Olenus (trilobite)

Olenus; Temporal range: 498.5–488.3 Ma PreꞒ Ꞓ O S D C P T J K Pg N Upper Cambrian
	Olenus micrurus from Trawsfynned, N-Wales, right free cheek displaced to left and turned 150°, pygidium missing
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Trilobita
Order:	Ptychopariida
Family:	Olenidae
Genus:	Olenus; Dalman, 1827[1] non Dejean, 1835 nomen nudum, ex Thomson, 1857 (= Trycherus, a beetle)[2]
species
	O. gibbosus (Wahlenberg, 1821) (type) synonym Entomostracites gibbosus; O. alpha Henningsmoen, 1957; O. altaicus Ivshin, 1962; O. apoxysomatus Jell et al., 1991 ; O. asiaticus Kobayashi, 1944; O. attenuatus (Boeck, 1838); O. austriacus Yang in Zhou et al., 1977; O. cataractes Salter, 1864; O. delicatus Öpik, 1963; O. dentatus Westergård, 1922; O. guizhouensis Lu and Chien in Yin and Li, 1978; O. haimantensis Reed, 1910 synonym Hundwarella haimantensis[3]; O. henningsmoeni Ahlgren and Ahlberg, 1996; O. micrurus Salter, 1849 ; O. mundus Lake, 1908; O. ogilviei Öpik, 1963; O. proximus Lazarenko, 1966; O. rotundatus Westergård, 1922; O. scanicus Westergård, 1922; O. sinensis Lu, 1964; O. solitarius (Westergård, 1922); O. transverses Westergård, 1922; O. truncatus (Brünnich, 1781); O. veles Rushton, 1983; O. wahlenbergi Westergård, 1922;

Olenus is a genus of Upper Cambrian ptychopariid trilobite.[4]

Distribution

Olenus can be found in the British Isles, India, Norway, Sweden, Denmark, Newfoundland, Texas, South Korea, and Australia.

O. haimantensis occurs in the Middle Cambrian of India (upper part of the Parahio Formation, left bank of the Parahio/Barachu river, above the Moopa camping ground, Spiti).[3]

Ecology

Fossils of Olenus are commonly found in dark mudstones, which were deposited on the seafloor in environments with low oxygen levels. The very wide side lobes (or pleurae) are thought to have shielded extended gills, which would have helped the animal absorb the maximum amount of oxygen possible in such an environment. Evidence also suggests that Olenus and its relatives may have developed a symbiotic relationship with sulfate-reducing bacteria, either by feeding on them or by absorbing nutrients directly from them.

Taxonomy

Species previously assigned to Olenus

O. forficula = Ceratopyge forficula
O. pecten = Ctenopyge pecten
O. punctatus = Asteropyge punctata
O. thompsoni = Olenellus thompsoni
O. zoppii = Dolerolenus zoppii

Species that may be confused with Olenus

The bug Anchodelphax olenus
The extant beetle Olenecamptus olenus.
The fossil waterbeetle Limnoxenus olenus.
The ant Polyrhachis olenus.
The ladyslipper orchid Paphiopedilum "Olenus".

Description

Size: Up to 1.5 in (4 cm) long. It had up to 15 thoracic segments, with a narrow axis and wide pleurae.

gollark: ++exec```haskellimport Data.Monoidimport Control.Applicativeimport Data.Listimport Control.Monadit = join.liftA2(<>)inits tailsallCombs xs = [1..] >>= \n -> mapM (const xs) [1..n]main = putStr . concat . take 10 . allCombs $ "gollark"```

gollark: Yesm

gollark: The params around the function are optional.

gollark: No they don't.

gollark: ++exec```haskellimport Data.Monoidimport Control.Applicativeimport Data.Listimport Control.Monadit = join.liftA2(<>)inits tailsmain = putStr $ it "gollark"```

References

Hwang, H.-A.; Choi, D.K. (2005). "Heterochrony of the Late Cambrian olenid trilobites from the Machari Formation, Yeongwol, Korea: implications for biostratigraphy and intercontinental correlation". Geosciences Journal. 9 (2): 215–222. doi:10.1007/bf02910580.
Poulsen, C. (1956). "Proposed use of the Plenary Powers to secure the availability of the generic names Olenus Dalman [1827], and Paradoxides Brogniard [1822] (Class Trilobita) for use in the sense in which these names are customarily employed". Bulletin of Zoological Nomenclature. 12 (1). Retrieved 2014-01-11.
Shah, S.K.; Parcha, S.K.; Raina, A.K. (1991). "Late Cambrian trilobites from Himalaya" (PDF). Journal of the Palaeontological Society of India. 36: 89–107.
Bodil Wesenberg Lauridsen & Arne Thorshøj Nielsen (2005). "The Upper Cambrian trilobite Olenus at Andarum, Sweden: a case of iterative evolution?". Palaeontology. 48 (5): 1041–1056. doi:10.1111/j.1475-4983.2005.00499.x.

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[1] Hwang, H.-A.; Choi, D.K. (2005). "Heterochrony of the Late Cambrian olenid trilobites from the Machari Formation, Yeongwol, Korea: implications for biostratigraphy and intercontinental correlation". Geosciences Journal. 9 (2): 215–222. doi:10.1007/bf02910580.

[2] Poulsen, C. (1956). "Proposed use of the Plenary Powers to secure the availability of the generic names Olenus Dalman [1827], and Paradoxides Brogniard [1822] (Class Trilobita) for use in the sense in which these names are customarily employed". Bulletin of Zoological Nomenclature. 12 (1). Retrieved 2014-01-11.

[SPR-3] Shah, S.K.; Parcha, S.K.; Raina, A.K. (1991). "Late Cambrian trilobites from Himalaya" (PDF). Journal of the Palaeontological Society of India. 36: 89–107.

[4] Bodil Wesenberg Lauridsen & Arne Thorshøj Nielsen (2005). "The Upper Cambrian trilobite Olenus at Andarum, Sweden: a case of iterative evolution?". Palaeontology. 48 (5): 1041–1056. doi:10.1111/j.1475-4983.2005.00499.x.