2016 in arthropod paleontology

This list of fossil arthropods described in 2016 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2016, as well as other significant discoveries and events related to arthropod paleontology that occurred in the year 2016.

List of years in arthropod paleontology
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In paleontology 2013 2014 2015 2016 2017 2018 2019
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Arachnids

Research

• A study on the phylogenetic relationships of the Eocene tick Ixodes succineus is published by Dunlop et al. (2016).[1]
• A study on the phylogenetic relationships of the Carboniferous species Archaeometa nephilina and Arachnometa tuberculata is published by Selden, Dunlop & Garwood (2016), who interpret these taxa as harvestmen rather than spiders.[2]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aenigmatarbus[3]	Gen. et sp. nov	Valid	Poschmann et al.	Carboniferous (late Gzhelian)		France	A member of Trigonotarbida of uncertain phylogenetic placement. The type species is Aenigmatarbus rasteli.
Afronothrus ornosae[4]	Sp. nov	Valid	Arillo, Subías & Sánchez-García	Early Cretaceous		Spain	An oribatid mite, a species of Afronothrus.
Archaeoananteroides[5]	Gen. et sp. nov	Valid	Lourenço & Velten	Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the family Buthidae. The type species is A. maderai.
Betaburmesebuthus bellus[6]	Sp. nov	Valid	Lourenço	Early Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the superfamily Buthoidea and the family Palaeoburmesebuthidae.	Betaburmesebuthus bellus
Betaburmesebuthus fleissneri[7]	Sp. nov	Valid	Lourenço & Velten	Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the superfamily Buthoidea and the family Palaeoburmesebuthidae.[6]
Betaburmesebuthus larafleissnerae[8]	Sp. nov	Valid	Lourenço & Velten	Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the superfamily Buthoidea and the family Palaeoburmesebuthidae.
Burmesescorpiops[9]	Gen. et sp. nov	Valid	Lourenço	Early Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the family Palaeoeuscorpiidae and the subfamily Archaeoscorpiopinae. The type species is Burmesescorpiops groehni.
Calyptostoma katyae[10]	Sp. nov	Valid	Konikiewicz, Wohltmann & Mąkol	Eocene	Baltic amber	Europe (Baltic Sea coast)	A mite belonging to the group Prostigmata, a species of Calyptostoma.
Carbolohmannia[11]	Gen. et sp. nov	Valid	Robin et al.	Carboniferous (ca. 320 mya)		China	An oribatid mite belonging to the group Mixonomata. The type species is Carbolohmannia maimaiphilus.
Chaerilobuthus gigantosternum[9]	Sp. nov	Valid	Lourenço	Early Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the family Chaerilobuthidae; a species of Chaerilobuthus.
Chaerilobuthus serratus[9]	Sp. nov	Valid	Lourenço	Early Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the family Chaerilobuthidae; a species of Chaerilobuthus.
Cheletomimus (Hemicheyletia) crinitus[12]	Sp. nov	Valid	Bochkov & Sidorchuk	Eocene	Baltic amber	Baltic Sea coast, most likely  Russia (Kaliningrad Oblast)	A mite belonging to the family Cheyletidae, a species of Cheletomimus.
Electroblemma[13]	Gen. et sp. nov	Valid	Selden, Zhang & Ren	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Tetrablemmidae. The type species is E. bifida.
Electrokoenenia[14]	Gen. et sp. nov	Valid	Engel & Huang in Engel et al.	Late Cretaceous (Cenomanian)		Myanmar	A microwhip scorpion. The type species is Electrokoenenia yaksha.
Hypovertex hispanicus[4]	Sp. nov	Valid	Arillo, Subías & Sánchez-García	Early Cretaceous		Spain	An oribatid mite, a species of Hypovertex.
Idmonarachne[15]	Gen. et sp. nov	Valid	Garwood et al.	Carboniferous (Gzhelian)		France	A member of Arachnida of uncertain phylogenetic placement, probably closely related to spiders. The type species is Idmonarachne brasieri.
Kronocharon prendinii adornellae[16]	Subsp. nov	Valid	Rossi & Merendino	Cretaceous	Burmese amber	Myanmar	A whip spider.
Nothrus vazquezae[4]	Sp. nov	Valid	Arillo, Subías & Sánchez-García	Early Cretaceous		Spain	An oribatid mite, a species of Nothrus.
?Opsieobuthus tungeri[17]	Sp. nov	Valid	Dunlop et al.	Early Permian (Sakmarian)	Leukersdorf Formation	Germany	A scorpion belonging to the family Centromachidae; possibly a species of Opsieobuthus.
Parageralinura marsiglioi[18]	Sp. nov	Valid	Selden, Dunlop & Simonetto	Carboniferous (Kasimovian–Gzhelian)	Probably Pizzul Formation	Italy	A member of Thelyphonida, a species of Parageralinura.
Paraliochthonius miomaya[19]	Sp. nov	Valid	Judson	Miocene		Mexico	A pseudoscorpion belonging to the family Chthoniidae; a species of Paraliochthonius.
Petrobunoides[20]	Gen. et sp. nov	Valid	Selden et al.	Late Cretaceous (early Cenomanian)	Burmese amber	Myanmar	A member of Opiliones belonging to the family Epedanidae. The type species is Petrobunoides sharmai.
Platyliodes sellnicki[4]	Sp. nov	Valid	Arillo, Subías & Sánchez-García	Early Cretaceous		Spain	An oribatid mite, a species of Platyliodes.
Porttrombidium gedanense[21]	Sp. nov	Valid	Konikiewicz, Sontag & Mąkol	Eocene (Lutetian)	Baltic amber	Russia (Kaliningrad Oblast)	A mite belonging to the group Parasitengona and the family Microtrombidiidae.
Protofeaella[22]	Gen. et sp. nov	Valid	Henderickx in Henderickx & Boone	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A pseudoscorpion. Originally classified as a member of the family Feaellidae;[22] Wolfe et al. (2016) only considered it to be a likely member of the pseudoscorpion crown group, probably a feaelloid.[23] The type species is P. peetersae.
Rhopalurus renelauerae[24]	Sp. nov	Valid	Lourenço in Lourenço & Velten	Miocene	Dominican amber	Dominican Republic	A scorpion belonging to the family Buthidae, a species of Rhopalurus.
Tenuelamellarea estefaniae[4]	Sp. nov	Valid	Arillo, Subías & Sánchez-García	Early Cretaceous		Spain	An oribatid mite, a species of Tenuelamellarea.

Crustaceans

Research

• Mesoprosopon triasinum is reinterpreted as a eumalacostracan larva by Hyžný, Haug & Haug (2016).[25]
• A study on the phylogenetic relationships of members of the genus Tealliocaris is published by Jones et al. (2016).[26]
• A study on the phylogenetic relationships of Antrimpos speciosus, indicating it was closely related to shrimps belonging to the genera Farfantepenaeus, Litopenaeus, Fenneropenaeus, Penaeus, Marsupenaeus and Melicertus, is published by Robalino et al. (2016).[27]
• A study on the shapes and sizes of eyes and ommatidia of Jurassic polychelidans is published by Audo et al. (2016).[28]
• New anatomical data on Hesslandona ventrospinata and Hesslandona angustata is published by Eriksson et al. (2016).[29]
• Early Miocene harpacticoid copepods belonging to five families (Canthocamptidae, Cletodidae, Darcythompsoniidae, Ectinosomatidae and Laophontidae) are described from Chiapas amber (Mexico) by Huys et al. (2016).[30]
• Fossil ephippia of members of the daphniid genus Ceriodaphnia are described from the Early Cretaceous (Aptian) Koonwarra Fossil Bed (Strzelecki Group; Victoria, Australia) by Hegna & Kotov (2016).[31]

New taxa

Malacostracans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acanthodromia zannatoi[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Dynomenidae.
Acanthogalathea broglioi[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A member of Galatheidae.
Acanthogalathea devecchii[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A member of Galatheidae.
Acanthogalathea paucispinosa[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A member of Galatheidae.
Aetocarcinus[33]	Gen. et comb. et sp. nov	Valid	Schweitzer et al.	Early Cretaceous (Albian)	Glen Rose Formation	United States	A member of Raninoida belonging to the family Orithopsidae. The type species is "Diaulax" roddai Bishop (1983); genus also includes new species Aetocarcinus muricatus.
Alponella[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the superfamily Goneplacoidea and the family Euryplacidae. The type species is A. paleogenica.
Amekicarcinus[34]	Gen. et sp. nov	Valid	Schweitzer, Odumodu & Feldmann	Eocene		Nigeria	A crab. Genus includes new species A. enigmaticus.
Arcantitanais[35]	Gen. et sp. nov	Valid	Sánchez-García et al.	Cretaceous (Albian–Cenomanian)	Charentese amber	France	A member of Tanaidacea of uncertain phylogenetic placement. The type species is A. turpis.
Arcticocarcinus[36]	Gen. et comb. nov	Valid	Schweitzer et al.	Paleocene (Danian)		Denmark	A crab belonging to the group Raninoida and the family Necrocarcinidae. A new genus for "Necrocarcinus" insignis Segerberg (1900).
Armadillopsis[35]	Gen. et sp. nov	Valid	Sánchez-García et al.	Late Cretaceous (Cenomanian)	Pyrenean amber	France	A member of Tanaidacea of uncertain phylogenetic placement. The type species is A. rara.
Atherfieldastacus[37]	Gen. et comb. nov	Valid	Simpson in Robin et al.	Early Cretaceous	Atherfield Clay Formation	Colombia[38]  Germany  Mexico  South Africa  Spain[39]  United Kingdom	A member of Glypheoidea belonging to the family Mecochiridae. The type species is "Meyeria" magna M'Coy (1849); genus also includes Atherfieldastacus mexicanus (Rathbun, 1935), Atherfieldastacus rapax (Harbort, 1905) and Atherfieldastacus schwartzi (Kitchin,1908) .
Aulavescus paintenensis[40]	Sp. nov	Valid	Feldmann et al.	Late Jurassic		Germany	A munidopsid galatheoid, a species of Aulavescus.
Barapseudia[41]	Gen. et sp. nov	Valid	Quayle	Eocene	Barton Clay Formation	United Kingdom	A member of Tanaidacea belonging to the family Apseudidae. The type species is Barapseudia prima.
Basadromia[42]	Gen. et sp. nov	Valid	Artal et al.	Eocene (Priabonian)		Spain	A member of Dromioidea, possibly a member of the family Dromiidae. The type species is Basadromia longifrons.
Bathynomus kominatoensis[43]	Sp. nov	Valid	Kato, Kurihara & Tokita	Late Miocene	Amatsu Formation	Japan	A giant isopod.
Biohermia[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the superfamily Homolodromioidea and the family Goniodromitidae. The type species is B. chalmasi.
Bittnereus depressus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Panopeidae.
Bittnereus tumidus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Panopeidae.
Bittnerilia granosa[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Parthenopidae.
Bolcagalathea[32]	Gen. et 3 sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A member of Galatheidae. The type species is B. corallina; genus also includes B. venetica and B. multispinosa.
Bolcapisa[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Epialtidae. The type species is B. giulianae.
Braggilambrus[45]	Gen. et sp. nov	Valid	De Angeli & Caporiondo	Eocene (Ypresian)		Italy	A crab belonging to the family Parthenopidae. The type species is B. tani.
Branchioplax cordata[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Mathildellidae.
Bylgia anjobea[46]	Sp. nov	Valid	Winkler	Late Jurassic	Painten Formation	Germany	A member of Penaeidae, a species of Bylgia.
Calappilia calculosa[47]	Sp. nov	Valid	Rumsey, Klompmaker & Portell	Late Eocene to early Oligocene		United States	A member of Calappidae, a species of Calappilia.
Caporiondolus[48]	Gen. et sp. nov	Valid	De Angeli	Eocene (Priabonian)		Italy	A crab belonging to the group Cyclodorippoida and the family Cymonomidae. The type species is Caporiondolus bericus.
Catillogalathea[49]	Gen. et 3 sp. nov	Valid	Robins et al.	Late Jurassic		Austria  Romania	A member of Galatheoidea. The type species is Catillogalathea falcula; genus also includes Catillogalathea patruliusi and Catillogalathea purcarenensis
Ceratiocaris winneshiekensis[50]	Sp. nov	Valid	Briggs et al.	Ordovician (Darriwilian)	Winneshiek Lagerstätte	United States
Charbelicaris[51]	Gen. et sp. nov	Valid	Haug et al.	Late Cretaceous (Cenomanian)	Hadjoula Lagerstätte	Lebanon	A member of Achelata. The type species is Charbelicaris maronites.
Corallioplax[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the superfamily Goneplacoidea and the family Euryplacidae. The type species is C. exigua.
Costacopluma mamethioupamei[52]	Sp. nov	Valid	Hyžný et al.	Late Cretaceous (Maastrichtian)	Cap de Naze Formation	Senegal	A retroplumid crab, a species of Costacopluma.
Cracchidynomene[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Dynomenidae. The type species is C. areolata.
Dardanus bayani[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A species of Dardanus.
Dardanus colosseus[53]	Sp. nov	Valid	Fraaije & Polkowsky	Eocene		Austria	A hermit crab, a species of Dardanus.
Dardanus suessi[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A species of Dardanus.
Discutiolira[49]	Gen. et comb. nov	Valid	Robins et al.	Late Jurassic		Austria  Czech Republic  Poland  Romania	A member of Galatheoidea. The type species is "Galathea" eutecta Moericke (1899); genus also includes "Galathea" moesica Muţiu & Bădăluţă (1971).
Disipia[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A porcelain crab. The type species is D. sorbinii.
Dromilites montenati[54]	Sp. nov	Valid[55]	Robin et al.	Paleocene (Danian)		France	A member of Dromiidae.
Dromiopsis ceratoi[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Dromiidae.
Dromiopsis longitudovata[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Dromiidae.
Dromiopsis parvula[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Dromiidae.
Dromiopsis vicetinus[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab.
Dynomene vetusta[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Dynomenidae.
Elektrocarcinus[36]	Gen. et comb. nov	Valid	Schweitzer et al.	Cretaceous (Albian to Maastrichtian)		United Kingdom  United States	A crab belonging to the group Raninoida and the family Necrocarcinidae. A new genus for "Necrocarcinus" pierrensis Rathbun (1917); genus also includes Elektrocarcinus davisi (Bishop, 1985), E. olsonorum (Bishop and Williams, 1991), E. texensis (Rathbun, 1935) and E. woodwardi (Bell, 1863).
Enoploclytia augustobonae[56]	Sp. nov	Valid	Devillez et al.	Early Cretaceous (Barremian)		France	A member of Erymidae.
Enoploclytia gigantea[56]	Sp. nov	Valid	Devillez et al.	Early Cretaceous (Albian)		United States	A member of Erymidae.
Eoacanthacaris[44]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A lobster. Genus includes new species E. tethysianus.
Eocarpilius lessineus[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Carpiliidae.
Eocharybdis rugosa[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Portunidae.
Eogeryon[57]	Gen. et sp. nov	Valid	Ossó	Late Cretaceous (Cenomanian)		Spain	A member of Portunoidea. The type species is E. elegius.
Eomunidopsis prealpina[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A squat lobster belonging to the family Munidopsidae.
Eosadayoshia[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A member of Munididae. The type species is E. bolcensis.
Eoxanthops[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Xanthidae. The type species is E. scutatus.
Eryma vocontii[56]	Sp. nov	Valid	Devillez et al.	Early Cretaceous (Albian)		France	A member of Erymidae.
Eumorphactaea convexa[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Pilumnidae.
Eurotanais pyrenaensis[35]	Sp. nov	Valid	Sánchez-García et al.	Late Cretaceous (Cenomanian)	Pyrenean amber	France	A member of Tanaidacea belonging the family Alavatanaidae.
Eurotanais seilacheri[35]	Sp. nov	Valid	Sánchez-García et al.	Late Cretaceous (Turonian)	Vendean amber	France	A member of Tanaidacea belonging the family Alavatanaidae.
Faxegalathea valeccensis[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A squat lobster. Originally assigned to the family Munidopsidae and to the genus Faxegalathea; subsequently transferred to the genus Tethysmunida by de Angeli & Ceccon (2017),[58] but this transfer was rejected by Beschin et al. (2019).[59]
Folguerolesia[60]	Gen. et comb. nov	Valid	Artal & Hyžný	Middle Eocene		Spain	A member of Leucosioidea belonging to the new family Folguerolesiidae; a new genus for "Typilobus" boscoi Vía Boada (1959).
Fornicaris[61]	Gen. et sp. nov	Valid	Wilson & Selden in Selden et al.	Late Triassic (Norian)	Dolomia di Forni Formation	Italy	A member of Asellota belonging to the family Paramunnidae. The type species is Fornicaris calligarisi.
Frontelata[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the superfamily Carpilioidea and the family Palaeoxanthopsidae. The type species is F. spinacomposita.
Gabaleryon[62]	Gen. et 2 sp. et comb. nov	Valid[63]	Audo et al.	Early Jurassic (Toarcian)		France  Germany  Italy  United Kingdom	A member of Polychelida. Genus includes new species Gabaleryon coquelae and Gabaleryon garassinoi, as well as "Eryon" moorei.
Galathea lovarica[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A species of Galathea.
Galatheites aiola[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones Štramberk Limestones	Austria  Czech Republic	A member of Galatheoidea, a species of Galatheites.
Galatheites diasema[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones	Austria	A member of Galatheoidea, a species of Galatheites.
Galatheites obtecta[49]	Sp. nov	Valid	Robins et al.	Late Jurassic		Romania	A member of Galatheoidea, a species of Galatheites.
Gecchelicarcinus zanderigoi[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Macropipidae.
Glabropilumnus trispinosus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Pilumnidae.
Glyphithyreus bendensis[34]	Sp. nov	Valid	Schweitzer, Odumodu & Feldmann	Eocene		Nigeria	A crab.
Graptocarcinus eocenicus[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Dynomenidae.
Haydnella granosa[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Xanthidae.
Hispanigalathea tithonia[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones	Austria	A member of Galatheoidea, a species of Hispanigalathea.
Hohensteiniella[64]	Gen. et sp. nov	Valid[65]	Haug et al.	Early Devonian	Hunsrück Slate	Germany	A crustacean with eumalacostracan affinities. The type species is H. engeli.
Iphiculus eliasi[66]	Sp. nov	Valid	Hyžný & Gross	Miocene	Florian Beds	Austria  Portugal[67]  Spain[67]	A crab belonging to the superfamily Leucosioidea and the family Iphiculidae.
Iraticaris[68]	Gen. et sp. nov	Valid	Adami-Rodrigues & Pazinato in Adami-Rodrigues, Pazinato & Pinto	Early Permian	Irati Formation	Brazil	A member of Pygocephalomorpha. The type species is I. damianii.
Karyosia[33]	Gen. et sp. nov	Valid	Schweitzer et al.	Early Cretaceous (Albian)	Glen Rose Formation Walnut Formation	United States	A member of Etyiidae. The type species is Karyosia apicava.
Kromtitis pseudolothi[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab.
Laevicarcinus serratus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Panopeidae.
Lemacola[49]	Gen. et 3 sp. nov	Valid	Robins et al.	Late Jurassic		Austria  Romania	A member of Galatheoidea. The type species is Lemacola jenniferae; genus also includes L. rossi and L. salia
Lessinipagurus boschettensis[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A hermit crab.
Lessinipagurus vasjamikuzi[69]	Sp. nov	Valid	Gašparič et al.	Eocene (late Bartonian–early Priabonian)		Croatia	A hermit crab related to members of the genus Xylopagurus.
Libinia amplissimus[70]	Sp. nov	Valid	Feldmann & Schweitzer	Miocene	St. Marys Formation	United States	A species of Libinia.
Liocarcinus priscus[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Carcinidae.
Lovaracarpilius[44]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Carpiliidae. The type species is L. incisus.
Lovaroides[44]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the superfamily Goneplacoidea and the family Carinocarcinoididae. The type species is L. elegans.
Macrocheira columbiaensis[71]	Sp. nov	Valid	Nyborg et al.	Early to middle Miocene	Astoria Formation	United States	A relative of the Japanese spider crab.
Macrocheira jayi[71]	Sp. nov	Valid	Nyborg et al.	Late Eocene to Oligocene	Hesquiat Formation	Canada	A relative of the Japanese spider crab.
Macrocheira sullivani[71]	Sp. nov	Valid	Nyborg et al.	Late Eocene to early Oligocene	Keasey Formation	United States	A relative of the Japanese spider crab.
Magyarcarcinus yebraensis[72]	Sp. nov	Valid	Domínguez & Ossó	Eocene (Bartonian)	Margas de Arguís Formation	Spain	A crab belonging to the superfamily Goneplacoidea, assigned to the new family Magyarcarcinidae.
Mesogalathea macra[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones	Austria	A member of Galatheoidea, a species of Mesogalathea.
Mesogalathea pyxis[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones	Austria	A member of Galatheoidea, a species of Mesogalathea.
Mesogalathea retusa[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones	Austria	A member of Galatheoidea, a species of Mesogalathea.
Metopocarcinus fragilis[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Panopeidae.
Miopipus zovensis[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Carcinidae.
Mithracia lovatoi[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Majidae.
Montemagralia[73]	Gen. et sp. nov	Valid	De Angeli & Ceccon	Eocene (Ypresian)		Italy	A crab belonging to the family Trapeziidae. The type species is M. lata.
Nanocassiope secretanae[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Xanthidae, a species of Nanocassiope.
Neocallichirus khadroensis[74]	Sp. nov	Valid	Hyžný & Charbonnier in Hyžný et al.	Paleocene (Danian)	Khadro Formation	Pakistan	A member of Callianassidae, a species of Neocallichirus.
Neocallichirus lakhraensis[74]	Sp. nov	Valid	Hyžný & Charbonnier in Hyžný et al.	Eocene (Ypresian)	Lakhra Formation	Pakistan	A member of Callianassidae, a species of Neocallichirus.
Nicoliscarcinus[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Cancridae. The type species is N. rotundatus.
Ophthalmoplax andina[75]	Sp. nov	Valid	Guzmán et al.	Late Cretaceous (late Campanian)	Lodolitas de Aguacaliente Formation	Colombia	A member of Macropipidae, a species of Ophthalmoplax.
Paguristes paucituberculatus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A species of Paguristes.
Palaeodromites xestos[33]	Sp. nov	Valid	Schweitzer et al.	Early Cretaceous (Albian)	Walnut Formation	United States	A member of Homolodromioidea belonging to the family Goniodromitidae, a species of Palaeodromites.
Palmyria levigata[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Domeciidae.
Panopeus postalensis[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Panopeidae, a species of Panopeus.
Paracancrinos[51]	Gen. et comb. nov	Valid	Haug et al.	Late Cretaceous (Cenomanian)		Lebanon	A member of Achelata. A new genus for "Cancrinos" libanensis Garassino & Schweigert (2006).
Paracapsulapagurus[76]	Gen. et sp. nov	Valid	Hyžný et al.	Late Cretaceous (Maastrichtian)	Cape de Naze Formation	Senegal	A hermit crab. The type species is P. poponguinensis.
Paracorallomursia[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Calappidae. The type species is P. medizzai.
Paradistefania denticulata[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the superfamily Homolodromioidea and the family Goniodromitidae.
Paradynomene antiqua[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Dynomenidae.
Paragalathea arcella[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones	Austria	A member of Galatheoidea, a species of Paragalathea.
Paragalathea crenarvina[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones Štramberk Limestones	Austria  Czech Republic	A member of Galatheoidea, a species of Paragalathea.
Paragalathea ternata[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones	Austria	A member of Galatheoidea, a species of Paragalathea.
Paragalathea vultuosona[49]	Sp. nov	Valid	Robins et al.	Late Jurassic	Ernstbrunn Limestones	Austria	A member of Galatheoidea, a species of Paragalathea.
Paraocalina silviae[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Carpiliidae.
Paraporcellana[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A porcelain crab. The type species is P. fabianii.
Parhalimede[44]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Galenidae. The type species is P. ornata.
Parthenope menini[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Parthenopidae.
Permocaris[68]	Gen. et sp. nov	Valid	Adami-Rodrigues, Pazinato & Pinto	Early Permian	Irati Formation	Brazil	A member of Pygocephalomorpha. The type species is P. purperae.
Petrochirus minutus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A member of Diogenidae.
Petrolisthes lineatus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A porcelain crab, a species of Petrolisthes.
Petrusia[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Plagusiidae. The type species is P. striata.
Phlyctenodes edwardsi[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Xanthidae, a species of Phlyctenodes.
Pilummede[77]	Gen. et sp. et comb. nov	Valid	Ossó & Clements	Eocene	Castle Hayne Limestone Formation Clairbone Group	United States	A member of Pilumnoidea of uncertain phylogenetic placement. The type species is P. penderensis; genus also includes "Lobonotus" bakeri Rathbun (1935).
Pilumnomimus dorsocarinatus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the superfamily Xanthoidea of uncertain phylogenetic placement.
Pilumnomimus miettoi[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the superfamily Xanthoidea of uncertain phylogenetic placement.
Pittinucaris[68]	Gen. et sp. nov	Valid	Adami-Rodrigues, Pazinato & Pinto	Early Permian	Irati Formation	Brazil	A member of Pygocephalomorpha. The type species is P. wuerdigae.
Planocorystes[78]	Gen. et sp. nov	Valid	Franţescu, Feldmann & Schweitzer	Cretaceous		United States	A crab belonging to the group Raninoida. The type species is P. robreidi.
Prealpicarcinus laisensis[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Pilumnidae.
Prealpiplax[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the superfamily Goneplacoidea and the family Euryplacidae. The type species is P. lessinea.
Prolecythocaris[79]	Gen. et sp. nov	Valid	Schweigert & Robins	Late Jurassic (early Kimmeridgian)		Germany	A crab belonging to the family Lecythocaridae. The type species is P. hauckei.
Protomunida pentaspinosa[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A member of Munididae.
Pseudocorallomursia[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Calappidae. The type species is P. barbierii.
Pseudolambrus antiquus[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Parthenopidae.
Pustulina colossea[56]	Sp. nov	Valid	Devillez et al.	Early Cretaceous (Hauterivian)		France	A member of Erymidae.
Pustulina occitana[56]	Sp. nov	Valid	Devillez et al.	Early Cretaceous (Berriasian)		France	A member of Erymidae.
Rama[32]	Gen. et sp. nov	Junior homonym	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Cancridae. The type species is R. lineatuberculata. The generic name is preoccupied by Rama Bleeker; de Angeli & Ceccon (2017) coined a replacement name Ramacarcinus.[58]
Ramozius[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Pseudoziidae. The type species is R. punctatus.
Raniliformis clampensis[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Raninidae.
Rathbunopon viai[80]	Sp. nov	Valid	González-León et al.	Early Cretaceous (Albian)		Spain	A homolodromioid crab belonging to the family Prosopidae.
Rosadromites[33]	Gen. et sp. nov	Valid	Schweitzer et al.	Early Cretaceous (Albian)	Glen Rose Formation	United States	A member of Homolodromioidea belonging to the family Longodromitidae. The type species is Rosadromites texensis.
Rugosanilia[44]	Gen. et comb. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Raninidae. The type species is "Raniliformis" rugosa De Angeli & Beschin (2007).
Sakaila italica[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A species of Sakaila.
Santeexanthus caporiondoi[44]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the superfamily Pseudozioidea and the family Pseudoziidae.
Scutata[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Tetraliidae. The type species is S. eocenica.
Serraphylctaena[49]	Gen. et comb. nov	Valid	Robins et al.	Cretaceous (Albian/Cenomanian)		Spain	A member of Galatheoidea. The type species is "Paragalathea" multisquamata Vía Boada (1981).
Spathanomus[48]	Gen. et sp. nov	Valid	De Angeli	Eocene (Priabonian)		Italy	A crab belonging to the group Cyclodorippoida and the family Cymonomidae. The type species is Spathanomus felicianensis.
Speocarcinus latus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Xanthidae, a species of Speocarcinus.
Tanidromites longinosa[81]	Sp. nov	Valid	Starzyk	Late Jurassic (Oxfordian)		Poland	A homolodromioid crab belonging to the family Tanidromitidae.
Tanidromites muelleri[82]	Sp. nov	Valid	Krobicki & Zatoń	Middle Jurassic (late Bajocian)		Poland	A homolodromioid crab belonging to the family Tanidromitidae.
Tanidromites schweitzerae[81]	Sp. nov	Valid	Starzyk	Late Jurassic (Oxfordian)		Poland	A homolodromioid crab belonging to the family Tanidromitidae.
Tanidromites wysokaensis[81]	Sp. nov	Disputed	Starzyk	Late Jurassic (Oxfordian)		Poland	A homolodromioid crab belonging to the family Tanidromitidae. Klompmaker et al. (2020) considered this species to be a junior synonym of Tanidromites scheffnerae Schweigert & Koppka (2011).[83]
Tethysastacus[56]	Gen. et comb. nov	Valid	Devillez et al.	Early Cretaceous (Valanginian)		France	A member of Erymidae. The type species is T. tithonius.
Tethyscarpilius[84]	Gen. et sp. et comb. nov	Valid	De Angeli & Alberti	Eocene		Italy  United States	A crab belonging to the family Carpiliidae. Genus includes new species T. bericus, as well as "Palaeocarpilius" brodkorbi Lewis & Ross (1965).
Tetralia minuta[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Tetraliidae.
Texicancer[78]	Gen. et comb. nov	Valid	Franţescu, Feldmann & Schweitzer	Cretaceous		United States	A crab belonging to the group Raninoida. The type species is "Necrocarcinus" renfroae Stenzel (1945).
Thalassina tsuyamensis[85]	Sp. nov	Valid	Ando & Kishimoto in Ando, Kishimoto & Kawano	Miocene	Yoshino Formation	Japan	A species of Thalassina.
Thalassina yamato[85]	Sp. nov	Valid	Ando & Kishimoto in Ando, Kishimoto & Kawano	Miocene	Kawachi Formation	Japan	A species of Thalassina.
Titanocarcinus schweitzerae[54]	Sp. nov	Valid[55]	Robin et al.	Paleocene (Danian)		France	A crab belonging to the family Tumidocarcinidae.
Tonneleryon[86]	Gen. et sp. nov	Valid	Audo	Early Jurassic (Toarcian)	Posidonia Shale	Germany	A member of Polychelida. The type species is T. schweigerti.
Tropicalia[32]	Gen. et sp. nov	Junior homonym	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Domeciidae. The type species is T. parva. The generic name is preoccupied by Tropicalia Kocak & Kemal (2008); de Angeli & Ceccon (2017) coined a replacement name Proticalia.[58]
Tuberosagalathea[49]	Gen. et comb. et 2 sp. nov	Valid	Robins et al.	Late Jurassic and Early Cretaceous (Hauterivian)	Ernstbrunn Limestones Štramberk Limestones	Austria  Czech Republic  France  Poland  Romania	A member of Galatheoidea. The type species is "Galathea" neojurensis Patrulius (1959); genus also includes "Galatheites" neocomiensis Van Straelen (1936) and "Eomunidopsis" portlandica Fraaye & Collins (1996), as well as new species Tuberosagalathea tornatilis and Tuberosagalathea antefixa.
Tymolus italicus[32]	Sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the group Cyclodorippoida and the family Cyclodorippidae.
Tytthotanais[35]	Gen. et sp. nov	Valid	Sánchez-García et al.	Late Cretaceous (Cenomanian)	Pyrenean amber	France	A member of Tanaidacea of uncertain phylogenetic placement. The type species is T. tenvis.
Vasconilia[49]	Gen. et comb. et sp. nov	Valid	Robins et al.	Late Jurassic and Early Cretaceous	Ernstbrunn Limestones Štramberk Limestones	Austria  Spain  Japan?	A member of Galatheoidea. The type species is "Galathea" ruizi Van Straelen (1940); genus also includes "Galatheites" straeleni Ruiz de Gaona (1943), as well as new species Vasconilia xystos. Genus might also include ?Vasconilia miyakoensis (Takeda & Fujiyama, 1983).
Ypresicorystes[32]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Ypresian)		Italy	A crab belonging to the family Corystidae. The type species is Y. expansus.
Zygastrocarcinus tricki[87]	Sp. nov	Valid	Nyborg et al.	Late Cretaceous (Campanian)	Pierre Shale	United States	A crab belonging to the family Homolidae.

Ostracods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Actinocythereis imbeensis[88]	Sp. nov	Valid	De Mattos Manica & Coimbra	Early Miocene	Pelotas Basin	Brazil	A member of the family Trachyleberididae, a species of Actinocythereis.
Amphizona? argentinensis[89]	Sp. nov	Valid	Salas	Devonian (Lochkovian)	Talacasto Formation	Argentina	A member of Podocopa belonging to the Beyrichiocopida and the family Arcyzonidae.
Aparchitellina magnicornigera[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation	Russia	A member of Beyrichicopida belonging to the family Aparchitellinidae.
Aparchitellina triangulata[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Rusanov Formation	Russia	A member of Beyrichicopida belonging to the family Aparchitellinidae.
Asciocythere raybatei[91]	Nom. nov	Valid	Kempf	Middle Jurassic		United Kingdom	A member of Schulerideidae; a replacement name for Asciocythere acuminata Bate (1964).
Bairdia sudrensis[92]	Sp. nov	Valid	Ied & Ismail	Miocene		Egypt	A member of the family Bairdiidae.
Bingeria fluida[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Krasnaya Bukhta Formation Ust’-Spokoinaya Formation	Russia	A member of Beyrichicopida belonging to the family Beyrichiidae.
Bingeria lucida[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Srednii Formation	Russia	A member of Beyrichicopida belonging to the family Beyrichiidae.
Bollia talacastensis[89]	Sp. nov	Valid	Salas	Devonian (Lochkovian)	Talacasto Formation	Argentina	A member of Podocopa belonging to the Beyrichiocopida and the family Bollidae.
Brachycythere smithsoniana[93]	Sp. nov	Valid	Antonietto et al.	Early Cretaceous (Aptian-Albian)	Riachuelo Formation	Brazil	A member of Podocopida belonging to the group Cytherocopina and the family Trachyleberididae.
Buntonia bassiounii[94]	Sp. nov	Valid	Morsi, Hewaidy & Samir	Eocene		Egypt	A member of Trachyleberididae, a species of Buntonia.
Buntonia posteroacuta[94]	Sp. nov	Valid	Morsi, Hewaidy & Samir	Eocene		Egypt	A member of Trachyleberididae, a species of Buntonia.
Bythoceratina brunomilhaui[91]	Nom. nov	Valid	Kempf	Miocene	Waitemata Basin	New Zealand	A member of Bythocytheridae; a replacement name for Bythoceratina robusta Milhau (1993).
Bythocythere agostinae[95]	Sp. nov	Valid	Sciuto	Pleistocene		Italy
Candona csikiiformis[96]	Sp. nov	Valid	Teterina	Miocene		Russia
Conchoprimitia brasiliensis[97]	Sp. nov	Valid	Adôrno & Salas in Adôrno et al.	Late Ordovician	Vila Maria Formation	Brazil	A member of Podocopa belonging to the order Beyrichiocopida and the family Conchoprimitiidae.
Cryptophyllus fossularis[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Ust’-Spokoinaya Formation	Russia
Cryptophyllus magnei[98]	Sp. nov	Valid	Maillet & Casier in Maillet, Danelian & Casier	Devonian (Givetian)	Fromelennes Formation	Belgium	A possible member of Palaeocopida belonging to the family Rhabdostichidae, a species of Cryptophyllus.
Ctenoloculinella papillifera[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation Rusanov Formation	Russia	A member of Hollinocopida belonging to the family Ctenoloculinidae.
Cytherella abdelshafyii[92]	Sp. nov	Valid	Ied & Ismail	Miocene		Egypt	A member of the family Cytherellidae.
Cytherellina subelliptica[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation Rusanov Formation	Russia	A member of Metacopida belonging to the family Healdiidae.
Cytheretta africana[92]	Sp. nov	Valid	Ied & Ismail	Miocene		Egypt	A member of the family Cytherettidae.
Cytheretta sinaensis[92]	Sp. nov	Valid	Ied & Ismail	Miocene		Egypt	A member of the family Cytherettidae.
Cytheropteron morsii[91]	Nom. nov	Valid	Kempf	Eocene		Egypt	A member of Cytheruridae, a species of Cytheropteron. The original specific name was Cytheropteron bicostatum, Morsi, Hewaidy & Samir (2016),[94] which turned out to be preoccupied by Cytheropteron bicostatum Brand (1990).
Cytheropteron nukhulensis[94]	Sp. nov	Valid	Morsi, Hewaidy & Samir	Eocene		Egypt	A member of Cytheruridae, a species of Cytheropteron.
Cytheropteron speijeri[94]	Sp. nov	Valid	Morsi, Hewaidy & Samir	Eocene		Egypt	A member of Cytheruridae, a species of Cytheropteron.
Digmocythere centroreticulata[94]	Sp. nov	Valid	Morsi, Hewaidy & Samir	Eocene		Egypt	A member of Trachyleberididae, a species of Digmocythere.
Ectonodoconcha[99]	Gen. et sp. nov	Valid	Ceolin & Whatley in Ceolin et al.	Paleocene (Danian)	Neuquén Basin	Argentina	A member of Cytheridae. The type species is Ectonodoconcha lepidotus.
Eobeyrichia nordensis[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Srednii Formation	Russia	A member of Beyrichicopida belonging to the family Beyrichiidae.
Eobeyrichia subeurina[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Samoilovich Formation Srednii Formation	Russia	A member of Beyrichicopida belonging to the family Beyrichiidae.
Eokloedenia reticulata[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Late Silurian	Krasnaya Bukhta Formation	Russia	A member of Beyrichicopida belonging to the family Kloedeniidae.
Eukloedeneterra[90]	Gen. et sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Samoilovich Formation Srednii Formation	Russia	A member of Kloedenellocopida belonging to the family Kloedenellidae. The type species is E. borealis.
Evlanella peranensis[100]	Sp. nov	Valid	Maillet in Maillet et al.	Devonian (Givetian)	Candás Formation	Spain	A species of Evlanella.
Evlanella tuberculifera[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Early Devonian	Al’banov Formation	Russia	A member of Kloedenellocopida belonging to the family Lichviniidae.
Gabonorygma[93]	Gen. et comb. et sp. nov	Valid	Antonietto et al.	Late Jurassic (Kimmeridgian–early Tithonian) and Early Cretaceous (Aptian-Albian)	Madiéla Formation Riachuelo Formation	Brazil  Denmark  Gabon	A member of Podocopida belonging to the group Cytherocopina and the family Krithidae. The type species is "Dicrorygma" brotzeni Christensen (1965); genus also includes new species G. sergipana.
Kozlowskiella? nuda[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation	Russia	A member of Beyrichicopida belonging to the family Aparchitellinidae.
Kuznetsoviella[101][102]	Nom. nov	Valid	Doweld	Early Cretaceous (Barremian)		Azerbaijan	A replacement name for Aenigma Kuznetsova (1957) and Kuznetsovia Doweld (2016) (both preoccupied).
Limnocythere alexanderi[103]	Sp. nov	Valid	Palacios-Fest, Cusminsky & McGlue	Late Quaternary		Argentina	A species of Limnocythere.
Limnocythere foresteri[103]	Sp. nov	Valid	Palacios-Fest, Cusminsky & McGlue	Late Quaternary		Argentina	A species of Limnocythere.
Limnocythere lysandrosi[103]	Sp. nov	Valid	Palacios-Fest, Cusminsky & McGlue	Late Quaternary		Argentina	A species of Limnocythere.
Limnocythere ruipunctifinalis[103]	Sp. nov	Valid	Palacios-Fest, Cusminsky & McGlue	Late Quaternary		Argentina	A species of Limnocythere.
Microxestoleberis scillae[95]	Sp. nov	Valid	Sciuto	Pleistocene		Italy
Mongolocypris kohi[104]	Sp. nov	Valid	Choi & Huh	Early Cretaceous (Albian)	Jinju Formation	South Korea	A non-marine ostracod, a species of Mongolocypris.
Mosolovina conifera[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation	Russia	A member of Beyrichicopida belonging to the family Aparchitellinidae.
Mosolovina costulifera[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation	Russia	A member of Beyrichicopida belonging to the family Aparchitellinidae.
Neohornibrookella nepeani[105]	Sp. nov	Valid	Warne & Whatley	Neogene		Australia
Neurocythere multiforma[106]	Sp. nov	Valid	Pais et al.	Middle Jurassic (Callovian)		Portugal
Nodoconcha polytorosa[99]	Sp. nov	Valid	Ceolin & Whatley in Ceolin et al.	Paleocene (Danian)	Neuquén Basin	Argentina	A member of Cytheridae, a species of Nodoconcha.
Nodoconcha sanniosis[99]	Sp. nov	Valid	Ceolin & Whatley in Ceolin et al.	Paleocene (Danian)	Neuquén Basin	Argentina	A member of Cytheridae, a species of Nodoconcha.
Nodoconcha upsilon[99]	Sp. nov	Valid	Ceolin & Whatley in Ceolin et al.	Paleocene (Danian)	Neuquén Basin	Argentina	A member of Cytheridae, a species of Nodoconcha.
Norditerria[90]	Gen. et 2 sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Krasnaya Bukhta Formation Ust’-Spokoinaya Formation	Russia	A member of Beyrichicopida belonging to the family Hexophthalmoididae. The type species is N. torosa; genus also includes N. simplex.
Noviportia cornuta[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Samoilovich Formation Srednii Formation	Russia	A member of Beyrichicopida belonging to the family Zygobolbidae.
Nyhamnella elongata[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Srednii Formation	Russia	A member of Kloedenellocopida belonging to the family Gotlandellidae.
Occultocythereis alsheikhlyi[91]	Nom. nov	Valid	Kempf	Late Cretaceous (Maastrichtian)		Iraq	A member of Trachyleberididae; a replacement name for Occultocythereis elongata Al-Sheikhly (1982).
Oertliella petraensis[107]	Sp. nov	Valid	Al-Sheikhly	Paleocene (Danian)	Taqiye Formation	Jordan	A member of the family Trachyleberididae, a species of Oertliella.
Paenaequina recta[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Devonian (Emsian)	Al’banov Formation	Russia	A member of Leperditicopida belonging to the family Schrenckiidae.
Palaeocytheridea (Malzevia)? dorsocostata[106]	Sp. nov	Valid	Pais et al.	Middle Jurassic (Callovian)		Portugal
Paragrenocythere monilis[107]	Sp. nov	Valid	Al-Sheikhly	Late Cretaceous (Maastrichtian)	Shiranish Formation	Iraq	A member of the family Trachyleberididae, a species of Paragrenocythere.
Parakrithe tayibaensis[94]	Sp. nov	Valid	Morsi, Hewaidy & Samir	Eocene		Egypt	A member of Krithidae, a species of Parakrithe.
Peloriops levisulcata[107]	Sp. nov	Valid	Al-Sheikhly	Late Cretaceous (late Campanian?-Maastrichtian)	Hartha Formation Shiranish Formation	Iraq	A member of the family Trachyleberididae, a species of Peloriops.
Perissocytheridea colini[108]	Sp. nov	Valid	Nogueira & Ramos	Oligocene-Miocene	Pirabas Formation	Brazil	An ostracod, a species of Perissocytheridea.
Perissocytheridea largulateralis[108]	Sp. nov	Valid	Nogueira & Ramos	Oligocene-Miocene	Pirabas Formation	Brazil	An ostracod, a species of Perissocytheridea.
Perissocytheridea pirabensis[108]	Sp. nov	Valid	Nogueira & Ramos	Oligocene-Miocene	Pirabas Formation	Brazil	An ostracod, a species of Perissocytheridea.
Perissocytheridea punctoreticulata[108]	Sp. nov	Valid	Nogueira & Ramos	Oligocene-Miocene	Pirabas Formation	Brazil	An ostracod, a species of Perissocytheridea.
Petrisigmoopsis? rotundum[89]	Sp. nov	Valid	Salas	Devonian (Lochkovian)	Los Espejos Formation Talacasto Formation	Argentina	A member of Podocopa belonging to the Beyrichiocopida and the family Bollidae.
Pircawayra antiqua[89]	Sp. nov	Valid	Salas	Devonian (Lochkovian)	Talacasto Formation	Argentina	A member of Podocopa belonging to the Beyrichiocopida and the superfamily Kloedenelloidea.
Platybolbina calva[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Late Silurian	Krasnaya Bukhta Formation	Russia	A member of Hollinocopida belonging to the family Eurychilinidae.
Praebythoceratina deltalata[93]	Sp. nov	Valid	Antonietto et al.	Early Cretaceous (Aptian-Albian)	Riachuelo Formation	Brazil	A member of Podocopida belonging to the group Cytherocopina and the family Bythocytheridae.
Praebythoceratina parascrobiculata[106]	Sp. nov	Valid	Pais et al.	Middle Jurassic (Callovian)		Portugal
Praeschuleridea lisensis[106]	Sp. nov	Valid	Pais et al.	Middle Jurassic (Callovian)		Portugal
Protobrachycythere[109]	Gen. et 2 sp. nov	Valid	Karpuk	Early Cretaceous (Barremian-Aptian)		Crimean Peninsula	A member of the family Protocytheridae. The type species is P. taurica; genus also includes P. aptica.
Pteriella? dubia[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Srednii Formation	Russia	A member of Hollinocopida belonging to the suborder Primitiopsicopina.
Reticulina ninurta[107]	Sp. nov	Valid	Al-Sheikhly	Middle-late Eocene	Avanah Formation Jaddala Formation	Iraq	A member of the family Trachyleberididae, a species of Reticulina.
Reticulina syriaensis[107]	Sp. nov	Valid	Al-Sheikhly	Paleocene	Aaliji Formation	Syria	A member of the family Trachyleberididae, a species of Reticulina.
Saccarchites tumefactus[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Late Silurian	Ust’-Spokoinaya Formation	Russia	A member of Beyrichicopida belonging to the family Ochescaphidae.
Samarella centrigibbera[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Rusanov Formation	Russia	A member of Metacopida belonging to the family Rishonidae.
Sapucariella[110]	Gen. et comb. et 3 sp. nov	Valid	Puckett, Andreu & Colin	Early Cretaceous (Aptian)-early Paleocene		Agulhas Bank  Algeria  Brazil  Cameroon  Egypt  Gabon  Ghana  India  Iran  Israel  Ivory Coast  Kuwait  Lebanon  Libya  Madagascar  Mali  Morocco  Niger  Nigeria  Oman  Saudi Arabia  Senegal  South Africa  Tanzania  Togo  Tunisia  United Arab Emirates	A member of the family Brachycytheridae. The type species is "Brachycythere" sapucariensis Krömmelbein (1964); genus also includes "Brachycythere" agulhasensis Dingle (1971), S. angulata (Grékoff, 1951), "Brachycythere" arabica Al-Furaih (1985), "Brachycythere" armata Reyment (1960), "Brachycythere" beershevaensis Honigstein (1984), "Brachycythere" dumoni Bismuth & Saint-Marc in Bismuth et al. (1981), "Brachycythere" ekpo Reyment (1960), "Brachycythere" glypta Al-Furaih (1985), "Brachycythere" ilamensis Emami (1990), "Brachycythere" iranensis Emami (1990), "Brachycythere" jodhpurensis Singh (1997), "Brachycythere" kulatturensis Guha (1971), "Brachycythere" labioforma Emami (1990), "Cytheridea" longicaudata Chapman (1904), "Brachycythere" meata Al-Furaih (1985), "Brachycythere" multidifferentis Nicolaidis & Piovesan in Piovesan et al. (2013), "Brachycythere" oguni Reyment (1960), "Brachycythere" posterotruncata Emami (1990), "Brachycythere" pucketti Babinot et al. (2009), "Brachycythere" reymenti Emami (1990), "Brachycythere" rotunda Dingle (1969), "Brachycythere" shamlani Al-Furaih (1985), "Brachycythere" sicarius Dingle (1980), "Brachycythere" sillakkudiensis Sastry et al. (1972), "Brachycythere" subtriangulata Singh & Porwal (1989), "Brachycythere" tumida Al-Furaih (1985), "Brachycythere" undosa Al-Furaih (1985) and "Brachycythere" ventrocomplanatus Delicio et al. (2000), as well as new species S. honigsteini, S. parvoangulata and S. pseudosapucariensis.
Satiellina paranaensis[97]	Sp. nov	Valid	Adôrno & Salas in Adôrno et al.	Late Ordovician	Vila Maria Formation	Brazil	A member of Podocopa belonging to the order Beyrichiocopida and the superfamily Drepanelloidea.
Sibiritia ? septentrionalis[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Golomyannyi Formation Srednii Formation Vodopad Formation	Russia	A member of Leperditicopida belonging to the family Sibiritiidae.
Signetopsis bolbilobata[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Ust’-Spokoinaya Formation	Russia	A member of Hollinocopida belonging to the family Signetopsidae.
Suinella[89]	Gen. et sp. nov	Valid	Salas	Devonian (Lochkovian)	Talacasto Formation	Argentina	A member of Podocopa belonging to the Beyrichiocopida and possibly to the family Primitiopsidae. The type species is S. huarpesi.
Sulcella? acuminata[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation	Russia	A member of Kloedenellocopida belonging to the family Mennerellidae.
Sulcella? annae[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation	Russia	A member of Kloedenellocopida belonging to the family Mennerellidae.
Sulcella? modesta[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation	Russia	A member of Kloedenellocopida belonging to the family Mennerellidae.
Timanella magna[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation	Russia	A member of Platycopida belonging to the family Ellesclavidae.
Timanella rara[90]	Sp. nov	Valid	Evdokimova in Abushik & Evdokimova	Devonian	Al’banov Formation	Russia	A member of Platycopida belonging to the family Ellesclavidae.
Tollitia navicula[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Golomyannyi Formation Srednii Formation	Russia	A member of Leperditicopida belonging to the family Tollitiidae.
Tricostabrachycythere[110]	Gen. et comb. nov	Valid	Puckett, Andreu & Colin	Cretaceous (Albian-Cenomanian)	Goru Formation	India  Morocco  Venezuela	A member of the family Brachycytheridae. The type species is "Veenia" kharatarensis Singh (1997).
Tvaerenella? ambigua[90]	Sp. nov	Valid	Abushik in Abushik & Evdokimova	Silurian	Srednii Formation	Russia	A member of Hollinocopida belonging to the family Piretellidae.
Virgulacytheridea posteroacuminata[106]	Sp. nov	Valid	Pais et al.	Middle Jurassic (Callovian)		Portugal
Xestoleberis morsiana[91]	Nom. nov	Valid	Kempf	Eocene		Egypt	A member of Xestoleberididae, a species of Xestoleberis. The original specific name was Xestoleberis posterotruncata Morsi, Hewaidy & Samir (2016),[94] which turned to be preoccupied by Xestoleberis posterotruncata Titterton & Whatley (2005).

Other crustaceans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Apocyclops californicus[111]	Sp. nov	Valid	Hołyńska, Leggitt & Kotov	Miocene (latest Burdigalian or earliest Langhian)	Barstow Formation	United States	A cyclopid copepod, a species of Apocyclops.
Arcuatoscalpellum[112]	Gen. et comb. nov	Valid	Gale	Cretaceous (Aptian to Maastrichtian)		France  United Kingdom	A goose barnacle belonging to the family Scalpellidae. The type species is "Scalpellum" arcuatum Darwin (1851); genus also includes "Scalpellum" (Arcoscalpellum) comptum Withers (1910) and "Scalpellum" trilineatum Darwin (1851).
Catherinum anglicum [112]	Sp. nov	Valid	Gale	Late Cretaceous (Campanian)		United Kingdom	A goose barnacle belonging to the family Scalpellidae, a species of Catherinum.
Cetopirus fragilis[113]	Sp. nov	Valid	Collareta et al.	Early Pleistocene		Italy	A whale barnacle, a species of Cetopirus.
Conopea bacata[114]	Sp. nov	Valid	Carriol in Carriol & Schneider	Miocene (Burdigalian)		Germany	A barnacle belonging to the family Archaeobalanidae, a species of Conopea.
Gesvesia[115]	Gen. et sp. nov	Valid	Gueriau et al.	Devonian (Famennian)		Belgium	A clam shrimp belonging to the group Spinicaudata. The type species is Gesvesia pernegrei.
Haltinnaias[115]	Gen. et sp. nov	Valid	Gueriau et al.	Devonian (Famennian)		Belgium	A member of Anostraca. The type species is Haltinnaias serrata.
Myolepas reussi[116]	Sp. nov	Valid	Kočí et al.	Late Cretaceous (late Cenomanian to early Turonian)	Bohemian Cretaceous Basin	Czech Republic	A goose barnacle.
Nestoria sikeshuensis[117]	Sp. nov	Valid	Teng et al.	Early Cretaceous	Tugulu Group	China	A clam shrimp belonging to the superfamily Eosestherioidea and the family Nestoriidae.
Pachyscalpellum heltzeli[118]	Sp. nov	Valid	Jagt, Verhesen & Goolaerts	Late Cretaceous (Maastrichtian)	El Haria Formation	Tunisia	A goose barnacle belonging to the family Scalpellidae, a species of Pachyscalpellum.
Plesiobrachylepas[119]	Gen. et sp. nov	Valid	Carriol in Carriol et al.	Eocene	Mo Clay Formation	Denmark	A barnacle belonging to the group Sessilia and the family Brachylepadidae. The type species is Plesiobrachylepas jutlandica.
Pseudoscapholeberis[120]	Gen. et sp. nov	Valid	Flössner & Fryer	Cretaceous (late Albian or earliest Cenomanian)		France	A member of Anomopoda. The type species is Pseudoscapholeberis enigmatica.
Scillaelepas danningeri[114]	Sp. nov	Valid	Carriol in Carriol & Schneider	Miocene (Burdigalian)	Ottnang Formation	Austria  Germany	A barnacle belonging to the family Calanticidae, a species of Scillaelepas.
Stipilepas[119]	Gen. et sp. nov	Valid	Carriol in Carriol et al.	Eocene	Mo Clay Formation	Denmark	A goose barnacle belonging to the group Scalpelliformes and the family Eolepadidae. The type species is Stipilepas molerensis.

Insects

Main article: 2016 in insect paleontology

Trilobites

Research

• Queues of the trilobite Trimerocephalus chopini are described from the Devonian (Famennian) of Poland by Błażejowski et al. (2016), who interpret these queues as representing mass migratory chains.[121]
• Disarticulated exoskeletons of the phacopids belonging to the species Omegops cornelius preserved within a nautiloid conch are described from the Devonian Hongguleleng Formation (China) by Zong, Fan & Gong (2016), who interpret the fossils as indicating that the trilobites had moulted within the living chamber of a dead nautiloid.[122]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Abarrandia[123]	Gen. et sp. nov	Valid	Zhou, Zhou & Yin	Ordovician (Darriwilian)	Zhuozishan Formation	China	A nileid. Genus includes new species A. wuhaiensis.
Acanthomicmacca franconica[124]	Sp. nov	Valid	Geyer	Cambrian		Germany	A member of Chengkouiidae.
Acanthomicmacca hupei[124]	Sp. nov	Valid	Geyer	Middle Cambrian	Jbel Wawrmast Formation	Morocco	A member of Chengkouiidae.
Acanthomicmacca mirandoides[124]	Sp. nov	Valid	Geyer	Cambrian			A member of Chengkouiidae.
Acanthomicmacca morrisoni[124]	Sp. nov	Valid	Geyer	Middle Cambrian	Jbel Wawrmast Formation	Morocco	A member of Chengkouiidae.
Acanthomicmacca rana[124]	Sp. nov	Valid	Geyer	Cambrian			A member of Chengkouiidae.
Acastoides poschmanni[125]	Sp. nov	Valid	Basse & Müller	Devonian (Emsian)	Rupbach Shale	Germany
Agasella prona[126]	Sp. nov	Valid	Salikhova in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the family Agraulidae.
Amphitryon xiushanensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Ampyx gongwusuensis[128]	Sp. nov	Valid	Lee et al.	Ordovician (Darriwilian)	Klimoli Formation	China
Ampyxinella latilimbata[123]	Sp. nov	Valid	Zhou, Zhou & Yin	Ordovician (Darriwilian)	Zhuozishan Formation	China
Amslamia[124]	Gen. et sp. nov	Valid	Geyer	Cambrian		Morocco	A member of Chengkouiidae. Genus includes new species A. rudkini.
Anaxarete[124]	Gen. et comb. nov	Junior homonym	Geyer	Cambrian			A member of Protoleninae; a new genus for "Micmacca" ellipsocephaloides var. senior (recombined as Anaxarete senior). The generic name is preoccupied by Anaxarete Gistl (1848).
Anechocephalus intermedius[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Anechocephalus rebeccaae[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Angustinouyia[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Inouyiidae.
Apatokephalus rugosus[131]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Asaphellus clarksoni[131]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Aulacopleura (Paraaulacopleura) lemkei[125]	Sp. nov	Valid	Basse & Müller	Devonian (Eifelian)	Rupbach Shale	Germany
Austerops legrandi[132]	Sp. nov	Valid	Khaldi et al.	Early Devonian	‘Chefar el Ahmar’ Formation	Algeria	A phacopid, a species of Austerops.
Australoscutellum talenti[133]	Sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia
Barrandeops chattertoni[132]	Sp. nov	Valid	Khaldi et al.	Early Devonian	‘Chefar el Ahmar’ Formation	Algeria	A phacopid. Originally described as a species of Barrandeops; Van Viersen & Holland (2016) considered the genus Barrandeops to be a junior synonym of the genus Morocops[134] and subsequently Van Viersen, Holland & Koppka (2017) transferred the species B. chattertoni to the genus Morocops.[135]
Barrandeops (Lahnops)[125]	Subgen. et sp. et nom. nov	Valid	Basse & Müller	Devonian	Leun Shale Rupbach Shale	Germany	The subgenus includes new species Barrandeops (Lahnops) steinmeyeri, as well as Barrandeops (Lahnops) burhennei (a replacement name for Phacops holzapfeli Burhenne).
Binodaspis nuda[126]	Sp. nov	Valid	Pegel in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the family Ptychopariidae.
Buitella concava[126]	Sp. nov	Valid	Pegel & Salikhova in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the subfamily Conokephalininae.
Camarella[130]	Gen. nov	Junior homonym	Zhu in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Solenopleuridae. The generic name is preoccupied by Camarella Dall (1877).
Carioides[136]	Gen. et sp. nov	Valid	Bushuev & Makarova	Cambrian	Chaya Formation	Russia	A member of Anomocaridae. Genus includes new species C. enodis.
Cernuolimbus ludvigseni[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Chlupacops[137]	Gen. et sp. et comb. nov	Valid	Feist in Feist, Mahboubi & Girard	Late Devonian		Algeria  Germany  Morocco	A member of Phacopidae. The type species is C. laticeps; genus also includes "Phacops" cryphoides Richter & Richter (1926) and possibly "Phacops" reichi Kegel (1931), "Phacops" koeneni Holzapfel (1895) and "Nephranops" spectabilis Meischner (1965).
Claycreekia[129]	Gen. et sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada	A member of the family Olenidae. Genus includes new species C. chrisorum.
Cliffia aitkeni[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Corrugatagnostus ningqiangensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Cybantyx? ergodes[133]	Sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia
Cyclopyge latilimbata[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Cyphaspides (Cyphaspides) malbertii[125]	Sp. nov	Valid	Basse & Müller	Devonian (Eifelian)	Rupbach Shale	Germany
Cyphaspides (Cyphaspides) weugi[125]	Sp. nov	Valid	Basse & Müller	Devonian (Eifelian)	Wissenbach Shale	Germany
Cyphaspis bluhmi[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Eifelian)	Taboumakhloûf Formation?	Morocco
Cyphaspis boninoi[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Eifelian)	Hanonet Formation	Belgium
Cyphaspis eximia[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Eifelian)		Morocco
Cyphaspis foumzguidensis[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Eifelian)	Timrhanrhart Formation	Morocco
Cyphaspis heisingi[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Pragian)	Assa Formation	Morocco
Cyphaspis ihmadii[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Givetian)	Bou Dîb Formation	Morocco
Cyphaspis juergenhollandi[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Emsian)	Amerboh Group	Morocco
Cyphaspis khraidensis[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Emsian)	Timrhanrhart Formation	Morocco
Cyphaspis kippingi[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Emsian)	Tazoulait Formation	Morocco
Cyphaspis kweberi[125]	Sp. nov	Valid	Basse & Müller	Devonian (Emsian)	Rupbach Shale	Germany
Cyphaspis lerougei[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Eifelian)	Khebchia Formation	Morocco
Cyphaspis maharchensis[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Pragian)	Ihandar Formation	Morocco
Cyphaspis smeenki[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Eifelian)	El Otfal Formation	Morocco
Cyphaspis tadachachtensis[134]	Sp. nov	Valid	Van Viersen & Holland	Devonian (Emsian)	Khebchia Formation	Morocco
Dechenella burmeisteri emstae[138]	Subsp. nov	Valid	Basse, Koch & Lemke	Devonian (Givetian)		Germany	A member of the family Proetidae.
Degamella interrupta[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Degamella latifrons[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Destombesina schumacherorum[125]	Sp. nov	Valid	Basse & Müller	Devonian (Emsian)	Rupbach Shale	Germany
Diademaproetus habenichti[125]	Sp. nov	Valid	Basse & Müller	Devonian (Emsian)	Rupbach Shale	Germany
Dislobosaspis angustiglossoides[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Dolabrapex[133]	Gen. et sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia	A member of Scutelluidae. Genus includes new species D. acomus.
Dorypyge? aliena[126]	Sp. nov	Valid	Pegel in Pegel et al.	Cambrian		Russia	A member of Dorypygidae.
Dunderbergia? edgecombei[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Eccaparadoxides? thorslundi[139]	Sp. nov	Valid	Rushton, Weidner & Ebbestad	Cambrian		Sweden	A member of Paradoxididae of uncertain phylogenetic placement, possibly a species of Eccaparadoxides.
Echidnops taphomimus[140]	Sp. nov	Valid	Rustán & Balseiro	Early Devonian	Talacasto Formation	Argentina
Effnaspis clavata[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Eiluroides[130]	Gen. nov	Valid	Yuan & Zhu in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Menocephalidae.
Encrinurella latifrons[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Enigmapyge[137]	Gen. et sp. nov	Valid	Feist in Feist, Mahboubi & Girard	Devonian (early Famennian)	“Argiles de Marhouma” Formation	Algeria	A member of Phacopidae. The type species is E. marhoumensis.
Erytheia[124]	Gen. et comb. nov	Valid	Geyer	Cambrian		Poland	A member of Chengkouiidae. Genus includes "Micmacca" klimontowi Orłowski (1985).
Eugonocare? phillipi[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Forchhammeria insueta[126]	Sp. nov	Valid	Salikhova in Pegel et al.	Cambrian		Russia	A member of Asaphida belonging to the family Anomocaridae.
Gaindella[126]	Gen. et sp. nov	Valid	Pegel in Pegel et al.	Cambrian		Russia	A member of Asaphida belonging to the family Pterocephaliidae. The type species is G. angusta.
Galbagnostus obscurus[123]	Sp. nov	Valid	Zhou, Zhou & Yin	Ordovician (Darriwilian)	Zhuozishan Formation	China
Geesops fabrei[132]	Sp. nov	Valid	Khaldi et al.	Early Devonian	‘Chefar el Ahmar’ Formation	Algeria	A phacopid. Originally described as a species of Geesops; Van Viersen, Holland & Koppka (2017) transferred the species to the genus Adrisiops.[135]
Gymnagnostus kobayashii[131]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Hadromeros hunanensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Helanshanaspis[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Ptychopariidae.
Hexacopyge scopulata[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Holdenia liangshanensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Hydrocephalus spinulosus[139]	Sp. nov	Valid	Rushton, Weidner & Ebbestad	Cambrian		Sweden	A member of Paradoxididae.
Iguidiella[124]	Gen. et sp. nov	Valid	Geyer	Cambrian			A member of Chengkouiidae. Genus includes new species I. schaeri.
Illaenoscutellum psephos[133]	Sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia
Illaenus slancyensis[141]	Sp. nov	Valid	Krylov	Ordovician	Kukruse Formation	Russia
Inouyia (Sulcinouyia)[130]	Subgen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Inouyiidae.
Iotoryx[133]	Gen. et sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia	A member of Scutelluidae. Genus includes new species I. clarksoni.
Irbuklina[126]	Gen. et 2 sp. nov	Valid	Egorova in Pegel et al.	Cambrian		Russia	A member of Asaphida of uncertain phylogenetic placement. The type species is I. candida; genus also includes I. rigida.
Irvingella dawnae[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Irvingella lata[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Irvingella orrensis[142]	Sp. nov	Valid	Westrop & Adrain	Cambrian	Orr Formation	United States
Japonoscutellum drakton[133]	Sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia
Japonoscutellum fractum[133]	Sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia
Japonoscutellum mawsonae[133]	Sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia
Juliaspis convexa[126]	Sp. nov	Valid	Pegel in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the family Namanoiidae.
Kanlingia mirabilis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Kanlingia reshiensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Kanlingia rhombica[123]	Sp. nov	Valid	Zhou, Zhou & Yin	Ordovician (Darriwilian)	Zhuozishan Formation	China
Kendallina greenensis[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Kindbladia wilsoni[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Kodymaspis expansa[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Kodymaspis maocaopuensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Koneprusia martini[125]	Sp. nov	Valid	Basse & Müller	Devonian (Emsian)	Rupbach Shale	Germany
Kosovopeltis avita[133]	Sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia
Lamproscutellum scoparium[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Leiolonchodomas[127]	Gen. et sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China	Genus includes new species L. dangmengouensis.
Lenacare sentum[126]	Sp. nov	Valid	Salikhova in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the family Alokistocaridae.
Linguagnostus aldanicus[143]	Sp. nov	Valid	Makarova & Bushuev	Cambrian	Chaya Formation	Russia	A member of Diplagnostidae.
Lioaspis[126]	Gen. et sp. nov	Valid	Lazarenko & Salikhova in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the family Asaphiscidae. The type species is L. infida.
Liopeishaniella[144]	Gen. et comb. nov	Valid	Bentley, Jago & Cooper	Cambrian (Drumian)		Antarctica (Victoria Land)	A polymerid trilobite. Genus includes "Liopeishania" angusta Wolfart (1994).
Liopelta paula[126]	Sp. nov	Valid	Salikhova in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the family Anomocarellidae.
Liquanella[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Asaphiscidae.
Lisogorites yaoi[123]	Sp. nov	Valid	Zhou, Zhou & Yin	Ordovician (Darriwilian)	Zhuozishan Formation	China	An asaphid.
Lonchocheirurus[127]	Gen. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Lonchodomas yichongiaoensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Metayuepingia riccardii[131]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Microparia (Heterocyclopyge) obsoleta[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Microparia (Heterocyclopyge) taoyuanensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Mioptychopyge semicircula[123]	Sp. nov	Valid	Zhou, Zhou & Yin	Ordovician (Darriwilian)	Zhuozishan Formation	China	An asaphid.
Nileus cingolanii[131]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Nileus erici[131]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Niuchangella angustilimbata[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Niuchangella yuanbaensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Niuxinshania[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Crepicephalidae.
Oedicybele caeca[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Ogygiocaris? iruyensis[131]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Olenaspella chrisnewi[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Olenoides sassikaspa[144]	Sp. nov	Valid	Bentley, Jago & Cooper	Cambrian (Drumian)		Antarctica (Victoria Land)
Onychopyge acenolazai[131]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Onychopyge gonzalezae[131]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina
Opsypharus pandanensis[133]	Sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia
Orientanomocare[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Anomocarellidae.
Orygmaspis (Parabolinoides) doriansmithi[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Orygmaspis (Parabolinoides) jenkinsi[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Orygmaspis (Parabolinoides) mckellari[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Orygmaspis (Parabolinoides) medispina[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Orygmaspis (Parabolinoides) morrisi[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Otarion fugitivum[145]	Sp. nov	Valid	McNamara & Feist	Devonian (Frasnian)	Virgin Hills Formation	Australia	A proetid, a species of Otarion.
Ovalocephalus pyriformis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Ovalocephalus shaanxiensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Palaeosunaspis[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Sunaspidae.
Panderia acutata[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Panderia lubrica[123]	Sp. nov	Valid	Zhou, Zhou & Yin	Ordovician (Darriwilian)	Zhuozishan Formation	China
Parabolinella clarisae[146]	Sp. nov	Valid	Monti, del Huerto Benítez & Ramirez	Ordovician (Tremadocian)		Argentina	A member of Olenidae.
Parabolinella pompadouris[146]	Sp. nov	Valid	Monti, del Huerto Benítez & Ramirez	Ordovician (Tremadocian)		Argentina	A member of Olenidae.
Paraperaspis striolatus[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Paraphillipsinella ciliensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Pararemopleurides[127]	Gen. et 3 sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China	Genus includes new species P. longispinus, P. schizophyllus and P. shaanxiensis.
Parashanxiella[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Proasaphiscidae.
Paraszeaspis[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Ignotogregatidae.
Parillaenus xui[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Pelitlina? corbachoi[125]	Sp. nov	Valid	Basse & Müller	Early Devonian		Spain
Perizostra[133]	Gen. et sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia	A member of Scutelluidae. Genus includes new species P. campbelli.
Phacops boudjemaai[132]	Sp. nov	Valid	Khaldi et al.	Early Devonian	‘Chefar el Ahmar’ Formation	Algeria	A phacopid. Originally described as a species of Phacops; Van Viersen, Holland & Koppka (2017) transferred the species to the genus Adrisiops.[135]
Platypeltoides hammondi[147]	Sp. nov	Valid	Corbacho & López-Soriano	Ordovician (late Tremadocian)	Fezouata formation	Morocco	A nileid.
Poulsenia venusta[126]	Sp. nov	Valid	Pegel in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the family Ptychopariidae.
Prolisania[130]	Gen. nov	Valid	Yuan & Zhu in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Lisaniidae.
Pseudopetigurus yuandunensis[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Psychopyge angeles[125]	Sp. nov	Valid	Basse & Müller	Devonian (Emsian)		Morocco
Qianlishania[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Solenopleuridae.
Quintonia[133]	Gen. et 2 sp. nov	Valid	Holloway & Lane	Silurian	Tomcat Creek limestone	Australia	A member of Scutelluidae. Genus includes new species Q. arata and Q. pavo.
Rasettiella[148]	Gen. et comb. et 2 sp. nov	Junior homonym	Wernette & Westrop	Cambrian		Canada	A dolichometopid. Genus includes species previously assigned to the genus Bathyuriscidella, as well as Rasettiella lupiae and Rasettiella sorgehanarum. The generic name is preoccupied by Rasettiella Özdikmen (2005).
Rhenocynproetus vanvierseni[125]	Sp. nov	Valid	Basse & Müller	Devonian (Emsian)	Rupbach Shale	Germany
Rickardsia[149]	Gen. et comb. nov	Valid	Storey, Thomas & Owens	Silurian		Czech Republic  Sweden  United Kingdom	A member of Dalmanitidae. The type species is "Dalmanites" mobergi Hede (1915); genus also includes "Delops" dermolac Šnajdr (1981) and "Phacops (Dalmania)" nobilis Thomas (1900).
Rina? mayskaya[136]	Sp. nov	Valid	Bushuev & Makarova	Cambrian	Chaya Formation	Russia	A member of Solenopleuridae.
Rorringtonia conica[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Rorringtonia minuta[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Rorringtonia obsoleta[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Rotundampyx[123]	Gen. et sp. nov	Valid	Zhou, Zhou & Yin	Ordovician (Darriwilian)	Zhuozishan Formation	China	A raphiophorid. Genus includes new species R. lasengmiaoensis.
Ryaneyreus[129]	Gen. et sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada	A member of the family Parabolinoididae. Genus includes news species R. zoeae.
Sagavia obscura[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Scabriscutellum (Rheiscutellum)[125]	Subgen. nov	Valid	Basse & Müller	Devonian (Emsian)	Rupbach Shale	Germany
Sciaspis[130]	Gen. nov	Valid	Zhu in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Ordosiidae.
Sibirolenus[124]	Gen. et comb. nov	Valid	Geyer	Cambrian		Russia	A member of Palaeolenidae; a new genus for "Micmacca" enormis.
Sinoanomocare[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Anomocaridae.
Strictaspis[126]	Gen. et sp. nov	Valid	Lazarenko & Salikhova in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the family Asaphiscidae. The type species is S. leve.
Sulcipagetia[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Eodiscidae.
Taenicephalus leei[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Taenicephalus westropi[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Tanglefootia[129]	Gen. et sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada	A member of the family Cheilocephalidae. Genus includes new species T. calvini.
Taosigouia[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Anomocarellidae.
Telephina truncata[127]	Sp. nov	Valid	Zhou, Zhou & Xiang	Ordovician	Pagoda Formation	China
Telopeltis intermedia[145]	Sp. nov	Valid	McNamara & Feist	Devonian (Frasnian)	Virgin Hills Formation	Australia	A species of Telopeltis.
Tomagnostus apertus[143]	Sp. nov	Valid	Makarova & Bushuev	Cambrian	Chaya Formation	Russia	A member of Ptychagnostidae.
Tongxinaspis ovata[123]	Sp. nov	Valid	Zhou, Zhou & Yin	Ordovician (Darriwilian)	Zhuozishan Formation	China	A dionidid.
Torleyiscutellum[150]	Gen. et sp. nov	Valid	Basse, Koch & Lemke	Devonian (early Givetian)		Germany	A scutelluid trilobite. Genus includes new species T. herwigorum.
Trifoliops cronierae[137]	Sp. nov	Valid	Feist in Feist, Mahboubi & Girard	Late Devonian	“Argiles de Marhouma” Formation	Algeria	A member of Phacopidae.
Trimerocephalus mahboubii[137]	Sp. nov	Valid	Feist in Feist, Mahboubi & Girard	Late Devonian	“Argiles de Marhouma” Formation	Algeria	A member of Phacopidae.
Tropidocoryphe werneri[125]	Sp. nov	Valid	Basse & Müller	Devonian (Emsian)	Rupbach Shale	Germany
Ulakhanella triangula[126]	Sp. nov	Valid	Pegel in Pegel et al.	Cambrian		Russia	A member of Ptychopariida belonging to the family Palaeolenidae.
Verkholenoides cognatus[126]	Sp. nov	Valid	Pegel & Salikhova in Pegel et al.	Cambrian		Russia	A member of Asaphida belonging to the family Pterocephaliidae.
Vicnepea[144]	Gen. nov	Valid	Bentley, Jago & Cooper	Cambrian (Drumian)		Antarctica (Victoria Land)	A polymerid trilobite.
Wuhaiaspis[130]	Gen. nov	Valid	Yuan & Zhang in Yuan, Zhang & Zhu	Cambrian		China	A member of the family Ptychopariidae.
Wujiajiania lyndasmithae[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada
Wujiajiania ricksmithi[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada

Other arthropods

Research

• A description of the ventral nerve cord in the early Cambrian euarthropod Chengjiangocaris kunmingensis is published by Yang et al. (2016).[151]
• A three-dimensionally preserved larva of Leanchoilia illecebrosa is described by Liu et al. (2016).[152]
• A study of the eye anatomy of the thylacocephalan (an arthropod of uncertain phylogenetic placement, possibly crustacean) Dollocaris is published by Vannier et al. (2016).[153]
• A study on the ability of bradoriid arthropods to close the shields of their carapace is published by Betts, Brock & Paterson (2016).[154]
• A study on the morphology of the digestive system and taphonomy of a variety of specimens of the Cambrian arthropod Campanamuta mantonae, known from the Sirius Passet Lagerstätte (Greenland), is published by Strang, Armstrong & Harper (2016).[155]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acanthomeridion anacanthus[156]	Sp. nov	Valid	Hou et al.	Early Cambrian		China
Agnostotes weugi[129]	Sp. nov	Valid	Chatterton & Gibb	Cambrian (Furongian)		Canada	A member of Agnostida.
Aquilonifer[157][158][159]	Gen. et sp. nov	Valid	Briggs et al.	Silurian (late Wenlock)	Herefordshire Lagerstätte	United Kingdom	A stem-mandibulate. The type species is Aquilonifer spinosus.
Archeallacma[160]	Gen. et sp. nov	Valid	Sánchez-García & Engel[161]	Early Cretaceous		Spain	A springtail belonging to the family Sminthuridae. The type species is A. dolichopoda.
Brachyaglaspis[162]	Gen. et sp. nov	Valid	Ortega-Hernández, Van Roy & Lerosey-Aubril	Ordovician (middle Floian)	Fezouata Konservat-Lagerstätte	Morocco	A member of Aglaspidida. The type species is Brachyaglaspis singularis.
Burmisotoma spinulifera[163]	Sp. nov	Valid	Sánchez-García & Engel	Early Cretaceous (late Albian)	Utrillas Group	Spain	A springtail belonging to the family Isotomidae.
Clypecaris serrata[164]	Sp. nov	Valid	Yang et al.	Cambrian	Hongjingshao Formation	China	A member of Deuteropoda of uncertain phylogenetic placement.
Cretokatianna[160]	Gen. et sp. nov	Valid	Sánchez-García & Engel[161]	Early Cretaceous		Spain	A springtail belonging to the family Katiannidae. The type species is C. bucculenta.
Decoracaris[50]	Gen. et sp. nov	Valid	Briggs et al.	Ordovician (Darriwilian)	Winneshiek Lagerstätte	United States	A bivalved arthropod of uncertain phylogenetic placement, possibly a thylacocephalan. The type species is Decoracaris hildebrandi.
Dyrnwynia[165]	Gen. et sp. nov	Valid	Legg	Ordovician (middle Darriwilian)	Llanfallteg Formation	United Kingdom	A member of Marrellida. The type species is Dyrnwynia conollyi.
Enosiaspis[166]	Gen. et sp. nov	Valid	Legg	Ordovician (Tremadocian)	Fezouata Formation	Morocco	A marrellomorph belonging to the group Acercostraca. The type species is Enosiaspis hrungnir.
Hipponicharion pearylandica[167]	Sp. nov	Valid	Peel et al.	Cambrian	Henson Gletscher Formation	Greenland	A member of Bradoriida. Originally described as a species of Hipponicharion; subsequently made the type species of a separate genus Navarana.[168]
Iosuperstes[50]	Gen. et sp. nov	Valid	Briggs et al.	Ordovician (Darriwilian)	Winneshiek Lagerstätte	United States	A bivalved arthropod of uncertain phylogenetic placement. The type species is Iosuperstes collisionis.
Katiannasminthurus[160]	Gen. et sp. nov	Valid	Sánchez-García & Engel[161]	Early Cretaceous		Spain	A springtail, possibly a member of the family Sminthuridae. The type species is K. xenopygus.
Liangshanella? nivalis[167]	Sp. nov	Valid	Peel et al.	Cambrian	Henson Gletscher Formation	Greenland	A member of Bradoriida.
Paleolimulus woodae[169]	Sp. nov	Valid	Lerner, Lucas & Mansky	Carboniferous (Tournaisian)	Horton Bluff Formation	Canada
Proisotoma communis[163]	Sp. nov	Valid	Sánchez-García & Engel	Early Cretaceous (late Albian)	Utrillas Group	Spain	A springtail belonging to the family Isotomidae.
Protoisotoma autrigoniensis[163]	Sp. nov	Valid	Sánchez-García & Engel	Early Cretaceous (late Albian)	Utrillas Group	Spain	A springtail belonging to the family Isotomidae.
Pseudosminthurides[160]	Gen. et sp. nov	Valid	Sánchez-García & Engel[161]	Early Cretaceous		Spain	A springtail belonging to the family Sminthurididae. The type species is P. stoechus.
Skryjagnostus usitatus[126]	Sp. nov	Valid	Salikhova in Pegel et al.	Cambrian		Russia	A member of Agnostida (a group of arthropods of uncertain phylogenetic placement, possibly trilobites).
Sphyrotheciscus[160]	Gen. et sp. nov	Valid	Sánchez-García & Engel[161]	Early Cretaceous		Spain	A springtail belonging to the family Sminthuridae. The type species is S. senectus.
Tomagnostus brantevikensis[170]	Sp. nov	Valid	Weidner & Nielsen	Middle Cambrian	Alum Shale Formation	Sweden	A member of Agnostida belonging to the family Ptychagnostidae.
Wisangocaris[171]	Gen. et sp. nov	Valid	Jago, García-Bellido & Gehling	Early Cambrian	Emu Bay Shale	Australia	A member of Chelicerata sharing many overall body features with Sanctacaris. The type species is Wisangocaris barbarahardyae.

References

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