferred this species to the ommatine genus Monticupes.[142]

Toksunius[141]

Gen. et comb. nov

Valid

Ponomarenko et al.

Late Triassic

 China

A new genus for "Ademosynoides" chinaticus Lin (1992).

Tomoderus balticus[161]

Sp. nov

Valid

Telnov

Late Eocene

 Latvia or  Lithuania

A tomoderine anthicid beetle found in Baltic amber, a species of Tomoderus.

Trixagus majusculus[162]

sp nov

Valid

Kovalev, Kirejtshuk & Nel

Lowermost Eocene

 France

A throscid beetle, a species of Trixagus.

Unda chifengensis[128]

sp nov

Valid

Wang, Zhang & Ponomarenko

Middle Jurassic

Daohugou Beds

 China

A trachypachid, a species of Unda.

Xenohimatium[163]

Gen. et sp. nov

Valid

Lyubarsky & Perkovsky

Late Eocene

Rovno amber

 Ukraine

A pleasing fungus beetle belonging to the subfamily Xenoscelinae. The type species is X. rovnense.

Diptera

Name Novelty Status Authors Age Unit Location Notes Images

Aetheapnomyia[164]

Gen. et comb. nov

Valid

Harbach & Greenwalt

Eocene

Europe

A mosquito found in Baltic Amber, a new genus for "Aedes" hoffeinsorum Szadziewski 1998.

Bibio anatolicus[165]

Sp. nov

Valid

Skartveit & Nel

Oligocene

 Turkey

A species of Bibio.

Bibio nigricosta[165]

Sp. nov

Valid

Skartveit & Nel

Oligocene

 Turkey

A species of Bibio.

Calosargus talbragarensis[166]

Sp. nov

Valid

Oberprieler & Yeates

Late Jurassic

 Australia

An archisargid brachyceran, a species of Calosargus.

Clarumreddera[167]

Gen. et 2 sp. nov

Valid

Fedotova & Perkovsky

Late Eocene

Rovno amber

 Ukraine

A cecidomyiid fly, with two species: C. korneyevi and C. conceptiva.

Culiseta kishenehn[164]

Sp. nov

Valid

Harbach & Greenwalt

Eocene

Kishenehn Formation

 United States

A mosquito, a species of Culiseta.

Culiseta lemniscata[164]

Sp. nov

Valid

Harbach & Greenwalt

Eocene

Kishenehn Formation

 United States

A mosquito, a species of Culiseta.

Curtonotum electrodominicum[168]

Sp. nov

Valid

Grimaldi & Kirk-Spriggs

Miocene

 Dominican Republic

A curtonotid found in Dominican amber, a species of Curtonotum.

Daohugosargus[169]

Gen. et comb. nov

Valid

Zhang

Middle Jurassic

Daohugou Beds

 China

An archisargid brachyceran; a new genus for "Sharasargus" eximius Zhang, Yang & Ren (2008).

Depressonotum[168]

Gen. et sp. nov

Valid

Grimaldi & Kirk-Spriggs

Miocene

 Dominican Republic

A curtonotid found in Dominican amber. The type species is Depressonotum priscum.

Dicranomyia (Caenolimonia) alexbrowni[170]

Sp. nov

Valid

Podenas & Poinar

Late Oligocene to early Miocene

 Mexico

A species of Dicranomyia.

Dicranomyia (Dicranomyia) chiapa[170]

Sp. nov

Valid

Podenas & Poinar

Late Oligocene to early Miocene

 Mexico

A species of Dicranomyia.

Dicranomyia (Dicranomyia) mexa[170]

Sp. nov

Valid

Podenas & Poinar

Late Oligocene to early Miocene

 Mexico

A species of Dicranomyia.

Dicranomyia (Dicranomyia) vella[170]

Sp. nov

Valid

Podenas & Poinar

Late Oligocene to early Miocene

 Mexico

A species of Dicranomyia.

Didactylomyia dlusskyi[171]

Sp. nov

Valid

Fedotova & Perkovsky

Late Eocene

 Ukraine

A cecidomyiid fly found in Rovno amber, a species of Didactylomyia.

Enischnomyia[172]

Gen. et sp. nov

Valid

Poinar & Brown

Burdigalian

Dominican amber

 Dominican Republic

A bat fly, type species E. stegosoma,
host for the Plasmodiidae Vetufebrus ovatus[173]

Enischnomyia stegosoma

Eoaedes[164]

Gen. et comb. nov

Valid

Harbach & Greenwalt

Eocene

Europe

A mosquito found in Baltic Amber, a new genus for "Aedes" damzeni Szadziewski 1998.

Eoptychopterina postica[174]

Sp. nov

Valid

Liu, Shih & Ren

Middle Jurassic

Jiulongshan Formation

 China

A ptychopterid, a species of Eoptychopterina.

Eotrichocera (Archaeotrichocera) spatiosa[174]

Sp. nov

Valid

Liu, Shih & Ren

Middle Jurassic

Jiulongshan Formation

 China

A trichocerid, a species of Eotrichocera.

Erioptera (?Mesocyphona) beata[170]

Sp. nov

Valid

Podenas & Poinar

Late Oligocene to early Miocene

 Mexico

A species of Erioptera.

Erioptera (Mesocyphona) diver[170]

Sp. nov

Valid

Podenas & Poinar

Late Oligocene to early Miocene

 Mexico

A species of Erioptera.

Erioptera (?Mesocyphona) sherry[170]

Sp. nov

Valid

Podenas & Poinar

Late Oligocene to early Miocene

 Mexico

A species of Erioptera.

Flagellisargus[175]

Gen. et 3 sp. nov

Valid

Zhang

Callovian or Oxfordian

Daohugou Beds

 China

An archisargid fly. Genus contains three species: Flagellisargus sinicus, F. venustus and F. robustus.

Groveromyia[176]

Gen. et 3 sp. et comb. nov

Valid

Fedotova & Perkovsky

Late Eocene

Rovno amber

 Ukraine

A cecidomyiid fly with 4 species
G. occlusa; G. digna, G. astrosa, and "Bryocrypta" concinna Fedotova (2005).

Helius (Helius) collemus[170]

Sp. nov

Valid

Podenas & Poinar

Late Oligocene to early Miocene

 Mexico

A species of Helius.

Jurochlus adustus[177]

Sp. nov

Valid

Lukashevich & Przhiboro

Mesozoic

 Mongolia

A chironomid, a species of Jurochlus.

Jurochlus limbatus[177]

Sp. nov

Valid

Lukashevich & Przhiboro

Mesozoic

 Mongolia

A chironomid, a species of Jurochlus.

Jurochlus lineatus[177]

Sp. nov

Valid

Lukashevich & Przhiboro

Mesozoic

 Mongolia

A chironomid, a species of Jurochlus.

Jurochlus trivittatus[177]

Sp. nov

Valid

Lukashevich & Przhiboro

Mesozoic

 Mongolia

A chironomid, a species of Jurochlus.

Lacrimyza[178]

Gen. et 2 sp. nov

Valid[179]

Roháček

Eocene

Europe

An anthomyzid. Genus contains two species: L. lacrimosa and L. christelae.

Laiyangitabanus[180]

Gen. et sp. nov

Valid

Zhang

Early Cretaceous

Laiyang Formation

 China

A horse-fly. The type species is Laiyangitabanus formosus.

Lithorhagio[181]

Gen. et sp. nov

Valid

Zhang & Li

Middle Jurassic or lowermost Upper Jurassic

Daohugou Beds

 China

A rhagionid. The type species is Lithorhagio megalocephalus.

Lygistorrhina caribbiana[182]

Sp. nov

Valid

Grund

Miocene

 Dominican Republic

A lygistorrhinid fungus gnat found in Dominican amber, a species of Lygistorrhina.

Mesorhyphus handlirschi[183]

Sp. nov

Valid

Lukashevich

Upper Jurassic

 Mongolia

An anisopodid bibionomorph, a species of Mesorhyphus.

Mesorhyphus hennigi[183]

Sp. nov

Valid

Lukashevich

Upper Jurassic

 Mongolia

An anisopodid bibionomorph, a species of Mesorhyphus.

Mesosciophila sigmoidea[184]

sp nov

Valid

Wang, Zhao & Ren

Middle Jurassic

Jiulongshan Formation

 China

A mesosciophilid nematoceran, a species of Mesosciophila.

Mesosolva zhangae[169]

Nom. nov

Valid

Zhang

Middle Jurassic

Daohugou Beds

 China

An archisargid brachyceran; a replacement name for Brevisolva daohugouensis Zhang et al. (2010).

Nemopalpus inexpectatus[185]

Sp. nov

Valid

Wagner in Wagner & Stuckenberg

Late Eocene

Baltic amber

Europe

A bruchomyiine psychodid fly. Originally described as a species of Nemopalpus, but subsequently transferred to the genus Palaeosycorax.[186]

Nemopalpus velteni[185]

Sp. nov

Valid

Wagner in Wagner & Stuckenberg

Cretaceous (Albian)

Burmese amber

 Myanmar

A bruchomyiine psychodid fly. Originally described as a species of Nemopalpus, but subsequently transferred to the genus Palaeoglaesum.[186]

Orientisargus[169]

Gen. et sp. nov

Valid

Zhang

Latest Middle Jurassic or earliest Late Jurassic

Daohugou Beds

 China

An orientisargid archisargoid brachyceran. The type species is Orientisargus illecebrosus.

Oryctochlus brundini[187]

Sp. nov

Valid

Lukashevich

Late Jurassic

 Mongolia

A podonomine chironomid, as species of Oryctochlus

Oryctochlus kaluginae[187]

Sp. nov

Valid

Lukashevich

Late Jurassic

 Mongolia

A podonomine chironomid, as species of Oryctochlus

Palaeopoecilostola eocenica[188]

Sp. nov

Valid

Kania & Krzemiński

Eocene

Europe

A limoniid found in Baltic amber, a species of Palaeopoecilostola.

Paramesosciophilodes aequus[184]

sp nov

Valid

Wang, Zhao & Ren

Middle Jurassic

Jiulongshan Formation

 China

A mesosciophilid nematoceran, a species of Paramesosciophilodes.

Penthetria beskonakensis[165]

Sp. nov

Valid

Skartveit & Nel

Oligocene

 Turkey

A species of Penthetria.

Probittacomorpha brisaci[189]

sp nov

Valid

Krzemiński, Kania & Nel

Late Miocene

 France

A ptychopterid, a species of Probittacomorpha.

Procramptonomyia kovalevi[183]

Sp. nov

Valid

Lukashevich

Upper Jurassic

 Kazakhstan
 Mongolia

A procramptonomyiid bibionomorph, a species of Procramptonomyia.

Procramptonomyia ponomarenkoi[183]

Sp. nov

Valid

Lukashevich

Upper Jurassic

 Mongolia

A procramptonomyiid bibionomorph, a species of Procramptonomyia.

Protanthomyza hennigi[178]

Sp. nov

Valid[179]

Roháček

Eocene

Europe

An anthomyzid, a species of Protanthomyza.

Protanthomyza hoffeinsorum[178]

Sp. nov

Valid[179]

Roháček

Eocene

Europe

An anthomyzid, a species of Protanthomyza.

Protanthomyza krylovi[178]

Sp. nov

Valid[179]

Roháček

Eocene

Europe

An anthomyzid, a species of Protanthomyza.

Protanthomyza loewi[178]

Sp. nov

Valid[179]

Roháček

Eocene

Europe

An anthomyzid, a species of Protanthomyza.

Protanthomyza meunieri[178]

Sp. nov

Valid[179]

Roháček

Eocene

Europe

An anthomyzid, a species of Protanthomyza.

Protanthomyza presli[178]

Sp. nov

Valid[179]

Roháček

Eocene

Europe

An anthomyzid, a species of Protanthomyza.

Protanthomyza tschirnhausi[178]

Sp. nov

Valid[179]

Roháček

Eocene

Europe

An anthomyzid, a species of Protanthomyza.

Protoculicoides ciliatus[190]

Sp. nov

Valid

Borkent

Cretaceous

Canadian amber

 Canada

A member of the family Ceratopogonidae. Originally described as a species of Protoculicoides; Borkent (2019) transferred this species to the genus Atriculicoides.[191]

Protorhagio parvus[181]

sp nov

Valid

Zhang & Li

Middle Jurassic or lowermost Upper Jurassic

Daohugou Beds

 China

A rhagionid, a species of Protorhagio.

Protorhyphus rohdendorfi[183]

Sp. nov

Valid

Lukashevich

Upper Jurassic

 Mongolia

A protorhyphid bibionomorph, a species of Protorhyphus.

Rovnodidactylomyia[171]

Gen. et 3 sp. et comb. nov

Valid

Fedotova & Perkovsky

Late Eocene

Europe

A cecidomyiid fly with 5 species
R. zosimovichi, R. sidorenkoi, R. iconica) in Rovno amber,
plus "Bryocrypta" girafa Meunier (1904) and "B." capitosa Meunier (1904) in Baltic amber.

Sinallomyia[169]

Nom. nov

Valid

Zhang

Early Cretaceous

Yixian Formation

 China

An archisargid brachyceran; a replacement name for Allomyia Ren (1998).

Sinocretomyia[180]

Gen. et sp. nov

Valid

Zhang

Early Cretaceous

Laiyang Formation

 China

An athericid. The type species is Sinocretomyia minuscula.

Stilobezzia pikei[190]

Sp. nov

Valid

Borkent

Cretaceous

Canadian amber

 Canada

A species of Stilobezzia.

Stomatosema iljieteugeniae[167]

Sp. nov

Valid

Fedotova & Perkovsky

Late Eocene

Rovno amber

 Ukraine

A cecidomyiid fly

Tipula paleopannonia[192]

Sp. nov

Valid

Engel & Gross

Miocene (ca. 11.3 Ma)

Paldau Formation

 Austria

A crane fly, a species of Tipula.

Ephemeroptera

Name Novelty Status Authors Age Unit Location Notes Images

Alexandrinia[193]

Gen. et sp.

Valid

Sinitshenkova & Vassilenko

Permian

Poldarsa Formation

 Russia
 United States

A mayfly. Species are A. gigantea and A. directum.

Hammephemera[194]

Gen. et sp. nov

Valid

Sinitshenkova in Bashkuev et al.

Early Triassic

 Germany

A sharephemerid mayfly. The type species is Hammephemera pulchra.

Misthodotes tshernovae[193]

sp nov

Valid

Sinitshenkova & Vassilenko

Upper Permian

Poldarsa Formation

 Russia

A mayfly, a species of Misthodotes.

Ponalex[195]

Gen. et sp. nov

Valid

Sinitshenkova & Aristov

Late Permian

Poldarsa Formation

 Russia

A protereismatid mayfly. The type species is Ponalex maximus.

Siphloplecton barabani[196]

Sp. nov

Valid

Staniczek & Godunko

Eocene

Europe

A metretopodid mayfly found in Baltic amber, a species of Siphloplecton.

Siphloplecton demoulini[196]

Sp. nov

Valid

Staniczek & Godunko

Eocene

Europe

A metretopodid mayfly found in Baltic amber, a species of Siphloplecton.

Siphloplecton hageni[196]

Sp. nov

Valid

Staniczek & Godunko

Eocene

Europe

A metretopodid mayfly found in Baltic amber, a species of Siphloplecton.

Siphloplecton picteti[196]

Sp. nov

Valid

Staniczek & Godunko

Eocene

Europe

A metretopodid mayfly found in Baltic amber, a species of Siphloplecton.

Hemiptera

Name Novelty Status Authors Age Unit Location Notes Images

Acaricoris robertae[197]

Sp. nov

Valid

Heiss & Poinar

Miocene

 Dominican Republic

An aradid found in Dominican amber, a species of Acaricoris.

Arctorthezia baltica[198]

Sp. nov

Valid

Vea & Grimaldi

Eocene

Europe

An ortheziid found in Eocene Baltic amber, a species of Arctorthezia.

Breviscutum[199]

Gen. et sp. nov

Valid[200]

Yao et al.

Cretaceous

 China

A primipentatomid pentatomoid. The type species is Breviscutum lunatum.

Burmacicada[201]

Gen. et sp. nov

Valid

Poinar & Kritsky

Cretaceous

Burmese amber

 Myanmar

A cicada. The type species is Burmacicada protera.

Burmorthezia[198]

Gen et 2 sp. nov

Valid

Vea & Grimaldi

Cretaceous (98 Ma)

Burmese amber

 Myanmar

An ortheziid. Genus contains two species: Burmorthezia kotejai and B. insolita.

Carventus bechlyi[197]

Sp. nov

Valid

Heiss & Poinar

Miocene

 Dominican Republic

An aradid found in Dominican amber, a species of Carventus.

Cicadocoris varians[202]

Sp. nov

Disputed

Dong, Yao & Ren

Middle Jurassic

Jiulongshan Formation

 China

A moss bug belonging to the family Progonocimicidae. Jiang & Huang (2017) considered this species to be a probable junior synonym of Cicadocoris sinensis Hong (1983).[203]

Cinemala[204]

Gen. et sp. nov

Valid

Shcherbakov

Middle or Late Jurassic

Bakhar Group

 Mongolia

A member of Cicadomorpha belonging to the family Hylicellidae. The type species is C. contrasta.

Cintux[205]

Gen. et sp. nov

Valid

Stroiński & Szwedo

Palaeocene

 France

A lophopid planthopper. The type species is Cintux menatensis.

Conjucella[204]

Gen. et sp. nov

Valid

Shcherbakov

Early Cretaceous

Zaza Formation

 Russia

A member of Cicadomorpha belonging to the family Hylicellidae. The type species is C. selene.

Dellashara[206]

Gen. et sp. nov

Valid

Shcherbakov

Late Jurassic

Shar Teg Sequence

 Mongolia

A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is D. tega.

Dominicicada[201]

Gen. et sp. nov

Valid

Poinar & Kritsky

 Dominican Republic

A cicada found in Dominican amber. The type species is Dominicicada youngi.

Edessa protera[207]

Sp. nov

Valid

Poinar & Thomas

Tertiary

 Mexico

A pentatomid stink bug found in Mexican amber, a species of Edessa.

Hispanocader[208]

Gen. et sp. nov

Valid

Golub, Popov & Arillo

Albian

Escucha Formation

 Spain

A relative of tingids. The type species is Hispanocader lisae.

Hoffeinsoria[209]

Gen. et sp. nov

Valid

Herczek & Popov

Eocene

Prussian Formation

Europe

An isometopine mirid found in Eocene Baltic amber. The type species is Hoffeinsoria robusta.

Jiania[210]

Gen. et 2 sp. nov

Valid

Wang & Szwedo in Wang, Szwedo & Zhang

Middle Jurassic

Daohugou Beds

 China

A sinoalid froghopper. Genus contains two species: Jiania crebra and Jiania gracila. Fu, Cai & Huang (2018) considered J. gracila to be a junior synonym of J crebra.[211]

Kachinocoris[212]

Gen. et sp. nov

Valid

Heiss

Cretaceous

Burmese amber

 Myanmar

A flat bug found in Cretaceous Burmese Amber. The type species is Kachinocoris brevipennis.

Kemobius[206]

Gen. et sp. nov

Valid

Shcherbakov

Middle Jurassic

Itat Formation

 Russia

A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is K. lux.

Kisa[206]

Gen. et sp. nov

Valid

Shcherbakov

Middle Jurassic

Itat Formation

 Russia

A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is K. fasciata.

Kubecola[206]

Gen. et sp. nov

Valid

Shcherbakov

Middle Jurassic

Itat Formation

 Russia

A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is K. guttatus.

Microaradus[213]

Gen. et sp. nov

Valid

Heiss & Poinar

Mesozoic

Burmese amber

 Myanmar

An archaearadine aradid found in Mesozoic Burmese amber. The type species is Microaradus anticus.

Minyscapheus[214]

Gen. et sp. nov.

Valid

Poinar, Kritsky & Brown

 Dominican Republic

A cicada found in Dominican amber. The type species is Minyscapheus dominicanus.

Mixorthezia dominicana[198]

Sp. nov

Valid

Vea & Grimaldi

Miocene

 Dominican Republic

An ortheziid found in Miocene Dominican amber, a species of Mixorthezia.

Mixorthezia kozari[198]

Sp. nov

Valid

Vea & Grimaldi

Miocene

 Dominican Republic

An ortheziid found in Miocene Dominican amber, a species of Mixorthezia.

Mongolocoris[215]

Gen. et comb. nov

Valid

Ryzhkova

Early Cretaceous

 Mongolia

An enicocorid leptopodomorph heteropteran, a new genus for "Enicocoris" tibialis Popov, 1986.

Myanmezira[213]

Gen. et sp. nov

Valid

Heiss & Poinar

Mesozoic

Burmese amber

 Myanmar

A mezirine aradid found in Mesozoic Burmese amber. The type species is Myanmezira longicornis.

Nesoproxius latocanus[197]

Sp. nov

Valid

Heiss & Poinar

Miocene

 Dominican Republic

An aradid found in Dominican amber, a species of Nesoproxius.

Notonecta vetula[216]

sp nov

Valid

Zhang, Yao & Ren

Late Jurassic

Chijinqiao Formation

 China

A notonectid,a species of Notonecta.

Oropentatoma[199]

Gen. et sp. nov

Valid[217]

Yao et al.

Cretaceous

 China

A primipentatomid pentatomoid. The type species is Oropentatoma epichara.

Poljanka hirsuta[218]

sp nov

Valid

Yang, Yao & Ren

Middle Jurassic

Jiulongshan Formation

 China

A member of Sternorrhyncha, a protopsyllidiid; a species of Poljanka.

Primipentatoma[199]

Gen. et 2 sp. nov

Valid[219]

Yao et al.

Cretaceous

 China

A primipentatomid pentatomoid. Genus contains two species: Primipentatoma peregrina and Primipentatoma fangi.

Proneoproxius[197]

Gen. et sp. nov

Valid

Heiss & Poinar

Miocene

 Dominican Republic

An aradid found in Dominican amber. The type species is Proneoproxius cornutus.

Purbecellus[204]

Gen. et comb. nov

Valid

Shcherbakov

Early Cretaceous

Purbeck Group

 United Kingdom

A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is "Cicadellium" psocus Westwood (1854).

Quadrocoris[199]

Gen. et sp. nov

Valid[220]

Yao et al.

Cretaceous

 China

A primipentatomid pentatomoid. The type species is Quadrocoris radius.

Sinaldocader rasnitsyni[221]

Sp. nov

Valid

Golub & Popov

Turonian

 Kazakhstan

A lace bug. Originally described as a species of Sinaldocader; Popov & Golub (2019) transferred this species to the separate genus Kzylcader.[222]

Sinoala[210]

Gen. et sp. nov

Valid

Wang & Szwedo in Wang, Szwedo & Zhang

Middle Jurassic

Daohugou Beds

 China

A sinoalid froghopper. The type species is Sinoala parallelivena.

Sunotettigarcta hirsuta[223]

sp nov

Valid

Li, Wang & Ren in Li et al.

Middle Jurassic

Daohugou Beds

 China

A tettigarctid, a species of Sunotettigarcta.

Ulanocoris[215]

Gen. et 2 sp. nov

Valid

Ryzhkova

Early Cretaceous

 Mongolia

An enicocorid leptopodomorph heteropteran. Genus contains two species: Ulanocoris femoralis and U. grandis.

Venustsalda[224]

Gen. et sp. nov

Valid

Zhang et al.

Lower Cretaceous

Yixian Formation

 China

A saldid heteropteran. The type species is Venustsalda locella.

Hymenoptera

Name Novelty Status Authors Age Unit Location Notes Images

Agapia[225]

Gen. et sp. nov

Valid

Kopylov

Late Cretaceous

 Russia

An ichneumonid. The type species is Agapia sukatchevae.

Agapteron[225]

Gen. et sp. nov

Valid

Kopylov

Late Cretaceous

 Russia

An ichneumonid. The type species is Agapteron popovi.

Archimyrmex wedmannae[226]

Sp. nov

Valid

Dlussky

Middle Eocene

Messel Pit

 Germany

A myrmeciine ant, a species of Archimyrmex.

Archimyrmex wedmannae

Anthophorula (Anthophorula) persephone[227]

Sp. nov

Valid

Engel in Engel et al.

Early Miocene

 Dominican Republic

An exomalopsine apid bee found in Dominican amber, a species of Anthophorula.

Atefia[228]

Gen. et sp. nov

Valid[229]

Krogmann et al.

Early Cretaceous

Crato Formation

 Brazil

A tenthredinoid hymenopteran sharing morphological characters with the families Pergidae and Diprionidae. The type species is Atefia rasnitsyni.

Boltonidris[230]

Gen. et sp. nov

Valid

Radchenko & Dlussky

Late Eocene

Rovno amber

 Ukraine

A stenammine myrmicine ant found in Rovno amber. The type species is Boltonidris mirabilis.

Bombus (Bombus) randeckensis[231]

Sp. nov

Valid

Wappler & Engel in Wappler et al.

Miocene

 Germany

A bumblebee.

Casaleia eocenica[232][233]

Sp nov

valid

Dlussky & Wedmann

Lutetian

Messel Formation

 Germany

An Amblyoponin ant.

Casaleia eocenica

Cascoscelio[5]

Gen. et sp. nov.

Valid

Poinar in Poinar & Buckley

Early Cretaceous

Burmese amber

 Myanmar

A platygastrid wasp,
type species is C. incassus.

Cascoscelio incassa

Cephalopone[232]

Gen. et 2 sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A poneromorph ant. Two species: C. potens & C. grandis.

Cephalopone grandis

Cratoenigma[234]

Gen. et sp. nov

Valid

Krogmann & Nél

Early Cretaceous (Aptian)

Crato Formation

 Brazil

A probable relative of xiphydriids and euhymenopterans. The type species is C. articulata.

Cretevania soplaensis[235]

Sp. nov

Valid

De La Fuente, Peñalver & Ortega-Blanco

Albian

 Spain

An evaniid, a species of Cretevania.

Cretorhyssalus[236]

Gen. et sp. nov

Valid

Belokobylskij

Cenomanian

Ola Formation

 Russia

A braconid wasp. Type species is Cretorhyssalus brevis.

Cyrtopone[232]

Gen. et 4 sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel Formation

 Germany

A ponerinae ant. Genus contains four species: Cyrtopone curiosa, C. elongata, C. microcephala and C. striata.

C. striata

Gulgonga[237]

Gen. et sp. nov

Valid

Oberprieler, Rasnitsyn & Brothers

Late Jurassic

Talbragar Fish Bed

 Australia

A praeaulacid evanioid. The type species is Gulgonga beattiei.

Haidomyrmex scimitarus[238]

Sp. nov

Valid

Barden & Grimaldi

Late Aptian to Early Cenomanian

Burmese amber

 Myanmar

A Sphecomyrmin ant

Haidomyrmex scimitarus

Haidomyrmex zigrasi[238]

Sp. nov

Valid

Barden & Grimaldi

Campanian

Burmese amber

 Myanmar

A Sphecomyrmin ant

Haidomyrmex zigrasi

Hyptia deansi[239]

Sp. nov

Valid

Jennings, Krogmann & Mew

Late Oligocene or early Miocene

 Mexico

An evaniid hymenopteran, a species of Hyptia.

Ibalia (Ibalia) electra [240]

Sp. nov

Valid

Engel & Liu

Eocene

Europe

An ibaliid found in Baltic amber, a species of Ibalia.

Lancepyris[241]

Gen. et sp. nov

Valid

Azevedo & Azar

Early Cretaceous

 Lebanon

A lancepyrine bethylid found in Early Cretaceous Lebanese amber. The type species is Lancepyris opertus.

Lonchodryinus balticus[242]

Sp. nov

Valid

Olmi & Guglielmino

Late Eocene

Europe

A dryinid found in Baltic amber, a species of Lonchodryinus.

Magadanobracon[236]

Gen. et 2 sp. nov

Valid

Belokobylskij

Cenomanian

Ola Formation

 Russia

A braconid wasp. The type species is M. rasnitsyni; a second species is M. zherikhini.

Messelepone[232]

Gen. et sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A poneromorph ant. The type species is Messelepone leptogenoides.

Messelepone leptogenoides

Mirabythus[243]

Gen. et 2 sp. nov

Valid

Cai, Shih & Ren in Cai et al.

Early Cretaceous

Yixian Formation

 China

A scolebythid chrysidoid. The genus contains two species: Mirabythus lechrius and M. liae.

Pachycondyla eocenica[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A Ponerinae ant.

Pachycondyla lutzi[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A ponerine ant

Pachycondyla? messeliana[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A ponerine ant.

Pachycondyla minuta[232][244]

Sp. nov

synonym

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A ponerine ant,
name occupied, renamed P. parvula.

Pachycondyla petiolosa[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A ponerine ant.

Pachycondyla petrosa[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A poneromorph ant, a species of Pachycondyla.

Platyxyela[245]

Gen. et sp. nov

Valid

Wang, Shih & Ren

Middle Jurassic

Jiulongshan Formation

 China

A macroxyeline xyelid sawfly. The type species is Platyxyela unica.

Praeichneumon dzhidensis[246]

sp nov

Valid

Kopylov

Lower Cretaceous

 Russia

A praeichneumonid ichneumonoid, a species of Praeichneumon.

Praeichneumon khamardabanicus[246]

sp nov

Valid

Kopylov

Lower Cretaceous

 Russia

A praeichneumonid ichneumonoid, a species of Praeichneumon.

Praeichneumon zakhaaminicus[246]

sp nov

Valid

Kopylov

Lower Cretaceous

 Russia

A praeichneumonid ichneumonoid, a species of Praeichneumon.

Prionomyrmex wappleri[226]

Sp. nov

Valid

Dlussky

Late Oligocene

 Germany

A myrmeciine ant.

Prionomyrmex wappleri

Protodryinus[242]

Gen. et sp. nov

Valid

Olmi & Guglielmino

Late Eocene

Europe

A dryinid found in Baltic amber. The type species is Protodryinus eocenicus.

Protopone? dubia[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel Formation

 Germany

A ponerine ant

Protopone? dubia

Protopone germanica[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A ponerine ant

Protopone germanica

Protopone magna[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A ponerine ant

Protopone magna

Protopone oculata[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A ponerine ant

Protopone oculata

Protopone sepulta[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A ponerine ant

Protopone sepulta

Protopone vetula[232]

Sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

A ponerine ant

Protopone vetula

Pseudectatomma[232]

Gen. et 2 sp. nov

Valid

Dlussky & Wedmann

Eocene

Messel pit

 Germany

An Ectatommin ant. Two species: P. eocenica and P. striatula.

Pseudectatomma eocenica

Trichelyon[247]

Gen. et sp. nov

Valid

Ortega-Blanco & Engel in Ortega-Blanco, Singh & Engel

Early Eocene (Ypresian)

 India

A rhysipoline exothecine braconid wasp found in Cambay amber. The type species is Trichelyon tadkeshwarense.

Urotryphon baikurensis[225]

Sp. nov

Valid

Kopylov

Late Cretaceous

 Russia

An ichneumonid, a species of Urotryphon.

Zatania electra[248]

Sp. nov

Valid

Lapolla, Kallal & Brady

Eocene or Miocene

Dominican amber

 Dominican Republic

A formicine ant, a species of Zatania.

Mecoptera

Name Novelty Status Authors Age Unit Location Notes Images

Baltipanorpa[249]

Gen. et sp. nov

Valid

Krzemiński & Soszyńska-Maj

Middle Eocene

 Poland

A mecopteran found in Baltic amber. The type species is Baltipanorpa damzeni.

Exilibittacus[250]

Gen. et sp. nov

Valid

Yang, Ren & Shih

Middle Jurassic

Jiulongshan Formation

 China

A hangingfly. The type species is Exilibittacus lii.

Jeholopsyche bella[251]

Sp. nov

Valid

Qiao, Shih & Ren

Early Cretaceous

Yixian Formation

 China

An aneuretopsychid mecopteran, a species of Jeholopsyche.

Jeholopsyche completa[251]

Sp. nov

Valid

Qiao, Shih & Ren

Early Cretaceous

Yixian Formation

 China

An aneuretopsychid mecopteran, a species of Jeholopsyche.

Jeholopsyche maxima[251]

Sp. nov

Valid

Qiao, Shih & Ren

Early Cretaceous

Yixian Formation

 China

An aneuretopsychid mecopteran, a species of Jeholopsyche.

Juracimbrophlebia[252]

Gen. et sp. nov

Valid

Wang, Labandeira, Shih & Ren in Wang et al.

Middle Jurassic (Bathonian or Callovian)

Jiulongshan Formation

 China

A cimbrophlebiid mecopteran, the wings and abdomen of which mimicked ginkgo leaves. The type species is Juracimbrophlebia ginkgofolia.

Megabittacus spatiosus[250]

Sp. nov

Valid

Yang, Ren & Shih

Late Jurassic or Early Cretaceous

Yixian Formation

 China

A hangingfly, a species of Megabittacus.

Orthophlebia nervulosa[253]

sp nov

Valid

Qiao, Shih & Ren

Middle Jurassic

Jiulongshan Formation

 China

A mecopteran, a species of Orthophlebia.

Orthophlebia riccardii[254]

Sp. nov

Valid

Petrulevičius & Ren

Middle Jurassic

Jiulongshan Formation

 China

A mecopteran. Originally described as a species of Orthophlebia, subsequently transferred to the separate genus Gigaphlebia.[255]

Orthophlebia stigmosa[253]

sp nov

Valid

Qiao, Shih & Ren

Middle Jurassic

Jiulongshan Formation

 China

A mecopteran. Originally described as a species of Orthophlebia, subsequently transferred to the separate genus Longiphlebia.[255]

Perfecticimbrophlebia[256]

Gen. et sp. nov

Valid

Yang, Shih & Ren in Yang et al.

Middle Jurassic

Jiulongshan Formation

 China

A cimbrophlebiid mecopteran. The type species is Perfecticimbrophlebia laetus.

Preanabittacus validus[256]

sp nov

Valid

Yang, Shih & Ren in Yang et al.

Middle Jurassic

Jiulongshan Formation

 China

A hangingfly, a species of Preanabittacus.

Neuroptera

Name Novelty Status Authors Age Unit Location Notes Images

Alloepipsychopsis[257]

Gen. et sp. nov

Valid

Makarkin, Yang, et al.

Barremian

Yixian Formation

 China

A psychopsid. Type species is Alloepipsychopsis lata.

Chorilingia[258]

Gen. et 4 sp. nov

Valid

Shi et al.

Middle Jurassic

Jiulongshan Formation

 China

A grammolingiid neuropteran. Genus contains four species: of C. euryptera, C. parvica, C. translucida and C. peregrina.

Guyiling[259]

Gen. et sp. nov

Valid

Shi et al.

Early Cretaceous

Yixian Formation

 China

A member of Myrmeleontiformia of uncertain phylogenetic placement. The type species is G. jianboni.

Hallucinochrysa[260]

Gen. et sp. nov

Valid

Pérez-de la Fuente, Delclòs, Peñalver & Engel in Pérez-de la Fuente et al.

Early Cretaceous (Albian)

 Spain

A chrysopoid neuropteran (a relative of the green lacewings). The type species is Hallucinochrysa diogenesi.

Kareninoides[261]

Gen. et sp. et comb. nov

Valid

Yang, Makarkin & Ren

Early Cretaceous

La Huerguina Formation
Yixian Formation

 China
 Spain

A mesochrysopid neuropteran. The type species is Kareninoides lii; genus might also contain "Armandochrysopa" inexpecta Nel et al. (2005).

Leptolingia oblonga[262]

Sp. nov

Valid

Khramov

Late Jurassic

 Mongolia

A grammolingiid neuropteran, a species of Leptolingia.

Litholingia longa[262]

Sp. nov

Valid

Khramov

Early or Middle Jurassic

 Kyrgyzstan

A grammolingiid neuropteran, a species of Litholingia.

Parakseneura[263]

Gen. et 11 sp. nov

Valid

Yang, Makarkin & Ren in Yang et al.

Middle Jurassic

Jiulongshan Formation

 China

A parakseneurid neuropteran. Genus contains 11 species.

Protoaristenymphes daohugouensis[261]

Sp. nov

Valid

Yang, Makarkin & Ren

Middle Jurassic

Daohugou Beds

 China

A mesochrysopid neuropteran, a species of Protoaristenymphes.

Protolingia[262]

Gen. et sp. nov

Valid

Khramov

Early or Middle Jurassic

 Kyrgyzstan

A grammolingiid neuropteran. The type species is Protolingia mira.

Pseudorapisma[263]

Gen. et 3 sp. nov

Valid

Yang, Makarkin & Ren in Yang et al.

Middle Jurassic

Jiulongshan Formation

 China

A parakseneurid neuropteran. Genus contains 3 species: P. jurassicum, P. maculatum and P. angustipenne.

Shuraboneura[263]

Gen. et sp. nov

Valid

Khramov & Makarkin in Yang et al.

Early or Middle Jurassic

Sogul Formation

 Kyrgyzstan

A parakseneurid neuropteran. The type species is Shuraboneura ovata.

Notoptera

Name Novelty Status Authors Age Unit Location Notes Images

Duoduo[264]

Gen. et sp. nov

Valid

Cui

Middle Jurassic

Daohugou Beds

 China

A putative relative of grylloblattids. The type species is Duoduo qianae.

Kitshuga[265]

Gen. et sp. nov

Valid

Aristov

Upper Permian

 Russia

A cacurgid grylloblattid. The type species is Kitshuga ryzhkovae.

Oborella brauckmanni[266]

sp nov

Valid

Prokop et al.

Early Permian

Meisenheim Formation

 Germany

A euryptilonid grylloblattid, a species of Oborella.

Oborella germanica[266]

sp nov

Valid

Prokop et al.

Early Permian

Meisenheim Formation

 Germany

A euryptilonid grylloblattid, a species of Oborella.

Odonatoptera

Name Novelty Status Authors Age Unit Location Notes Images

Austroprotolindenia[267]

Gen. et sp. nov

Valid

Beattie & Nel

Late Jurassic

Talbragar Fossil Fish Bed

 Australia

A dragonfly. The type species is Austroprotolindenia jurassica.

Balticoagrion[268]

Gen. et sp. nov

Valid

Bechly

Eocene

Europe

A damselfly found in Baltic amber. The type species is Balticoagrion paulyi.

Decoraeshna[269]

Gen. et sp. nov

Valid

Li et al.

Early Cretaceous

Yixian Formation

 China

A progobiaeshnid dragonfly. The type species is Decoraeshna preciosus.

Germanostenolestes[270]

Gen. et sp. nov

Valid

Nel & Fleck

Middle Miocene

 Germany

A sieblosiid epiproctophoran odonate. The type species is Germanostenolestes lutzi.

Karatawia sinica[271]

Sp. nov

Valid

Li, Nel & Ren in Li et al.

Middle Jurassic

Jiulongshan Formation

 China

A campterophlebiid odonate,a species of Karatawia.

Mongoliaeshna exiguusens[269]

Sp. nov

Valid

Li et al.

Early Cretaceous

Yixian Formation

 China

A progobiaeshnid dragonfly, a species of Mongoliaeshna.

Mongoliaeshna hadrens[269]

Sp. nov

Valid

Li et al.

Early Cretaceous

Yixian Formation

 China

A progobiaeshnid dragonfly, a species of Mongoliaeshna.

Oligomazon[272]

Gen. et comb. nov

Valid

Zessin & Brauckmann

Carboniferous (Pennsylvanian)

 United States

A member of Meganisoptera, a paralogid; a new genus for "Oligotypus" makowskii Carpenter & Richardson (1971).

Paragilsonia[273]

Gen. et sp. nov (but see notes)

Valid

Zhang, Hong & Su in Su et al.

Carboniferous (Namurian)

 China

A member of Odonatoptera; the type species is Paragilsonia orientalis. Li et al. (2013) synonymized Paragilsonia with the genus Tupus Sellards (1906); however, they retained P. orientalis as a distinct species within the latter genus.[274]

Paralogobora[272]

Gen. et sp. nov

Valid

Zessin & Brauckmann

Early Permian

 Czech Republic

A member of Odonata. The type species is Paralogobora guentherpetersi.

Pseudocymatophlebia boda[275]

Sp. nov

Valid

Li, Nel & Ren in Li et al.

Early Cretaceous

 China

An aktassiid odonate, a species of Pseudocymatophlebia.

Sinierasiptera[273]

Gen. et sp. nov (but see notes)

Valid

Zhang, Hong & Su in Su et al.

Carboniferous (Namurian)

 China

A member of Odonatoptera; the type species is Sinierasiptera jini. Li et al. (2013) synonymized Sinierasiptera with the genus Erasipterella Brauckmann (1983); however, they retained S. jini as a distinct species within the latter genus.[274]

Sinojagoria cancellosa[269]

Sp. nov

Valid

Li et al.

Early Cretaceous

Yixian Formation

 China

A gomphaeschnid dragonfly, a species of Sinojagoria.

Sinojagoria magna[269]

Sp. nov

Valid

Li et al.

Early Cretaceous

Yixian Formation

 China

A gomphaeschnid dragonfly, a species of Sinojagoria.

Orthopterida

Name Novelty Status Authors Age Unit Location Notes Images

Archaboilus musicus[276]

Sp. nov

Valid

Gu, Engel and Ren in Gu et al.

Bathonian to Callovian

Jiulongshan Formation

 China

A haglid orthopteran.

Eoerianthus[277]

Gen. et sp. nov

Valid

Gorochov in Gorochov & Labandeira

Eocene

Green River Formation

 United States

A member of the family Eumastacidae. The type species is E. eocaenicus.

Eogryllus[277]

Gen. et 2 sp. nov

Valid

Gorochov in Gorochov & Labandeira

Eocene

Green River Formation

 United States

A member of the tribe Gryllini. The type species is E. unicolor; genus might also include Eogryllus? elongatus.

Eomogoplistes[277]

Gen. et sp. nov

Valid

Gorochov in Gorochov & Labandeira

Eocene

Green River Formation

 United States

A member of the family Mogoplistidae. The type species is E. longipennis.

Eotetrix[277]

Gen. et sp. nov

Valid

Gorochov in Gorochov & Labandeira

Eocene

Green River Formation

 United States

A member of the family Tetrigidae. The type species is E. unicornis''.

Eotrella[277]

Gen. et sp. nov

Valid

Gorochov in Gorochov & Labandeira

Eocene

Green River Formation

 United States

A spider cricket. The type species is E. mira''.

Eozacla[277]

Gen. et 2 sp. nov

Valid

Gorochov in Gorochov & Labandeira

Eocene

Green River Formation

 United States

A spider cricket. The type species is E. arachnomorpha; genus also includes E. problematica.

Heterologus duyiwuer[278]

Sp. nov

Valid

Béthoux, Gu & Ren

Late Carboniferous (Namurian)

Tupo Formation

 China

An archaeorthopteran (stem-orthopteran), a species of Heterologus.

Hirsutgerarus[279]

Gen. et sp. nov

Valid

Zessin

Carboniferous (Stephanian)

 Germany

A gerarid panorthopteran. The type species is Hirsutgerarus saxonicus.

Liassophyllum caii[280]

sp nov

Valid

Gu & Ren in Gu, Qiao, & Ren

Bathonian to Callovian

Jiulongshan Formation

 China

A haglid orthopteran, a species of Liassophyllum.

Miamia maimai[281]

Sp. nov

Valid

Béthoux et al.

Late Carboniferous (early Pennsylvanian)

 China

A stem–orthopteran insect, a species of Miamia.

Protogryllus (Protogryllus) lakshmi[282]

Sp. nov

Valid

Pérez-de La Fuente et al.

Middle Jurassic

Kota Formation

 India

A protogryllid grylloid ensiferan, a species of Protogryllus.

Pteromogoplistes[277]

Gen. et sp. nov

Valid

Gorochov in Gorochov & Labandeira

Eocene

Green River Formation

 United States

A member of the family Mogoplistidae. The type species is P. grandis; genus might also include Pronemobius smithii Scudder (1890).

Pterotriamescaptor? americanus[277]

Sp. nov

Valid

Gorochov in Gorochov & Labandeira

Eocene

Green River Formation

 United States

A mole cricket.

Sunoptera[283]

Gen. et sp. nov

Valid

Hong & Li

Middle Triassic

Tongchuan Formation

 China

A sunopterid "protorthopteran". The type species is Sunoptera orientalis.

Tettoraptor[277]

Gen. et sp. nov

Valid

Gorochov in Gorochov & Labandeira

Eocene

Green River Formation

 United States

A member of the infraorder Tettigoniidea of uncertain phylogenetic placement. The type species is T. maculatus''.

Raphidioptera

Name Novelty Status Authors Age Unit Location Notes Images

Alavaraphidia[284]

Gen et sp. nov

Valid

Pérez-de la Fuente et al.

Albian

Escucha Formation

 Spain

A Mesoraphidiid snakefly. The sole species is A. imperterrita.

Alavaraphidia imperterrita

Amarantoraphidia[284]

Gen et sp. nov

Valid

Pérez-de la Fuente et al.

Albian

Escucha Formation

 Spain

A Mesoraphidiid snakefly. The sole species is A. ventolina.

Amarantoraphidia ventolina

Necroraphidia[284]

Gen. et sp.

Valid

Pérez-de la Fuente et al.

Albian

Las Peñosas Formation

 Spain

A Mesoraphidiid snakefly. The only species is N. arcuata.

Necroraphidia arcuata

Styporaphidia? hispanica[284]

sp. nov

Valid

Pérez-de la Fuente et al.

Albian

Escucha Formation

 Spain

A Mesoraphidiid snakefly. The second species in Styporaphidia.

Styporaphidia? hispanica

other insects

Name Novelty Status Authors Age Unit Location Notes Images

Anebos[285]

Gen. et sp. nov

Valid

Garwood et al.

Late Pennsylvanian

Assise de Montceau Formation

 France

A polyneopteran insect of uncertain affinities. The type species is Anebos phrixos.

Bechala[286]

Gen. et sp. nov

Valid

Ilger & Brauckmann

Late Carboniferous

 Germany

A bechalid megasecopteran. Type species is Bechala sommeri.

Bruayaphis[287]

Gen. et sp. nov

Valid

Nel, Szwedo & Roques in Nel et al.

Late Carboniferous

 France

A member of Paraneoptera of uncertain phylogenetic placement. The type species is Bruayaphis oudardi.

Cucullistriga[288]

Gen. et sp. nov

Valid

Aristov & Rasnitsyn

Kungurian

 Russia

A member of Grylloblattida/Eoblattida belonging to the family Idelinellidae; a relative of ice crawlers or cockroaches.[289] Type species is Cucullistriga cucullata.

Darekia sanguinea[290]

Gen. et sp. nov

Valid

Prokop et al.

Upper Carboniferous

Upper Silesian Coal Basin

 Poland

A stem winged insect.

Eochauliodes[291]

Gen. et sp. nov

Valid

Liu et al.

Middle Jurassic (Bathonian-Callovian)

Jiulongshan Formation

 China

A fishfly. The type species is Eochauliodes striolatus.

Eospilopsyllus[292]

Gen. et sp. nov

Valid

Beaucournu & Perrichot in Perrichot, Beaucournu & Velten

Early Miocene

 Dominican Republic

A pulicid flea found in Dominican amber. The type species is Eospilopsyllus kobberti.

Geinitzia aristovi[293]

sp nov

Valid

Cui, Storozhenko & Ren

Middle Jurassic

Jiulongshan Formation

 China

A geinitziid, a member of Polyneoptera/Gryllones belonging to the group Reculida;[289] a species of Geinitzia.

Gymnopollisthrips[294]

Gen. et 2 sp. nov.

Valid

Peñalver, Nel & Nel in Peñalver et al.

Early Cretaceous

 Spain

A melanthripid thrip. Genus contains two species: Gymnopollisthrips minor and G. maior.

Liadotaulius daohugouensis[295]

sp nov

Valid

Wu & Huang

Middle Jurassic

Daohugou Beds

 China

A caddisfly, a species of Liadotaulius.

Netoxena[296]

Nom. nov

Valid

Sohn in Sohn et al.

Early Cretaceous (late Aptian)

Crato Formation

 Brazil

A lepidopteran, possibly a member of Eolepidopterigidae; a replacement name for Xena Martins-Neto (1999).

Pseudopulex[297]

Gen. et 2 sp. nov

Valid

Gao, Shih, & Ren in Gao et al.

Middle Jurassic to Early Cretaceous

 China

A flea-like insect. The genus contains 2 species: P. jurassicus and P. magnus.

Rasstriga[288]

Gen. et sp. nov

Valid

Aristov & Rasnitsyn

Upper Carboniferous (Desmoinesian)

 United States

A member of Grylloblattida/Eoblattida probably belonging to the family Idelinellidae; a relative of ice crawlers or cockroaches.[289] Its type species is Rasstriga americana.

Scutistriga[288]

Gen. et sp. nov

Valid

Aristov & Rasnitsyn

Kungurian

 Russia

A member of Grylloblattida/Eoblattida belonging to the family Idelinellidae; a relative of ice crawlers or cockroaches.[289] Its type species is Scutistriga scutata.

Sharasialis[298]

Gen. et sp. nov

Valid

Ponomarenko

Late Jurassic

 Mongolia

An alderfly. The type species is Sharasialis fusiformis.

Sinoprotodiplatys[299]

Gen. et sp. nov

Valid

Nel et al.

Early Cretaceous

Yixian Formation

 China

An earwig belonging to the family Protodiplatyidae. The type species is Sinoprotodiplatys zhangi.

Strigulla[288]

Gen. et comb. nov

Valid

Aristov & Rasnitsyn

Kungurian

 Russia

A member of Grylloblattida/Eoblattida belonging to the family Idelinellidae; a relative of ice crawlers or cockroaches.[289] A new genus for "Euryptilon" cuculiophoris (Aristov, 2002).

Strudiella[300]

Gen. et sp. nov.

Valid

Garrouste et al.

Late Devonian

 Belgium

The first complete Late Devonian insect, probably terrestrial. The type species is Strudiella devonica.

Zdenekia silesiensis[290]

Species

Valid

Prokop et al.

Upper Carboniferous

Upper Silesian Coal Basin

 Poland

A stem winged insect.

Eurypterids

Name Novelty Status Authors Age Unit Location Notes Images

Adelophthalmus dubius[301]

Sp. nov

Valid

Shpinev

Middle Devonian

 Russia

A eurypterid, a species of Adelophthalmus.

Adelophthalmus kamyshtensis[301]

Sp. nov

Valid

Shpinev

Middle Devonian

 Russia

A eurypterid, a species of Adelophthalmus.

Drepanopterus odontospathus[302]

Sp. nov

Valid

Lamsdell

Early Devonian

 Canada

A eurypterid.

Nanahughmilleria notosibirica[303]

Sp. nov

Valid

Shpinev

Devonian

 Russia

A eurypterid, a species of Nanahughmilleria.

Parahughmilleria longa[303]

Sp. nov

Valid

Shpinev

Devonian

 Russia

A eurypterid, a species of Parahughmilleria.

Stylonuroides orientalis[303]

Sp. nov

Valid

Shpinev

Devonian

 Russia

A eurypterid, a species of Stylonuroides.

Trilobites

Name Novelty Status Authors Age Unit Location Notes Images

Abanomocarella[304]

Gen. et sp. et comb. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Anomocarellidae. Genus includes new species A. quadrata, as well as A. humilis (Resser & Endo in Endo & Resser, 1937) and A. pyriformis (Resser & Endo in Endo & Resser, 1937).

Acmarhachis kindlei[305]

Sp nov

Valid

Westrop & Eoff

Late Cambrian

Cow Head Group

 Canada

An agnostid, a species of Acmarhachis.

Acmarhachis whittingtoni[305]

Sp nov

Valid

Westrop & Eoff

Late Cambrian

Cow Head Group

 Canada

An agnostid, a species of Acmarhachis.

Angsiduoa jiulongshanensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Ignotogregatidae.

Angsiduoa shanxiensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Ignotogregatidae.

Angsiduoa transversa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Ignotogregatidae.

Anomocarella? antiqua[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Anomocarellidae.

Anomocarella jiulongshanensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Anomocarellidae.

Anomocarella similis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Anomocarellidae.

Aponileus aasei[306]

Sp nov

Valid

Adrain & McAdams

Early Ordovician (late Floian)

Fillmore Formation

 United States

A bathyurid trilobite, a species of Aponileus.

Aponileus belkaae[306]

Sp nov

Valid

Adrain & McAdams

Early Ordovician (late Floian)

Wah Wah Formation

 United States

A bathyurid trilobite, a species of Aponileus.

Aponileus laikaae[306]

Sp nov

Valid

Adrain & McAdams

Early Ordovician (late Floian)

Fillmore Formation

 United States

A bathyurid trilobite, a species of Aponileus.

Aponileus strelkaae[306]

Sp nov

Valid

Adrain & McAdams

Early Ordovician (late Floian)

Wah Wah Formation

 United States

A bathyurid trilobite, a species of Aponileus.

Aponileus ugolekae[306]

Sp nov

Valid

Adrain & McAdams

Early Ordovician (late Floian)

Wah Wah Formation

 United States

A bathyurid trilobite, a species of Aponileus.

Aponileus? veterokae[306]

Sp nov

Valid

Adrain & McAdams

Early Ordovician (late Floian)

Wah Wah Formation

 United States

A bathyurid trilobite, a possible species of Aponileus.

Archaeagnostus typicus[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Peronopsidae.

Archegonus (Phillibole) nehdenensis africanus[307]

Subsp. nov

Valid

Hahn, Müller & Becker

Carboniferous (Mississippian)

 Morocco

A member of Phillipsiidae belonging to the subfamily Archegoninae, a subspecies of Archegonus nehdenensis.

Asaphellus cuervoae[308]

Sp. nov

Disputed

Corbacho & López-Soriano

Ordovician (Floian)

Upper Fezouata Formation

 Morocco

A member of Asaphidae, a species of Asaphellus. Gutiérrez-Marco et al. (2017) considered this species to be likely based on poorly preserved and probably retouched specimens of Asaphellus stubbsi.[309]

Astycoryphe arduinnae[310]

Sp nov

Valid

Van Viersen, van Rossum & Prescher

Middle Devonian

 Belgium

A member of Proetida.

Australoscutellum iota[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Australoscutellum.

Australoscutellum microps[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Australoscutellum.

Australoscutellum pulex[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Australoscutellum.

Australoscutellum trica[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Australoscutellum.

Baltagnostus yaojiayuensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Diplagnostidae.

Baojingia (Eobaojingia)[304]

Subgen. et 2 sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Lisaniidae. The subgenus includes Baojingia (Eobaojingia) dubia and Baojingia (Eobaojingia) ezhuangensis, as well as Baojingia miaogouensis Mong in Zhou et al. (1977).

Borenoria[311]

gen et sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite. The type species is Borenoria cirrita.

Bumastoides graffhami[312]

Sp nov

Valid

Carlucci et al.

Ordovician

 United States

An illaenid trilobite, a species of Bumastoides.

Bumastoides kimmswickensis[312]

Sp nov

Valid

Carlucci et al.

Ordovician

 United States

An illaenid trilobite, a species of Bumastoides.

Bumastoides moundensis[312]

Sp nov

Valid

Carlucci et al.

Ordovician

 United States

An illaenid trilobite, a species of Bumastoides.

Calymene harringtoni[313]

Sp. nov

Valid

Tortello in Tortello et al.

Early Silurian

Vargas Peña Formation

 Paraguay

A species of Calymene.

Chlupacula (Chlupacula) erfoudensis[307]

Sp. nov

Valid

Hahn, Müller & Becker

Carboniferous (Mississippian)

 Morocco

A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Chlupacula.

Crepicephalina exsculpta[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Crepicephalina obscura[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Crepicephalina pulchra[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Crepicephalina? rara[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Crepicephalina woqishanensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Cummingella boikoi[314]

sp nov

Valid

Mychko

Sakmarian

 Russia

A phillipsiid, a species of Cummingella.

Cyrtoproetus (Crassibole) acrops[307]

Sp. nov

Valid

Hahn, Müller & Becker

Carboniferous (Mississippian)

 Morocco

A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Cyrtoproetus.

Dalejeproetus owensi[315]

Sp nov

Valid

Johnson & Fortey

Early Devonian (Pragian)

Ihandar Formation

 Morocco

A proetid trilobite, a species of Dalejeproetus.

Dalejeproetus sagaouii[315]

Sp nov

Valid

Johnson & Fortey

Early Devonian (Pragian)

Ihandar Formation

 Morocco

A proetid trilobite, a species of Dalejeproetus.

Dameselloides[304]

Gen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Damesellidae. Genus includes new species D. sinensis.

Danjiangella shandongensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Dechenelloides tazoultensis[307]

Sp. nov

Valid

Hahn, Müller & Becker

Carboniferous (Mississippian)

 Morocco

A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Dechenelloides.

Decoroscutellum planes[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Decoroscutellum.

Derikaspis conicus[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Derikaspis ruichengensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Diacoryphe (Archaeocoryphe) maiderensis[307]

Sp. nov

Valid

Hahn, Müller & Becker

Carboniferous (Mississippian)

 Morocco

A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Diacoryphe.

Dindymene whittingtoni[316]

Sp nov

Valid

Owen & Bruton

Late Ordovician

 United States

A species of Dindymene.

Dorypyge cylindrica[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Dorypyge zhangxianensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Dunopyge seiberti[317]

Sp. nov

Valid

Basse & Müller

Early Devonian

Seifen Beds

 Germany

A member of Acastidae belonging to the subfamily Asteropyginae, a species of Dunopyge.

Eilura trapezialis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Elaphraella? taebaeksanensis[318]

Sp. nov

Valid

Park & Choi

Cambrian (middle Furongian)

Sesong Formation

 South Korea

A member of the family Shumardiidae.

Emmrichella punctata[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Ptychopariidae.

Eokoldinia[304]

Gen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Plethopeltidae. Genus includes new species E. porifera.

Eomapania[304]

Gen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Mapaniidae. Genus includes new species E. subquadrata.

Eoscutellum annosum[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Eoscutellum.

Eymekops pristina[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Eymekops transversa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Grandioculus macroculus[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Grandioculus (Protohedina) quadraticephala[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Gumunsoia[319]

Gen. et sp. nov

Valid

Park, Sohn & Choi

Cambrian (middle Furongian)

Sesong Formation

 South Korea

A member of Leiostegioidea belonging to the family Kaolishaniidae. Genus includes new species G. triangularis.

Gushanella[304]

Gen. et 2 sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Solenopleuridae. Genus includes new species G. quadrata and G. truncata.

Heyelingella crassa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Honania porifera[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Huayuanaspis angustilimbata[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Idioura brevispina[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Illaenoscutellum tryo[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Illaenoscutellum.

Inouyella rara[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Inouyella shiliuyuanensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Inouyia (Bulbinouyia)[304]

Subgen. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Inouyiidae. The subgenus includes Inouyia kangzhuangensis (Zhang & Wang, 1985) and I. lubrica (Zhou in Zhou et al., 1982).

Inouyia latilimbata[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Iranoleesia pustulosa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Iranoleesia (Proasaphiscina) rogosa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Japonoscutellum borenorense[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Japonoscutellum.

Japonoscutellum crassum[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Japonoscutellum.

Japonoscutellum diascetum[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Japonoscutellum.

Japonoscutellum gephyricum[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Japonoscutellum.

Japonoscutellum guttulatum[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Japonoscutellum.

Japonoscutellum serangicum[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Japonoscutellum.

Japonoscutellum strabo[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Japonoscutellum.

Jiulongshania shandongensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Diceratocephalidae.

Koptura (Eokoptura)[304]

Subgen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Anomocaridae. The subgenus includes new species Koptura (Eokoptura) bella, as well as Koptura quadrata (Resser & Endo in Endo & Resser, 1937).

Koptura intermedia[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Anomocaridae.

Kosovopeltis trepida[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Kosovopeltis.

Kosovopeltis xynon[311]

sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite, a species of Kosovopeltis.

Lepidoproetus lahceni[315]

Sp nov

Valid

Johnson & Fortey

Early Devonian (Pragian)

Ihandar Formation

 Morocco

A proetid trilobite, a species of Lepidoproetus.

Lepidoproetus maharchianus[315]

Sp nov

Valid

Johnson & Fortey

Early Devonian (Pragian)

Ihandar Formation

 Morocco

A proetid trilobite, a species of Lepidoproetus.

Lepidoproetus splendens[315]

Sp nov

Valid

Johnson & Fortey

Early Devonian (Pragian)

Ihandar Formation

 Morocco

A proetid trilobite, a species of Lepidoproetus.

Liowutaishania laevigata[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Lisania elliptica[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Longiproetus meiopsifius[310]

Sp nov

Valid

Van Viersen, van Rossum & Prescher

Middle Devonian

 Belgium

A member of Proetida.

Lunania[304]

Gen. nov

Junior homonym

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Menocephalidae. The generic name is preoccupied by Lunania Chow (1957).

Maotunia transversa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Mapania (Mapanella)[304]

Subgen. et 2 sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Mapaniidae. The subgenus includes new species M. (Mapanella) transversa and M. (Mapanella) prisca, as well as Mapania blackwelderi (Resser & Endo in Endo & Resser, 1937).

Mapania subcylindrica[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Mapanopsis conicus[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Maurotarion megacephalum[320]

Sp. nov.

Valid

Rustán & Vaccari

Early Devonian (Pragian)

Talacasto Formation

 Argentina

An otarionine aulacopleurid, a species of Maurotarion.

Megagraulos similis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Menocephalina[304]

Gen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Menocephalidae. Genus includes new species M. longispinosa.

Menocephalites (Hutoushania)[304]

Subgen. et 2 sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Menocephalidae. The subgenus includes Menocephalites (Hutoushania) triangulata and M. (Hutoushania) zaozhuangansis.

Menocephalites porosus[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Menocephalites primitivus[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Menocephalites simplex[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Metanomocarella primitiva[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Micagnostus semicircularis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Mirrabooka[311]

gen et sp nov

Valid

Holloway and Lane

Silurian

 Australia

A scutelluid trilobite. The type species is Mirrabooka harryi.

Monkaspis conica[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Ohleum magreani[321]

Sp nov

Valid

Taghon, Bonino & Mottequin

Devonian (Eifelian)

Jemelle Formation

 Belgium

A member of Lichida belonging to the family Lichidae.

Pagetides qianensis[322]

Sp nov

Valid

Zhang & Clarkson

Cambrian

 China

An eodiscoid trilobite, a species of Pagetides.

Paleonelsonia granulosa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Ptychopariidae.

Paleonelsonia sulcata[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Ptychopariidae.

Paracoosia langgongsiensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Anomocaridae.

Paralevisia subglobosa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Parayujinia pulchra[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Peishania abnormis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Peishania obscura[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Peronopsis jiulongshanensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Peronopsis universalis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Persia[323]

Gen. et sp. nov

Valid

Lerosey-Aubril

Carboniferous (Tournaisian)

 Iran

The type species is Persia praecox.

Pilletina seifenensis[317]

Sp. nov

Valid

Basse & Müller

Early Devonian

Seifen Beds

 Germany

A member of Acastidae belonging to the subfamily Asteropyginae, a species of Pilletina.

Pingluia granulosa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Plectocrania elegans[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Plectocrania intermedia[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Plesigangderria latilimbata[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Podoliproetus mirdani[315]

Sp nov

Valid

Johnson & Fortey

Early Devonian (Pragian)

Ihandar Formation

 Morocco

A proetid trilobite, a species of Podoliproetus.

Podoliproetus sinespina[315]

Sp nov

Valid

Johnson & Fortey

Early Devonian (Pragian)

Ihandar Formation

 Morocco

A proetid trilobite, a species of Podoliproetus.

Praeymekops[304]

Gen. et sp. et comb. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Proasaphiscidae. Genus includes new species P. minutus, as well as P. sinensis (Mansuy, 1916).

Proasaphiscus laiwuensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Proetina ihamadii[315]

Sp nov

Valid

Johnson & Fortey

Early Devonian (Pragian)

Ihandar Formation

 Morocco

A proetid trilobite, a species of Proetina.

Pseudoolenoides fossilmountainensis[324]

Sp nov

Valid

Adrain, McAdams & Karim

Middle Ordovician

Kanosh Formation

 United States

A bathyurid, a species of Pseudoolenoides.

Pseudoolenoides oilcreekensis[324]

Sp nov

Valid

Adrain, McAdams & Karim

Middle Ordovician

Oil Creek Formation

 United States

A bathyurid, a species of Pseudoolenoides.

Pseudoolenoides pogonipensis[324]

Sp nov

Valid

Adrain, McAdams & Karim

Middle Ordovician

Kanosh Formation

 United States

A bathyurid, a species of Pseudoolenoides.

Pseudopeishania[304]

Gen. et 3 sp. et comb. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Anomocarellidae. Genus includes new species P. brevis, P. similis and P. woqishanensis, as well as P. angustata (Endo, 1944).

Pseudoperonopsis elegans[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Pseudophillipsia parvizii[323]

Sp. nov

Valid

Lerosey-Aubril

Permian

 Iran

A species of Pseudophillipsia sensu lato.

Pseudotaitzuia angustilimbata[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Asaphida belonging to the family Ordosiidae.

Pseudotaitzuia rara[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Asaphida belonging to the family Ordosiidae.

Pudoproetus zhorae[307]

Sp. nov

Valid

Hahn, Müller & Becker

Carboniferous (Mississippian)

 Morocco

A member of Proetidae, a species of Pudoproetus.

Raragnostus intermedius[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Raymondaspis grandigena[325]

sp nov

Valid

Stein & Bergström

Middle Ordovician

 Sweden

A styginid, a species of Raymondaspis.

Reediella mitchelli tafilaltensis[307]

Subsp. nov

Valid

Hahn, Müller & Becker

Carboniferous (Mississippian)

 Morocco

A member of Phillipsiidae belonging to the subfamily Bollandiinae, a subspecies of Reediella mitchelli.

Ruichengocephalus[304]

Gen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Ptychopariidae. Genus includes new species R. typicus.

Saimachia yichangensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Schopfaspis? graciai[326]

sp nov

Valid

Esteve et al..

Middle Cambrian

 Spain

An alokistocarid, possibly a species of Schopfaspis.

Sdzuyella conica[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belongig to the family Holocephalidae.

Securifrons[311]

gen et comb nov

Valid

Holloway and Lane

Ordovician to Silurian

Scutelluid trilobite. species- S. borealis, S. cunctatum, S. major and ?S. umbonatus.

Shiliuyuania[304]

Gen. et 2 sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Crepicephalidae. Genus includes new species S. cylindrica and S. quadrata.

Solenoparia zhangxianensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Solenoparina bella[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Sphaerophthalmus arcus[327]

Sp nov

Valid

Høyberget & Bruton

Cambrian (Furongian)

Alum Shale Formation

 Norway

A member of the family Olenidae.

Sudanomocarina elliptica[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Sudanomocarina hutoushanensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Symphysops stevaninae[328]

Sp. nov

Valid

López-Soriano & Corbacho

Late Ordovician

Lower Ktaoua Formation

 Morocco

A member of Cyclopygidae, a species of Symphysops.

Szeaspis (Pseudoszeaspis)[304]

Subgen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Anomocarellidae. The subgenus includes new species Szeaspis (Pseudoszeaspis) conicus, as well as Szeaspis huananensis Peng, Babcock & Lin (2004) and Szeaspis pustulosa (Resser & Endo in Endo & Resser, 1937).

Szeaspis crassus[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Anomocarellidae.

Taitzehoia huolianzhaiensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Damesellidae.

Teinistion jiulongshanense[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Damesellidae.

Teinistion quadratum[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Damesellidae.

Teinistion yangjiazhuangense[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Damesellidae.

Tomagnostus (Paratomagnostus)[304]

Subgen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Ptychagnostidae. The subgenus includes Tomagnostus (Paratomagnostus) ezhuangensis.

Trachoparia elongata[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Trachoparia intermedia[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Trachoparia spinosa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Triangulopyge[327]

Gen. et comb. nov

Valid

Høyberget & Bruton

Cambrian

 Norway
 Sweden
 United Kingdom

A member of the family Olenidae. The type species is "Olenus" humilis Phillips (1848); genus also includes "Sphaerophthalmus" majusculus Linnarsson (1880) and T. major (Lake, 1913).

Tsunyidiscus pengshuiensis[322]

Sp nov

Valid

Zhang & Clarkson

Cambrian

 China

An eodiscoid trilobite, a species of Tsunyidiscus.

Ulania concava[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Utagnostus primitivus[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Clavagnostidae.

Utagnostus sulcatus[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Clavagnostidae.

Woqishanaspis[304]

Gen. et comb. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Ignotogregatidae. Genus includes W. trapezoidalis (Lin, 1984).

Wutaishanaspis yangjiazhuangensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Xenolorenzella granulosa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Yichengocephalus[304]

Gen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Solenopleuridae. Genus includes new species Y. latus.

Yongwolia quadrata[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Yushugouia ezhuangensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Zhangxiaspis[304]

Gen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Ptychopariidae. Genus includes new species Z. subcylindrica.

Zhongtiaoshanella[304]

Gen. et sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Crepicephalidae. Genus includes new species Z. brevispinosa.

Zhongweia convexa[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

Zhujia yangjiazhuangensis[304]

Sp. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of Ptychopariida belonging to the family Anomocarellidae.

Zichuanaspis[304]

Gen. et 2 sp. et comb. nov

Valid

Yuan et al.

Cambrian

Changhia Formation

 China

A member of the family Leiostegiidae. Genus includes Zichuanaspis acerius (Walcott, 1905) and Z. catinus (Resser & Endo in Endo & Resser, 1937), as well as new species Z. yangjiazhuangensis and possibly Z? latilimbata.

Others

Name Novelty Status Authors Age Unit Location Notes Images

Australimicola[329]

Gen. et sp.

Valid

Paterson, García-Bellido and Edgecombe

Early Cambrian

The Emu Bay Shale Konservat-Lagerstätte

 Australia

An artiopodan arthropod. The type species is Australimicola spriggi.

Kangacaris shui[330]

Sp. nov

Valid

Zhang, Fu & Dai

Lower Cambrian

Chengjiang lagerstätte

 China

A species of Kangacaris.

Luohuilinella[331]

Gen. et sp. nov

Valid

Zhang, Fu & Dai

Lower Cambrian

Chengjiang Lagerstätte

 China

A possible member of Xandarellida. The type species is Luohuilinella rarus.

Nereocaris[332]

Gen. et sp. nov

Valid

Legg et al.

Cambrian

Burgess Shale Formation

 Canada

A bivalved arthropod, one of the basalmost known arthropods. The type species is Nereocaris exilis.

Pleuralata[333]

Gen. et sp. nov

Valid

McCoy, Strother and Briggs

Silurian (Llandovery)

Tuscarora Formation

 United States

An arthropod of uncertain phylogenetic affinities, a probable tracemaker of Arthrophycus alleghaniensis trace fossils. The type species is Pleuralata spinosa.

Scolopocryptops simojovelensis[334]

Sp. nov

Valid

Edgecombe et al.

Miocene

 Mexico

A scolopocryptopid scolopendromorph centipede, a species of Scolopocryptops.

Sinoleperditia hamadai[335]

Sp. nov

Valid

Tanaka, Ono and Maeda

Early Devonian

 Japan

A member of Leperditicopida (a group of arthropods of uncertain affinities, classified as ostracods or excluded from this group by different authors), a species of Sinoleperditia.

Sinoleperditia huyeni[336]

Sp. nov

Valid

Tanaka, Phong and Komatsu

Early Devonian

 Vietnam

A member of Leperditicopida, a species of Sinoleperditia.

Squamacula buckorum[329]

Species

Valid

Paterson, García-Bellido and Edgecombe

Early Cambrian

The Emu Bay Shale Konservat-Lagerstätte

 Australia

An artiopodan arthropod, a species of Squamacula.

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  252. Yongjie Wang; Conrad C. Labandeira; Chungkun Shih; Qiaoling Ding; Chen Wang; Yunyun Zhao; Dong Ren (2012). "Jurassic mimicry between a hangingfly and a ginkgo from China". Proceedings of the National Academy of Sciences of the United States of America. 109 (50): 20514–20519. doi:10.1073/pnas.1205517109. PMC 3528590. PMID 23184994.
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  255. Agnieszka Soszyńska-Maj; Wiesław Krzemiński; Katarzyna Kopeć; Yizi Cao; Dong Ren (2018). "Large Jurassic scorpionflies belonging to a new subfamily of the family Orthophlebiidae (Mecoptera)". Annales Zoologici. 68 (1): 85–92. doi:10.3161/00034541ANZ2018.68.1.004.
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  257. Vladimir N. Makarkin; Qiang Yang; YuanYuan Peng; Dong Ren (2012). "A comparative overview of the neuropteran assemblage of the Lower Cretaceous Yixian Formation (China), with description of a new genus of Psychopsidae (Insecta: Neuroptera)". Cretaceous Research. 35: 57–68. doi:10.1016/j.cretres.2011.11.013.
  258. Chao-Fan Shi; Yong-Jie Wang; Qiang Yang; Dong Ren (2012). "Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia, China". Alcheringa: An Australasian Journal of Palaeontology. 36 (3): 309–318. doi:10.1080/03115518.2012.644994.
  259. Chaofan Shi; Olivier Béthoux; Chungkun Shih; Dong Ren (2012). "Guyiling jianboni gen. et sp.n., an antlion-like lacewing, illuminating homologies and transformations in Neuroptera wing venation". Systematic Entomology. 37 (3): 617–631. doi:10.1111/j.1365-3113.2012.00633.x.
  260. Ricardo Pérez-de la Fuente; Xavier Delclòs; Enrique Peñalver; Mariela Speranza; Jacek Wierzchos; Carmen Ascaso; Michael S. Engel (2012). "Early evolution and ecology of camouflage in insects". Proceedings of the National Academy of Sciences of the United States of America. 109 (52): 21414–21419. doi:10.1073/pnas.1213775110. PMC 3535654. PMID 23236135.
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  263. Qiang Yang; Vladimir N. Makarkin; Shaun L. Winterton; Alexander V. Khramov; Dong Ren (2012). "A Remarkable New Family of Jurassic Insects (Neuroptera) with Primitive Wing Venation and Its Phylogenetic Position in Neuropterida". PLOS ONE. 7 (9): e44762. doi:10.1371/journal.pone.0044762. PMC 3445537. PMID 23028608.
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  267. Beattie, R.G.; Nel, A. (2012). "A new dragonfly, Austroprotolindenia jurassica (Odonata: Anisoptera), from the Upper Jurassic of Australia". Alcheringa: An Australasian Journal of Palaeontology. 36 (2): 189–193. doi:10.1080/03115518.2011.598055.
  268. Günter Bechly (2012). "An interesting new fossil relict damselfly (Odonata: Zygoptera: Coenagrionoidea) from Eocene Baltic amber" (PDF). Palaeodiversity. 5: 51–55.
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  279. Wolfgang Zessin (2012). "Eine weitere neue Geraride, Hirsutgerarus saxonicus n. gen. n. sp. (Insecta: Panorthoptera: Geraridae), aus dem Oberkarbon (Stephanium C) von Plötz bei Halle (Deutschland)" (PDF). Virgo, Mitteilungsblatt des Entomologischen Vereins Mecklenburg. 15 (1): 10–17.
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