Paleobiota of the Ciechocinek Formation

The Ciechocinek Formation, is a Jurassic (lower to middle Toarcian) geologic formation that extends from Grimmen, Germany, to Poland.[1] Dinosaur remains are among the fossils that have been recovered from the formation, including the Thyreorporan Emausaurus and others which have not yet been referred to a specific genus. On Poland, the main basin if charaserized mostly by lack of marine microfauna in the Częstochowa-Zawiercie area. It is mainly composed by Foraminiferans: Ammodiscus glumaceous, Trochammina? sp., Lagenammina sp., Saccammina? sp. and Dentalina sp. (On Pabianice, Łutowiec and Żarki), as well as the Ostracodan Liasina sp.[2] The crustacean Estheria is relatively abundant, along traces of Diplocraterion and the inconclusive remains of Clams, Snails and Fish teeth (On Żuki, Gorzków, Choroń and Bobolice). The discussed settlement complex, due to its characteristic lithological education with floristic and faunistic analogies, allows without difficulty to parallelize with similarly developed sediments in the Polish Lowlands.[3]

Terrestrial environment of Łęczna (Lublin, Poland), based on the Bogdanka Coal Mine Flora. Dinosaurs are based on material found on various locations of the formation

Dinoflagellates

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Genus Species Location Material Notes Images

Nannoceratopsis[4]

  • Nannoceratopsis senex
  • Nannoceratopsis triceras
  • Nannoceratopsis raunsgardii
  • Nannoceratopsis ridingii
  • Kozłowice borehole.
  • Boroszów borehole

Dinocysts

A Dinophyceae Dinoflagellatan, type member of the family Nannoceratopsiaceae. The large amount of Cysts of the genus point to more diversified marine palaeoenvironments. N. senex is the most abundant.

Luehndea[4]

  • Luehndea spinosa
  • Kozłowice borehole.
  • Boroszów borehole

Dinocysts

A Dinophyceae Dinoflagellatan, type member of the family Luehndeoideae. Presence of Luehndea spinosa suggests Late Pliensbachian–earliest Toarcian age of studied assemblages.

Fungi

Genus Species Location Material Notes Images

Xylophagous Fungi[5]

Indeterminate

  • Brody-Lubienia borehole.[5]
  • Mechowo borehole
  • Gorzow Wielkopolski borehole

Fungal spores of different sizes associated with wood

Associated with a high rate of organic burial, the presence of Fungal Matter increased on the Uppermost layers of the Drzewica Formation, with a continue deposition between the T-OAE extincion, and several ups and downs on the Ciechocinek Strata, related with local climate and humidity changes. This is rather a reflection of the efficiency of terrestrial biodegradation.[5] Measured increasing of temperature favoured local fungal-mediated decomposition of plant litter, specifically of normally resistant wood.[5]

Xylophagous Fungi like Fomitopsis are probably the main origin for the Fungal Spores found on the Ciechocinek Formation

Sporonites[6]

  • Sporonites neddenii
  • Poręba

Spores

Fungal Spores with non diagnosed affinities, probably related with Pucciniomycetes inside Agaricomycotina. It is found associated with Pollen and Spores, interpreted as some sort of parasitism.

Sporonites was probably related with plant parasite fungus, such as Gymnosporangium

Invertebrata

Ichnofossils

Genus Species Location Material Notes Images

Planolites[7]

  • Planolites montanus
  • Planolites beverleyensis
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole

Burrowing and track ichnofossils.

Burrow-like ichnofossils. Planolites is really common in all types of the Ciechocinek Formation deposits. It is referred to vermiform deposit-feeders, mainly Polychaetes, producing active Fodinichnia. It is controversial, since is considered a strictly a junior synonym of Palaeophycus.[8]

Example of Planolites fossil

Palaeophycus[7]

  • Palaeophycus tubularis
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole

Burrowing and track ichnofossils.

Burrow-like ichnofossils. Palaeophycus is less common than Planolites in deposits of the Ciechocinek Formation. On the Kozlowice outcrop however there are numerous specimens occur, interpreted as the result of passive filling of polychaete burrows.[7]

Example of Palaeophycus fossil

Helminthopsis[7]

  • Helminthopsis isp.
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole

Burrowing and track ichnofossils.

Burrow-like ichnofossils. There are a few specimens in the Kozlowice outcrop. It is interpreted as a grazing trail or Fodinichnia, produced at shallow depth in sediment by Polychaetes and Priapulids.[7]

Example of Helminthopsis fossil

Gyrochorte[7]

  • Gyrochorte isp.
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole

Burrowing and track ichnofossils.

Burrow-like ichnofossils. Gyrochorte is interpreted as a result of active digging on the sediment by deposit-feeding worm-like animal, probably an Annelid or similar kinds of creatures.[7] On the Kozlowice strata only a few specimens where observed.

Protovirgularia[7]

  • Protovirgularia isp.
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole

Bilobate trace fossil

Bottom Trace Fossils. Protovirgularia is a Repichnia form, ascribed to the activity of Bivalves, leaving a trace due to the rhythmic action of a foot.[7]

Spongeliomorpha[7]

  • Spongeliomorpha isp.
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole

Burrows and associated traces

Burrow-like ichnofossils. Spongeliomorpha is believed to come from the domicile of Crustaceans: Anomuras (Probably Eocarcinoidea) and Decapodas (Probably Glypheidae), created as they dig in a firm, semiconsolidated substrate.[7] Local Spongeliomorpha could point to a transgressional Sea water impulse following a short episode of regression.[7]

Diplocraterion[7]

  • Diplocraterion parallelum
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole

Burrows and associated traces

Burrow-like ichnofossils. Most Diplocraterion show only protrusive spreit, like the local ones, produced under predominantly erosive conditions where the organism was constantly burrowing deeper into the substrate as sediment was eroded from the top. It can be Made by crustaceans, annelids or other benthic fauna.[7]

Diplocraterion parallelum diagram

Annelida

Genus Species Stratigraphic position Material Notes Images

Dictyothylakos[9]

  • Dictyothylakos pesslerae
  • Dictyothylakos sp. "Sing.,1964"
  • Dictyothylakos sp.
  • Silesian Coal Basin

Leech cocoons

Hirudinea cocoons, identified with palynological residues. The cocoons Dictyothylakos are common on flooded basin sediments, and implies not only the presence of parasitic leeches, but also the presence of large hosts nearby, that is proven on the case of Ciechocinek thanks to the dinosaur fossils from the German realm.

Example of Leech Cocoon

Brachiopoda

Genus Species Stratigraphic position Material Notes Images

Discina[10][11]

  • Discina cornucopiae
  • Green Series
  • Braunschweig

Cunchs

A Brachiopodan, member of Discinidae inside Discinida. The classification of the Discinidae is rather treated with confusion, due to the description and the identification of either extinct and extant genera and species.

Bivalvia

Genus Species Stratigraphic position Material Notes Images

Stalagmina[10][11]

  • Stalagmina koeneni
  • Green Series
  • Braunschweig

Cunchs

A Clam, member of Inoceramidae inside Myalinida. This genus resemble the Modern Pearl Oysters, although it was more likely a clam. The specimens are rather complete.

Parainoceramya[10][11]

  • Parainoceramya dubia
  • Green Series
  • Braunschweig

Cunchs

A Clam, member of Inoceramidae inside Myalinida. Pseudomytiloides dubius is a possible junior synonym. This genus resemble the Modern Pearl Oysters, although it was more likely a clam. The specimens are rather complete. It is the most common bivalve found locally.

Parainoceramya

Lucina[10][11]

  • Lucina plana
  • Green Series
  • Braunschweig

Cunchs

A Clam, member of Lucinidae inside Lucinida. Very abundant on the layers

Goniomya[10][11]

  • Goniomya rhombifera
  • Green Series
  • Braunschweig

Cunchs

A Clam, member of Pholadomyidae inside Pholadomyida. Rather common, but less abundant than other local genera

Neaera[10][11]

  • Neaera kayseri
  • Green Series
  • Braunschweig
  • Silesian Coal Basin

Cunchs

A Clam, member of Cuspidariidae inside Anomalodesmata. Marginal Marine to Mangrove swamp mollusc fauna, present on a rather large degree of salinities.

Gastropoda

Genus Species Stratigraphic position Material Notes Images

Cerithium[10][11]

  • Cerithium armatum
  • Cerithium roeceri
  • Green Series
  • Braunschweig

Cunchs

A Sea Snail, member of Cerithiinae inside Caenogastropoda. The main recovered gasteropod from the Green Series. Extant genus.

Cephalopoda

Genus Species Stratigraphic position Material Notes Images

Eleganticeras[12][13][14][15]

  • Eleganticeras elegantulum
  • Green Series
  • Boulder from Ahrensburg

Shells

A Hildoceratidae Ammonite.

Eleganticeras

Tiltoniceras[11][16]

  • Tiltoniceras antiquum
  • Tiltoniceras sp.
  • Green Series
  • Boulder from Ahrensburg

Shells

A Hildoceratidae Ammonite.

Grammoceras[16][12][11][15]

  • Grammoceras thouarcense
  • Grammoceras striatulum
  • Green Series
  • Boulder from Ahrensburg

Shells

A Hildoceratidae Ammonite.

Phylloceras[16][12][11][15]

  • Phylloceras heterophyllum
  • Green Series
  • Boulder from Ahrensburg

Shells

A Phylloceratidae Ammonite. Among the largest ammonites found on the Green Series, with specimens over 60 cm

Lobolytoceras[17][18]

  • Lobolytoceras siemensi
  • Green Series
  • Boulder from Ahrensburg

Shells

A Lytoceratidae Ammonite.

Lytoceras[16][12][11]

  • Lytoceras siemensi
  • Green Series
  • Boulder from Ahrensburg

Shells

A Lytoceratidae Ammonite.

Pseudolioceras[16][12][11]

  • Pseudolioceras lythense
  • Green Series
  • Boulder from Ahrensburg

Shells

A Harpoceratinae Ammonite.

Pseudolioceras

Dactylioceras[19][16][12][11][15]

  • Dactylioceras athleticum
  • Dactylioceras temperatum
  • Dactylioceras attenuatum
  • Dactylioceras tenuicostatum
  • Dactylioceras semicelatum
  • Dactylioceras cf. aequistriatum
  • Green Series
  • Boulder from Ahrensburg

Shells

A Dactylioceratinae Ammonite. Most common ammonite found on the Green Series and the different erratic boulders, as is the most common on the German realm, on the north and the south, with several specimens of different sizes.

Peronoceras[16][12][11][15]

  • Peronoceras fibulatum
  • Green Series
  • Boulder from Ahrensburg

Shells

A Dactylioceratinae Ammonite.

Parapassaloteuthis[10][16][12][11]

  • Parapassaloteuthis zieteni
  • Green Series
  • Braunschweig

Multiple Specimens.

A Passaloteuthididae Belemnoidean.

Cuspiteuthis[10][16][12]

  • Cuspiteuthis acuaria
  • Green Series
  • Braunschweig

Multiple Specimens.

A Megateuthididae Belemnoidean.

Acrocoelites[10][16][12]

  • Acrocoelites pyramidalis
  • Green Series
  • Braunschweig

Multiple Specimens.

A Megateuthididae Belemnoidean.

Beloteuthis[10][16][12]

  • Beloteuthis schubleri
  • Green Series
  • Braunschweig

Multiple Specimens.

A Beloteuthidae Mesoteuthoidean. Is a relatively small genus.

Beloteuthis

Teudopsis[10][16][12][20]

  • Teudopsis bollensis
  • Green Series
  • Braunschweig

Multiple Specimens.

A Teudopsidae Vampyropodan. Related to the modern Vampyroteuthis infernalis.

Jeletzkyteuthis[21][22]

  • Jeletzkyteuthis coriaceus
  • Green Series
  • Braunschweig

Multiple Specimens.

A Loligosepiidae Loligosepiidan (Vampyromorpha). Related to the modern Vampyroteuthis infernalis. Gladii of Loligosepia can be distinguished from Jeletzkyteuthis by the transition lateral field/hyperbolar zone. Described originally as Belopeltis bollensis.

Echinodermata

Genus Species Stratigraphic position Material Notes Images

Paleocoma[23]

  • Palaeocoma milleri
  • Green Series

Various specimens.

An Ophiuroidean Echinodermatan. It is relatively less abundant than in coeval strata.

Palaeocoma

Ophioderma[23]

  • Ophioderma dentata
  • Green Series

Various specimens.

An Ophiodermatidae Echinodermatan. Still alive today. On the Green Series, is related to deep basinal deposits.

Archastropecten[23]

  • Archastropecten hastingiae
  • Green Series

Various specimens.

An Astropectinidae Asteroidean. Related to freshwater debris, that probably caused changes to salinity and mass mortality of Echinoderms.

Archastropecten

Chladocrinus[23]

  • Chladocrinus basaltiformis
  • Green Series

Multiple specimens.

An Isocrinidae Crinoidean.

Echinoidea[23]

  • Echinoidea Indeterminate
  • Green Series

Spines from two Echinoidea Species

Sea Urchin Echinodermatan.

Crustacea

Genus Species Stratigraphic position Material Notes Images

Posidinia[24][16][12][11]

  • Posidinia opalina
  • Green Series

Various specimens.

An Eocarcinoidean Anomuran. It is among the oldest found Crabs worldwide. The genus is known locally mostly due to fragmentary specimens, related to less saline conditions.

Glyphea[16][12][11][24]

  • Glyphea liasina
  • Glyphea muensteri
  • Green Series
  • Silesian Coal Basin

Various specimens.

A Glypheidae Decapodan. The most complete Crustacean found on the formation

Proeryon[25]

  • cf. Proeryon sp.
  • Green Series (Grimmen)

Single specimen with preserved upper torax and chelae

A Proeryoninae Polychelidan.

Proeryon

Lioestheria[26]

  • Lioestheria sp.
  • Silesian Coal Basin

Docens of Specimens

A Clam shrimp (Phyllopodan) of the family Lioestheriidae. They fed on detritus, being very small slow moving, nektonic organisms that filter fed as they floated. The presence of Lioestheria marks the appearance of less saline conditions, as this is a mostly freshwater genus. The local Phyllopods are related with a great amount of freshwater debris (specially plants), and suggest seasonal changes on the rivers on the Toarcian Polish Basin.

Liasina[26]

  • Liasina lanceolata
  • Brody-Lubienia borehole

Valves

An Ostracodan of the family Pontocyprididae. Small marine ostracods related with abundant Green Algae environments

Ambigocythere[27]

  • Ambigocythere concentricostata
  • Green Series (Brandenburg)

Cunchs

A Marine Ostracodan with incertade Sedis assignation. The specimens of this genus are rather fargmentary.

Infracytheropteron[28]

  • Infracytheropteron supraliasicum
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Protostomia. The specimens of this genus are rather fragmentary and of uncertain nature.

Ogmoconchella[29][30][31]

  • Ogmoconchella adenticulata
  • Ogmoconchella aequalis
  • Ogmoconchella secunda
  • Green Series (Dobbertin)
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Healdiidae inside Podocopida. This genus is the main reported on the marine facies of the Dobbertin Clay Pit.

Ogmoconcha[32]

  • Ogmoconcha amalthei
  • Green Series (Dobbertin)
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Healdiidae inside Podocopida. Is probably present on all the Clay pits, altrougth the other locations specimens wheren´t published.

Ledahia[33]

  • Ledahia conviniens
  • Green Series (Dobbertin)
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Healdiidae inside Podocopida. One of the genus that reflect better the migration patterns of ostracodans on the Pliensbachian-Toarcian boundary.

Pseudohealdia[33]

  • Pseudohealdia pseudoespina
  • Green Series (Dobbertin)
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Healdiidae inside Podocopida. The genus is rare on the layers.

Hermiella[30]

  • Hermiella circumvallata
  • Hermiella intercedens
  • Green Series (Brandenburg)

Cunchs

A Marine Ostracodan of the family Healdiidae inside Podocopida. This genus is the main reported on the marine facies of the Dobbertin Clay Pit.

Cytheropteron[34][35]

  • Cytheropteron diversum
  • Green Series (Grimmen)

Cunchs

A Marine Ostracodan of the family Cytheruridae inside Podocopida. Is rare and the specimens found are rather incomplete.

Nanacythere[34][35]

  • Nanacythere (D.) firma
  • Nanacythere (D.) fissicosta
  • Nanacythere (G.) minor
  • Nanacythere (N.) simplex
  • Green Series (Grimmen)

Cunchs

A Marine Ostracodan of the family Cytheruridae inside Podocopida. The most abundant genus on the Grimmen Clay Pit.

Procytherura[36]

  • Procytherura suebica
  • Green Series (Grimmen)

Cunchs

A Marine Ostracodan of the family Cytheruridae inside Podocopida. A genus with well preserved specimens locally.

Gramannicythere[37][38]

  • Gramannicythere bachi
  • Gramannicythere coniuncta
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Protocytheridae inside Podocopida. Common, and associated to benthonic deposits. This genus maybe was able to resist relative changes in salinity.

Kinkelinella[27]

  • Kinkelinella procera
  • Green Series (Grimmen)

Cunchs

A Marine Ostracodan of the family Protocytheridae inside Podocopida. A genus related with Fish fossils and anoxic bottoms.

Bairdia[39][40]

  • Bairdia donzei
  • Bairdia guttulae
  • Bairdia michelseni
  • Bairdia praehilda
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Bairdiidae inside Bairdioidea. Abundant and diverse, is found associated with Ammonite shells.

Bairdiacypris[39][40]

  • Bairdiacypris anisica brevis
  • Bairdiacypris triasica postera
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Bairdiidae inside Bairdioidea. Less abundant than the genus Bairdia, is present on layers where Wood debris is more abundant.

Fabalacypris[39][40]

  • Fabalacypris symmetrica
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Bairdiidae inside Bairdioidea. Know only from the Green Series, is a rather unusual and complex genus.

Isobythocypris[39][40]

  • Isobythocypris cylindrica
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Bairdiidae inside Bairdioidea. It is a relatively abundant genus, but based on incomplete material.

Polycope[28]

  • Polycope tenuireticulata
  • Green Series (Thüringen)

Cunchs

A Marine Ostracodan of the family Polycopidae inside Cladocopina. Scarce but well preserved specimens.

Arachnida

Genus Species Stratigraphic position Material Notes Images

Seppo[41]

  • Seppo koponeni
  • Green Series

Single Incomplete Specimen.

A possible member of the superfamily Palpimanoidea.[41] It is the first confirmed spider from the lower jurassic, and a rare find, probably washed to the sea due to a hard wind related to hurricane action, present due to the measured moonsonal conditions of the formation, as on the rest of the Lower-Middle Toarcian strata. Probably a ground dewelling predator that hunted the abundant insect fauna present on the layers.[41] With a robust and well-armed legs I, directed forwards give the suggestion that they were preycapture appendages, a morphology typical of a sit-and-wait predator, while the short legs III are more typical on web spiders, especially Orbweavers, but also found on Palpimanoids, but not on that that are common substrate dwellers, that had legs more equal.[41] Seppo was probably not a habitual ground dweller, with armoured front legs related to capturing dangerous prey, such as many palpimanoids today are Araneophagous, for example.[41]

Seppo koponeni reconstruction on a Tree, as suggested for it's Leg III morphology

Insecta

Insects are a common terrestrial animals that where proabaly drifted to the sea due to Moonsonal conditions present on the Ciechocinek Formation.[42]

Genus Species Stratigraphic position Material Notes Images

Eolepidopterigidae[43]

  • Eolepidopterigidae indet.
  • Green Series

Wing Scales

A Lepidoptera (Moth) of the family Eolepidopterigidae. They are essential fossils for the Development of the color on Lepidopterans.

Zalmonites[44]

  • Zalmonites geinitzi
  • Green Series

Several Specimens

A Grasshopper.

Panorpidium[45][46]

  • Panorpidium geinitzi
  • Panorpidium magna
  • Panorpidium minima
  • Panorpidium media
  • Panorpidium reticulata
  • Panorpidium mesostena
  • Green Series

Several Specimens

A Grasshopper of the family Elcanidae. The species P. magna is among the largest Orthopterans of the Jurassic, while P. minima is among the Smallest.

Parelcana[44][47][48]

  • Parelcana tenuis
  • Green Series

Several Specimens

A Grasshopper of the family Elcanidae.

Parelcana[45][48]

  • Parelcana tenuis
  • Green Series

Several Specimens

A Grasshopper of the family Elcanidae.

Synelcana[45][48]

  • Synelcana muelleri
  • Green Series

Several Specimens

A Grasshopper of the family Elcanidae.

Locustopsis[45][48]

  • Locustopsis sp 1
  • Locustopsis sp 2
  • Locustopsis sp 3
  • Locustopsis sp 4
  • Locustopsis gyra
  • Locustopsis bernstorffi
  • Locustopsis sippeli
  • ?Locustopsis dubia
  • Locustopsis bernstorffi
  • Locustopsis nana
  • Locustopsis reducta
  • Locustopsis elongata
  • Locustopsis elegans
  • Locustopsis pulchella
  • Locustopsis lacera
  • Locustopsis procera
  • Locustopsis mecklenburgica
  • Locustopsis maculosa
  • Green Series

Several Specimens

A Grasshopper of the family Locustopsidae.

Plesioschwinzia[48][49]

  • Plesioschwinzia thalassophila
  • Green Series

Several Specimens

A Grasshopper of the family Locustopsidae.

Schwinzia[45][48]

  • Schwinzia sola
  • Green Series

Several Specimens

A Grasshopper of the family Locustopsidae.

Locustophanes[45][48]

  • Locustophanes rhipidophorus
  • Green Series

Several Specimens

A Grasshopper of the family Locustidae.

Protochaeta[45][48]

  • Protochaeta masculina
  • Protochaeta lanceolata
  • Green Series

Several Specimens

A Grasshopper of the family Regiatidae.

Protogryllus[47][48]

  • Protogryllus dobbertinensis
  • Protogryllus germanicus
  • Protogryllus sp.
  • Protogryllus vicinus
  • Protogryllus liadis
  • Protogryllus major
  • Protogryllus stenobasis
  • Protogryllus femina
  • Protogryllus acutipennis
  • Protogryllus minor
  • Protogryllus irregularis
  • Green Series

Several Specimens

A Cricket of the family Protogryllidae.

Griphopteron[50]

  • Griphopteron molle
  • Green Series

Several Specimens

A Cricket of the family Blattogryllidae.

Durnovaria[50]

  • Durnovaria cf. parallela
  • Green Series

Several Specimens

A Stick Insect of the family Aerophasmidae.

Aenne[51]

  • Aenne liasina
  • Green Series

Wings

The oldest known non biting midge (Chironomidae)

Dobbertiniopteryx[52]

  • Dobbertiniopteryx capniomimus
  • Green Series

Wings

A Stonefly of the family Protostomia

Mesotipula[53][54]

  • Mesotipula siggiae
  • Mesotipula vicina
  • Mesotipula cf. vicina
  • Mesotipula mirabilis
  • Mesotipula curvata
  • Mesotipula geinitzi
  • Mesotipula minuta
  • Mesotipula lapidaria
  • Mesotipula parva
  • Green Series

Several Specimens

A Crane Fly of the family Limoniidae.

Architipula[53][47][48]

  • Architipula stigmatica
  • Architipula latipennis
  • Architipula elegans
  • Architipula seebachi
  • Architipula nana
  • Architipula pusilla
  • Architipula pulla
  • Architipula debilis
  • Architipula simplex
  • Architipula intermedia
  • Architipula obliqua
  • Architipula areolata
  • Architipula maior
  • Architipula conspicua
  • Architipula clara
  • Architipula dubia
  • Green Series

Several Specimens

A Crane Fly of the family Limoniidae. A. nana can be the smallest Crane fly of the Jurassic.

Eotipula[44]

  • Eotipula defuncta
  • Eotipula mortua
  • Eotipula coarctata
  • Green Series

Several Specimens

A Crane Fly of the family Limoniidae.

Phryganidium[44]

  • Phryganidium minimum
  • Green Series

Several Specimens

A Crane Fly of the family Limoniidae.

Grimmenia[53]

  • Grimmenia prima
  • Grimmenia secunda
  • Green Series

Several Specimens

A Crane Fly of the family Limoniidae.

Rhaetomyia[50]

  • Rhaetomyia herrigi
  • Green Series

Several Specimens

A Phantom midge of the family Chaoboridae.

Grimmyia[50]

  • Grimmyia baltica
  • Green Series

Several Specimens

A Snipe Fly of the family Rhagionidae.

Liassobrachyceron[55]

  • Liassobrachyceron kotejai
  • Green Series

Several Specimens

A Snipe Fly of the family Rhagionidae.

Palaeobrachyceron[50]

  • Palaeobrachyceron willmanni
  • Green Series

Several Specimens

A Snipe Fly of the family Rhagionidae.

Antefungivora[50]

  • Antefungivora germanica
  • Green Series

Several Specimens

A Fly of the family Antefungivoridae.

Archibio[50]

  • Archibio mycetophilinus
  • Green Series

Several Specimens

A Fly of the family Antefungivoridae.

Pleciofungivora[50]

  • Pleciofungivora liassica
  • Green Series

Several Specimens

A Fly of the family Pleciofungivoridae.

Archirhyphus[50]

  • Archirhyphus geinitzi
  • Green Series

Several Specimens

A Fly of the family Protorhyphidae.

Protorhyphus[47]

  • Protorhyphus simplex
  • Green Series

Several Specimens

A Fly of the family Protorhyphidae.

Protobrachyceron[56]

  • Protobrachyceron liasinum
  • Green Series

Several Specimens

A Fly of the family Protobrachyceridae.

Heterorhyphus[50]

  • Heterorhyphus triangularis
  • Green Series

Several Specimens

A Fly of the family Heterorhyphidae.

Eoditomyia[50]

  • Eoditomyia primitiva
  • Green Series

Several Specimens

A Fly of the family Eoditomyidae.

Archipleciomima[50]

  • Archipleciomima sp.
  • Archipleciomima germanica
  • Green Series

Several Specimens

A Fly.

Protoplecia[50]

  • Protoplecia klafacki
  • Protoplecia liasina
  • Green Series

Several Specimens

A Fly of the family Protopleciidae.

Mailotrichocera[57]

  • Mailotrichocera mikereichi
  • Mailotrichocera variabilis
  • Mailotrichocera zessini
  • Green Series

Several Specimens

A Winter Crane Fly of the family Trichoceridae.

Nannotanyderus[58]

  • Nannotanyderus grimmenensis
  • Nannotanyderus krzeminskii
  • Green Series

Several Specimens

A primitive Crane Fly of the family Tanyderidae.

Eoptychoptera[47]

  • Eoptychoptera simplex
  • Green Series

Several Specimens

A Phantom crane fly of the family Ptychopteridae.

Eolimnobia[47]

  • Eolimnobia geinitzi
  • Green Series

Several Specimens

A Phantom crane fly of the family Ptychopteridae.

Tanypsycha[58]

  • Tanypsycha connexa
  • Green Series

Several Specimens

A Moth fly of the family Psychodidae.

Liassopsychodina[58]

  • Liassopsychodina pommerana
  • Green Series

Several Specimens

A Moth fly of the family Psychodidae.

Mesorhyphus[56]

  • Mesorhyphus nanus
  • Green Series

Several Specimens

A Wood Gnat of the family Anisopodidae.

Metatrichopteridium[59]

  • Metatrichopteridium confusum
  • Green Series

Several Specimens

A rare Fly of the family Hennigmatidae.

Orthophlebia[47]

  • Orthophlebia germanica
  • Orthophlebia elongata
  • Orthophlebia fuscipennis
  • Orthophlebia limnophila
  • Orthophlebia cf. limnophila
  • Orthophlebia radialis
  • Orthophlebia liadis
  • Orthophlebia maculata
  • Orthophlebia vicina
  • Orthophlebia brunsvicensis
  • Green Series

Several Specimens

A Mecopteran of the family Orthophlebiidae.

Orthophlebia fossil

Neorthophlebia[47]

  • Neorthophlebia maculipennis
  • Neorthophlebia megapolitana
  • Neorthophlebia pallida
  • Neorthophlebia debilis
  • Neorthophlebia simillima
  • Neorthophlebia medialis
  • Neorthophlebia stigmatica
  • Neorthophlebia acutipennis
  • Green Series

Several Specimens

A Mecopteran of the family Bittacidae.

Parabittacus[60]

  • Parabittacus analis
  • Green Series

Several Specimens

A Mecopteran of the family Bittacidae.

Mesobittacus[47]

  • Mesobittacus minutus
  • Green Series

Several Specimens

A Mecopteran of the family Bittacidae.

Pseudopolycentropus[44]

  • Pseudopolycentropus triangularis
  • Pseudopolycentropus perlaeformis
  • Pseudopolycentropus sp.
  • Green Series

Several Specimens

A Mecopteran of the family Pseudopolycentropodidae.

Mesochorista?[44]

  • Mesochorista sinuata
  • Green Series

Several Specimens

A Mecopteran of the family Permochoristidae.

Liasocoris[61]

  • Liasocoris hainmuelleri
  • Green Series

Several Specimens

A Hemipteran.

Fulgoridium[50][47]

  • Fulgoridium egens
  • Fulgoridium lapideum
  • Fulgoridium dubium
  • Fulgoridium latum
  • Fulgoridium venosum
  • Fulgoridium pallidum
  • Fulgoridium geinitzi
  • Fulgoridium spilographum
  • Fulgoridium oligoneurum
  • Fulgoridium regulare
  • Fulgoridium graphipterum
  • Fulgoridium inconspicuum
  • Fulgoridium quadrisignatum
  • Fulgoridium breviradiatum
  • Fulgoridium modestum
  • Fulgoridium anale
  • Fulgoridium marginepunctatum
  • Fulgoridium picturatum
  • Fulgoridium anomalum
  • Fulgoridium stigmaticum
  • Fulgoridium punctatum
  • Fulgoridium mortuum
  • Fulgoridium alatum
  • Fulgoridium nodosum
  • Fulgoridium megapolitanum
  • Fulgoridium pulchrum
  • Fulgoridium remotum
  • Fulgoridium multivenosum
  • Fulgoridium debile
  • Fulgoridium trifurcatum
  • Fulgoridium litorale
  • Fulgoridium parvispilum
  • Fulgoridium retractum
  • Fulgoridium polyneurum
  • Fulgoridium oligospilum
  • Fulgoridium nubeculum
  • Fulgoridium obtusum
  • Fulgoridium grave
  • Fulgoridium fenestratum
  • Fulgoridium breve
  • Fulgoridium dilutum
  • Fulgoridium multipunctatum
  • Fulgoridium ampliatum
  • Fulgoridium clavatum
  • Fulgoridium nebulosum
  • Fulgoridium vicinum
  • Fulgoridium curvipenne
  • Fulgoridium elegantulum
  • Fulgoridium defunctum
  • Fulgoridium plicatum
  • Fulgoridium exhumatum
  • Fulgoridium angulosum
  • Fulgoridium acutum
  • Fulgoridium rotundatum
  • Fulgoridium ancylla
  • Fulgoridium bifurcatum
  • Fulgoridium inaequale
  • Fulgoridium brachyptilum
  • Fulgoridium intercalatum
  • Fulgoridium balticum
  • Green Series

Several Specimens

A Hemipteran of the family Fulgoridiidae. The colossal abundance of the genus maybe it's related to a preference for seashore habitats. Some specimens are indentinguible, making possible some species synonymous.

Metafulgoridium[47]

  • Metafulgoridium spilotum
  • Metafulgoridium ampliatum
  • Metafulgoridium graptum
  • Margaroptilon germanicum
  • Green Series

Several Specimens

A Hemipteran of the family Fulgoridiidae.

Archiconiopteryx[62]

  • Archiconiopteryx liasina
  • Green Series

Several Specimens

A Hemipteran of the family Archiconiopterygidae.

Liadopsylla[63]

  • Liadopsylla geinitzi
  • Green Series

Several Specimens

A Hemipteran of the family Liadopsyllidae.

Hadrocoris[56]

  • Hadrocoris scutellaris
  • Green Series

Several Specimens

A Hemipteran of the family Hadrocoridae.

Progonocimex[56]

  • Progonocimex jurassicus
  • Progonocimex liasinus
  • Green Series

Several Specimens

A Hemipteran of the family Progonocimicidae.

Eocercopis[47]

  • Eocercopis ancyloptera
  • Eocercopis similis
  • Green Series

Several Specimens

A Hemipteran of the family Progonocimicidae.

Archicercopis[47]

  • Archicercopis falcatus
  • Green Series

Several Specimens

A Hemipteran of the family Progonocimicidae.

Archegocimex[44]

  • Archegocimex geinitzi
  • Archegocimex primitiva
  • Archegocimex liadis
  • Green Series

Several Specimens

A Hemipteran of the family Archegocimicidae.

Progonocoris[56]

  • Progonocoris pictus
  • Green Series

Several Specimens

A Hemipteran of the family Archegocimicidae.

Anosmus[47]

  • Anosmus spilopterus
  • Green Series

Several Specimens

A Hemipteran of the family Archegocimicidae.

Diatillus[63]

  • Diatillus debilis
  • Green Series

Several Specimens

A Hemipteran of the family Archegocimicidae.

Pachymeridium[64]

  • Pachymeridium dubium
  • Green Series

Several Specimens

A Hemipteran of the family Pachymeridiidae.

Sisyrocoris[63]

  • Sisyrocoris rudis
  • Green Series

Several Specimens

A Hemipteran of the family Pachymeridiidae.

Hypocimex[47]

  • Hypocimex membranaceus
  • Green Series

Several Specimens

A Hemipteran of the family Pachymeridiidae.

Apsicoria[47]

  • Apsicoria semideleta
  • Green Series

Several Specimens

A Hemipteran of the family Pachymeridiidae.

Cathalus[47]

  • Cathalus alutaceus
  • Green Series

Several Specimens

A Hemipteran of the family Pachymeridiidae.

Psychrocoris[56]

  • Psychrocoris cuneifera
  • Green Series

Several Specimens

A Hemipteran of the family Pachymeridiidae.

Cuneocoris[56]

  • Cuneocoris geinitzi
  • Green Series

Several Specimens

A Hemipteran of the family Cuneocoridae.

Apopnus[63]

  • Apopnus magniclavus
  • Green Series

Several Specimens

A Hemipteran of the family Naucoridae.

Aphlebocoris[44]

  • Aphlebocoris punctata
  • Aphlebocoris nana
  • Green Series

Several Specimens

A Hemipteran of the family Naucoridae.

Archijassus[44][65]

  • Archijassus heeri
  • Archijassus pachyneura
  • Green Series

Several Specimens

A Hemipteran of the family Archijassidae.

Ardela[50][65]

  • Ardela grimmenensis
  • Green Series

Several Specimens

A Hemipteran of the family Archijassidae.

Liassocercopis[50][65]

  • Liassocercopis schnicki
  • Green Series

Several Specimens

A Hemipteran of the family Hylicellidae.

Toarcopsyllidium[50][65]

  • Toarcopsyllidium ernsti
  • Green Series

Several Specimens

A Hemipteran of the family Protopsyllidiidae.

Grimmenaphis[50][65]

  • Grimmenaphis magnifica
  • Green Series

Several Specimens

A Hemipteran of the family Oviparosiphidae.

Grimmaratavites[66]

  • Grimmaratavites mirabilis
  • Green Series

Several Specimens

A Hemipteran of the family Karatavitidae.

Liassochrysa[67]

  • Liassochrysa stigmatica
  • Green Series

Several Specimens

A Neuropteran of the family Mantispidae. It is the earliest know Chrysopid

Prohemerobius[68]

  • Prohemerobius prodromus
  • Prohemerobius ovatus
  • Prohemerobius anomalus
  • Prohemerobius costalis
  • Prohemerobius persimilis
  • Prohemerobius geinitzi
  • Prohemerobius liasinus
  • Prohemerobius major
  • Prohemerobius parvulus
  • Prohemerobius geinitzianus
  • Prohemerobius dilaroides
  • Prohemerobius chryseus
  • Prohemerobius latus
  • Green Series

Several Specimens

A Neuropteran of the family Prohemerobiidae.

Prohemerobius at the bottom

Mesosmylina[69]

  • Mesosmylina falcifera
  • Green Series

Several Specimens

A Neuropteran of the family Osmylidae.

Polyosmylus[50]

  • Polyosmylus excelsus
  • Green Series

Several Specimens

A Neuropteran.

Mesoleon[50]

  • Mesoleon dobbertinianus
  • Green Series

Several Specimens

A Neuropteran.

Loxophleps[47][50]

  • Loxophleps costalis
  • Green Series

Several Specimens

A Neuropteran.

Melaneimon[47][50]

  • Melaneimon dubium
  • Green Series

Several Specimens

A Neuropteran.

Melamnous[47]

  • Melamnous indistinctus
  • Green Series

Several Specimens

A Neuropteran.

Solenoptilon[47]

  • Solenoptilon kochi
  • Green Series

Several Specimens

A Neuropteran of the Family Solenoptilidae.

Epigambria[47]

  • Epigambria longipennis
  • Green Series

Several Specimens

A Neuropteran of the Family Epigambriidae.

Apeirophlebia[47]

  • Apeirophlebia grandis
  • Green Series

Several Specimens

A Neuropteran of the Family Psychopsidae. An unexpectec giant Silky lacewing, with a size up to 7 cm.

Magnasupplephlebia[70][71]

  • Magnasupplephlebia kallweita
  • Magnasupplephlebia intercalaria
  • Green Series

Several Specimens

An Odonatan. Very large dragonfly, with a wingspan of 13 cm

Liadothemis[47]

  • Liadothemis hydrodictyon
  • Liadothemis major
  • Liadothemis geinitzi
  • Liadothemis insignis
  • Green Series

Several Specimens

An Odonatan.

Petrothemis[47]

  • Petrothemis singularis
  • Green Series

Several Specimens

An Odonatan.

Parelthothemis[47]

  • Parelthothemis dobbertinensis
  • Green Series

Several Specimens

An Odonatan.

Anomothemis[47]

  • Anomothemis brevistigma
  • Green Series

Several Specimens

An Odonatan.

Rhabdothemis[47]

  • Rhabdothemis strigivena
  • Green Series

Several Specimens

An Odonatan.

Temnostigma[47]

  • Temnostigma singulare
  • Green Series

Several Specimens

An Odonatan.

Pycnothemis[47]

  • Pycnothemis densa
  • Green Series

Several Specimens

An Odonatan.

Archithemis[47]

  • Archithemis brodiei
  • Green Series

Several Specimens

An Odonatan of the family Archithemistidae.

Selenothemis[47]

  • Selenothemis liadis
  • Green Series

Several Specimens

An Odonatan of the family Selenothemistidae.

Heterothemis[72]

  • Heterothemis brodiei
  • Green Series
  • Boulder from Ahrensburg

Several Specimens

An Odonatan of the family Liassogomphidae.

Anisozygopteron[47]

  • Anisozygopteron geinitzianum
  • Green Series

Several Specimens

An Odonatan of the family Myopophlebiidae.

Dialothemis[73]

  • Dialothemis dubia
  • Green Series

Several Specimens

An Odonatan of the family Selenothemistidae.

Eosagrion[56]

  • Eosagrion risi
  • Green Series

Several Specimens

An Odonatan of the family Eosagrionidae.

Turanopteron[50]

  • Turanopteron pommerana
  • Green Series

Several Specimens

An Odonatan of the family Asiopteridae.

Liadobracona[74]

  • Liadobracona raduhna
  • Green Series

Several Specimens

A Hymenopteran of the family Ephialtitidae.

Brigittepterus[75]

  • Brigittepterus brauckmanni
  • Green Series

Several Specimens

A Hymenopteran of the family Ephialtitidae.

Xyelula[75]

  • Xyelula benderi
  • Green Series

Several Specimens

A Hymenopteran of the family Sepulcidae.

Mesoblattina[76]

  • Mesoblattina protypa
  • Green Series

Several Specimens

A Cockroach of the family Mesoblattinidae.

Caloblattina[76][77]

  • Caloblattina mathildae
  • Green Series

Several Specimens

A Cockroach of the family Caloblattinidae.

Liadoblattina[76][78]

  • Liadoblattina blakei
  • Green Series

Several Specimens

A Cockroach of the family Raphidiomimidae.

Eublattula[47][76]

  • Eublattula crassivena
  • Green Series

Several Specimens

A Cockroach. Blattulidae is an extinct cockroach family which was widely distributed around the world and lasted from Late Triassic to Cretaceous.

Blattula[72][76]

  • Blattula langfeldti
  • Blattula dubia
  • Green Series
  • Boulder from Ahrensburg

Several Specimens

A Cockroach. Blattulidae is an extinct cockroach family which was widely distributed around the world and lasted from Late Triassic to Cretaceous.

Dicronemoura[50]

  • Dicronemoura furcata
  • Green Series

Several Specimens

A Plecopteran of the family Perlariopseidae.

Liassopsocus[50]

  • Liassopsocus lanceolatus
  • Green Series

Several Specimens

A Permopsocidan of the family Psocidiidae.

Archipsylla[50]

  • Archipsylla primitiva
  • Green Series

Several Specimens

A Permopsocidan of the family Archipsyllidae.

Undacypha[50]

  • Undacypha europaea
  • Green Series

Several Specimens

A Dicondylian of uncertaing Placement.

Parnidium[44]

  • Parnidium frechi
  • Parnidium geinitzi
  • Green Series

Several Specimens

A Coleopteran.

Polypamon[79]

  • Polypamon byrrhoides
  • Green Series

Several Specimens

A Coleopteran.

Bathygerus[79]

  • Bathygerus bellus
  • Bathygerus divergens
  • Green Series

Several Specimens

A Coleopteran.

Plastonebria[79]

  • Plastonebria scudderi
  • Green Series

Several Specimens

A Coleopteran.

Plastobuprestites[79]

  • Plastobuprestites elegans
  • Green Series

Several Specimens

A Coleopteran.

Nannoodes[79]

  • Nannoodes pseudocistela
  • Green Series

Several Specimens

A Coleopteran.

Pseudocyphon[79]

  • Pseudocyphon geinitzi
  • Green Series

Several Specimens

A Coleopteran.

Keleusticus[79]

  • Keleusticus zirkeli
  • Green Series

Several Specimens

A Coleopteran.

Allognosis[79]

  • Allognosis nitens
  • Green Series

Several Specimens

A Coleopteran.

Enamma[79]

  • Enamma striatum
  • Green Series

Several Specimens

A Coleopteran.

Pseudoprionites[79]

  • Pseudoprionites liasinus
  • Green Series

Several Specimens

A Coleopteran.

Nebrioides[79]

  • Nebrioides dobbertinensis
  • Green Series

Several Specimens

A Coleopteran.

Paracurculium[79]

  • Paracurculium punctatum
  • Green Series

Several Specimens

A Coleopteran.

Apioderes[79]

  • Apioderes punctatus
  • Green Series

Several Specimens

A Coleopteran.

Bareus[79]

  • Bareus strigipennis
  • Green Series

Several Specimens

A Coleopteran.

Anypostatus[79]

  • Anypostatus taurus
  • Green Series

Several Specimens

A Coleopteran.

Periboloptera[79]

  • Periboloptera rotunda
  • Green Series

Several Specimens

A Coleopteran.

Clinomerus[79]

  • Clinomerus laticollis
  • Green Series

Several Specimens

A Coleopteran.

Anomerus[79]

  • Anomerus punctifer
  • Green Series

Several Specimens

A Coleopteran.

Masselytron[79]

  • Masselytron quinquestriatum
  • Green Series

Several Specimens

A Coleopteran.

Oxytoroptera[79]

  • Oxytoroptera mediocris
  • Green Series

Several Specimens

A Coleopteran.

Ecthlimma[47]

  • Ecthlimma forficuloides
  • Green Series

Several Specimens

A Coleopteran.

Onkedodimus[47]

  • Onkedodimus discicollis
  • Green Series

Several Specimens

A Coleopteran.

Apsychus[47]

  • Apsychus alutaceus
  • Green Series

Several Specimens

A Coleopteran.

Gyrinulopsis[47]

  • Gyrinulopsis nanu
  • Green Series

Several Specimens

A Coleopteran.

Anancaeon[47]

  • Anancaeon microcephalum
  • Green Series

Several Specimens

A Coleopteran.

Hydrobiites[47]

  • Hydrobiites dobbertinensis
  • Hydrobiites punctatostriatus
  • Green Series

Several Specimens

A Coleopteran of the family Permosynidae.

Thoracotes[44]

  • Thoracotes dubius
  • Green Series

Several Specimens

A Coleopteran of the family Trogossitidae.

Eurynucha[44]

  • Eurynucha pseudobuprestis
  • Green Series

Several Specimens

A Coleopteran of the family Buprestidae.

Coptogyrinus[44]

  • Coptogyrinus scutellatus
  • Green Series

Several Specimens

A Coleopteran of the family Gyrinidae.

Carabites[44]

  • Carabites dubius
  • Carabites geinitzi
  • Green Series

Several Specimens

A Coleopteran of the family Carabidae.

Nele[50]

  • Nele jurassica
  • Green Series

Several Specimens

A Grylloblattodean of the family Bajanzhargalanidae.

Vertebrata

Fishes

Actinopteri

Genus Species Stratigraphic position Material Notes Images

Saurorhynchus[80]

  • Saurorhynchus hauffi
  • Green Series

Multiple specimens.

The youngest representative of the family Saurichthyidae, known for its large jaws, similar to modern Belonidae.

Grimmenichthys[81]

  • Grimmenichthys ansorgei
  • Cf. Grimmenichthys ansorgei
  • Green Series

GG 439/1, articulated, but incompletely preserved specimen

A member of the Pholidophoriformes.

Pholidophoriformes[81]

  • Pholidophoriformes indet.
  • Green Series

GG 439/2

Non assigned to an especific genus

Pholidophorus[82]

  • Pholidophorus sp.
  • Ahrensburg erratic assemblage[82]

Fragmentary remains

Non assigned to an especific species

Pachycormiformes[81]

  • Pachycormiformes indet.
  • Green Series

Unknown, only cited.

Non assigned to an especific genus

Leptolepis[83][84]

  • Leptolepis coryphaenoides
  • Leptolepis normandica
  • Leptolepis jaegeri
  • Leptolepis sp.
  • Green Series
  • Ahrensburg erratic assemblage[82]
  • GG 431/3 slightly disarticulated, incomplete head
  • GG 431/7 subadult, almost complete
  • NRM P 6091 (cast of MNH P. 23834) fragmentary head
  • GG 431/19 isolated head with pectoral girdle
  • GG 431/20 incomplete head

Type member of the Leptolepidae.

Leptolepidae[83][2][3]

Indeterminate

  • Green Series
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Parkoszowice borehole
  • GG 431/5a, b isolated head with missing postcranial bones.[83]
  • Assigned Teeth

Non assigned to a concrete genus.

Proleptolepis[83]

Proleptolepis sp.

  • Green Series
  • Boroszów Borehole
  • Parkoszowice borehole
  • MV 202612, fragmentary head .[83]
  • Assigned Teeth

Member of the Leptolepidae.

Tetragonolepis[2][3]

  • Tetragonolepis semicincta
  • Boroszów Borehole
  • Parkoszowice borehole
  • Fragmentary Head
  • Assigned Teeth

Member of the Semionotidae.

Lepidotes[85]

  • Lepidotes elvensis
  • Green Series
  • Ahrensburg erratic assemblage[82]

Various specimens

Member of the Semionotidae.

Lepisosteiformes[2][3]

  • Lepisosteiformes Indeterminate
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Parkoszowice borehole

Teeth & Scales

Member of the Semionotidae.

Grimmenodon[86]

  • Grimmenodon aureum
  • Green Series

GG 437, Fragmentary Skull

Member of the Pycnodontiformes.

Dapedium[87][88]

  • Dapedium pholidotum
  • Green Series

Fragmentary specimen.

Member of the Dapediidae.[88]

Gyrosteus[89]

  • Gyrosteus mirabilis
  • Ahrensburg erratic assemblage

GPIH 4864, Hyomandibula

Member of the Chondrosteidae. This find, which probably originates from the western Baltic basin between Bornholm Island (Denmark) and northeastern Germany, markedly expands the known range of this chondrosteid taxon across the northern part of the strait connecting the Boreal Sea with the Tethys Ocean during the Early Jurassic.[89] The relatively small size of the hyomandibula compared to the largest English material (ca. 40% smaller) may be related to a younger ontogenetic stage of the individual.[89]

Chondrichthyes

Genus Species Stratigraphic position Material Notes Images

Hybodus[88]

  • Hybodus hauffianus
  • Green Series (Grimmen)
  • Ahrensburg erratic assemblage[82]
  • Fragmentary Fin spine
  • Supposed coprolithes
  • Teeth

Member of the Hybodontiformes. Related to Hybodus hauffianus and other genera from the southern of Germany

Synechodontiformes[2][3]

  • Synechodontiformes Indeterminate
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Parkoszowice borehole

Teeth

Member of the Squalomorphii. Small to medium benthonic sharks.

Hemiscylliidae?[2][3]

  • Hemiscylliidae? Indeterminate
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Parkoszowice borehole

Teeth

Member of the Orectolobiformes. Small benthonic sharks.

Sarcopterygii

Genus Species Location Material Notes Images

Coelacanthidae[2][3]

Cf. Coelacanthidae

  • Boroszów Borehole

Scales

A Coelacanth, member of the family Coelacanthiformes inside Actinistia. The scales recovered are homologous to that found on the Triassic coelacanths Axelia & Wimania.

Ptychoceratodus[2][3]

Ptychoceratodus rectangulus

  • Brody-Lubienia borehole
  • Boroszów Borehole
  • Parkoszowice borehole

Teeth

A Lungfish, type member of the family Ptychoceratodontidae inside Dipnoi. A Lungfish of relative small size, related with Freshwater Debris, found associated with Deltaic Deposits.

Ceratodus[2][3]

  • Ceratodus cf. rectangularis
  • Brody-Lubienia borehole
  • Parkoszowice borehole
  • Teeth & Scales
  • Asiggned fragmentary Teeth

A Lungfish, member of the Neoceratodontidae inside Ceratodontiformes. A Lungfish, abundant on Europe since the Hettangian to the Bajocian on the Jurassic. The Species are known by a few scarce fragments, and are related with Triassic species found along Germany.

Illustration of Ceratodus by Heinrich Harder

Reptiles

Ichthyosaurs

Genus Species Stratigraphic position Material Notes Images

Stenopterygius[90][88]

  • Stenopterygius cf. quadriscissus
  • Green Series

Posterior left half of the cranium.

A Icthyosaur of the family Stenopterygiidae inside Thunnosauria. A common Toarcian Ichthyosaur, present on multiple layers. The rather exquisite level of preservation has led to know even the coloration.

Restoration

Neoichthyosauria[91]

  • Neoichthyosauria indet.
  • Green Series

Four articulated tail vertebrae.

An indeterminate ichthyosaur, has been assigned to the species Stenopterygius longifrons

Ophthalmosauridae[92][93][94]

  • Ophthalmosauridae indet.
  • Green Series

Partial skull and associated postcranial elements preserved in a concretion

An indeterminate ichthyosaur. It has an expanded basipterygoid processes on the basisphenoid, only currently known in members of the Ophthalmosauridae

Ichthyosauria[88]

  • Ichthyosauria indet.
  • Green Series

Presacral centrum, ?sacral centrum

An indeterminate ichthyosaur.

Ichthyosauria[95][96][97]

  • Ichthyosauria indet.
  • Ahrensburg Erratic Boulders

Skull, coracoid and associated rib fragment

An indeterminate ichthyosaur. Assigned to Ichthyosaurus sp., but also suggested affinities to "Leptopterygius" (= Temnodontosaurus) platyodon.

Sauropterygia

Genus Species Stratigraphic position Material Notes Images

Plesiosauria[88][98]

  • Plesiosauria indeterminate
  • Cf. Plesiosauria indeterminate
  • Green Series

Three caudal vertebrae, Gastralia, Cervical rib and other postcranial elements

Non assigned to a concrete genus, probably are related to the Plesiosaurian genera from the Posidonia Shale

Plesiosauroidea[88][98]

  • Plesiosauroidea indeterminate
  • Green Series

Cervical vertebra, Caudal centrum & Phalanx

Non assigned to a concrete genus, probably are related to the Plesiosaurian genera from the Posidonia Shale

Microcleididae[98]

  • Microcleididae indeterminate
  • Forst Hagen gravel pit

Isolated tooth crown & isolated cervical vertebrae

Non assigned to a concrete genus, probably are related to the Plesiosaurian genus Microcleidus or to Seeleyosaurus

Seeleyosaurus?[98]

  • Seeleyosaurus? sp.
  • Forst Hagen gravel pit

Three articulated dorsal vertebrae

Non assigned to a concrete genus, probably are related to the Plesiosaurian genus Microcleidus

Meyerasaurus?[98]

  • Meyerasaurus sp.
  • Vierbergen near Ahrensburg

Incomplete coracoid and associated rib

Material referred to the family Rhomaleosauridae

Crocodyliformes

Genus Species Stratigraphic position Material Notes Images

Goniopholididae?[88]

  • Goniopholididae indeterminate
  • Green Series

Cervical vertebra

Affinities with Goniopholididae inside Neosuchia. Non assigned to a concrete genus, originally labeled as Mesoeucrocodylia indet. can be the earliest representative of the group on Europe.

Thalattosuchia[88]

  • Thalattosuchia indet
  • Green Series
  • Vierbergen near Ahrensburg

Partial rostrum with teeth

Affinities with Thalattosuchia inside Neosuchia. Probably related to Pelagosaurus

Steneosaurus[88][99]

  • Steneosaurus bollensis
  • Green Series

Incomplete skull and associated osteoderm

Affinities with Thalattosuchia inside Neosuchia. A marine crocodrylomorph with a diet probably based on fish.[100] It is the best known member of the family Teleosauriade.

Theropoda

Genus Species Stratigraphic position Material Notes Images

Orionides?[101][102]

Orionides? Indeterminate

  • Forst Hagen gravel pit

Dorsal Vertebrae

A possible Tetanurae theropod. The affinities of the Specimen aren't clear due to its fragmentary nature. Has been classified as Saurischia indeterminate, although shows clearly characters of the Orionides group (concave articular surfaces and a dished lateral pleurocoel, remnants of the neural arch and postzygapophyses).[101] The vertebrae centrum measures 80 mm, implying a medium-sized theropod (~5 m long).[101] Can be related with Yunyangosaurus.

Sauropoda

Genus Species Stratigraphic position Material Notes Images

Gravisauria[102][103]

Gravisauria Indeterminate[103]

  • Green Series
  • Iliac preacetabular process
  • Distal right pubis
  • Proximal left ischium
  • Proximal right ischium

A Gravisaurian Sauropod. The specimen is believed to be a juvenile, based on the ossification and unfused spine. Has affinities with the genus Tazoudasaurus and it is clearly distinctive form the also toarcian Ohmdenosaurus, who is thought to be more basal.[103] The pelvic girdle elements can be clearly placed among the Sauropoda, on account of the presence of an elongated and strongly dorsally expanded iliac preacetabular process a possible relative.[103] The ischia GG411/3-4 resemble those of Tazoudasaurus in exhibiting a subtriangular iliac peduncle which create a short anteriorly directed expansion to reach the medial acetabular rim, and are slightly less developed than the Genus Barapasaurus.[103]

Barapasaurus can be a close relative of the Grimmen Sauropod

Gravisauria[102][103]

Gravisauria Indeterminate[103]

  • Green Series
  • Single dorsal neural spine

A Gravisaurian Sauropod. The specimen is believed to be a juvenile (its comparatively small size is indicative of belonging to a not fully grown individual) and probably related to the Asian genus Zizhongosaurus, as shares characters with the neural spine V9067.1.[103] GG412 within Gravisauria is indicated by the presence of a well-developed spinodiapophyseal lamina.[103]

Thyreophora

Genus Species Stratigraphic position Material Notes Images

Emausaurus[104][105]

  • Emausaurus ernsti
  • Green Series

Right side of the skull, the right lower jaw, caudal vertebrae, neural arches, a radius, a metatarsal, a claw, fragments of ribs, scutes and plates.[104]

A Thyreophoran ornithischian. Its juvenile status makes controversial its phylogeny.[104]

Emausaurus attacked by a theropod

Plantae

Megaespores

Genus Species Stratigraphic position Material Notes Images

Rogalskaisporites[6][106]

  • Rogalskaisporites cicatricosus
  • Blanowice Coals
  • Spores

Affinities with Sphagnopsida inside Sphagnales. Pollen nearly identical to that one found associated with the modern moss genus Sphagnum. Moss related to high humid environments.

Bacutriletes[107][108]

  • Bacutriletes spicatus
  • Bacutriletes clavatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lycophyta inside Tracheophyta. A relative abundant genus, specially on deltaic deposits. Low herbaceous Lycophyte flora.

Acanthotriletes[109]

  • Acanthotriletes levidensis
  • Acanthotriletes minor
  • Acanthotriletes osmundiformis
  • Acanthotriletes rarispinosus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lycophyta inside Tracheophyta. This Spores, rather abundant are correlated with humid settings, and come from mostly herbaceous-arbustive plants.

Hughesisporites[109]

  • Hughesisporites pustulatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lycophyta inside Tracheophyta. Spores from low herbaceous flora, linked mostly to humid environments with abundant freshwater.

Eehitriletes[107][108][110][111][112]

  • Eehitriletes hispidus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lycophyta inside Tracheophyta. Low herbaceous flora from humid envrironments

Hughesisporites[107][108][110][111][112]

  • Hughesisporites stillarus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lycophyta inside Tracheophyta. Abundant on humid strata, mostly related with deltaic facies.

Aneuletes[107][108][110][111][112]

  • Aneuletes patera
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lycophyta inside Tracheophyta. Rather rare than other similar genera, found mostly on deltaic facies.

Trachytriletes[109]

  • Trachytriletes dichotomus
  • Trachytriletes filiciformis
  • Trachytriletes lineatus
  • Trachytriletes rugosus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lycopodiaceae inside Lycopodiopsida. Represents herbaceous Lycophytes of small to medium size (10–40 cm), that are found mostly on deltaic deposits.

Lycopodiumsporites[109]

  • Lycopodiumsporites gristhorpensis
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lycopodiaceae inside Lycopodiopsida. Resemble spores of the modern genus Lycopodium. If it belongs to a similar genus, represent low herbaceous flora spores.

Reticulatisporites[109]

  • Reticulatisporites triangulus
  • Reticulatisporites flavus
  • Reticulatisporites pallidus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lycopodiaceae inside Lycopodiopsida. Resembles the modern genera Lycopodiella and Huperzia, characteristic small to medium (20–90 cm) herbaceous Lycopites, found on environments with abundant water.

Biharisporites[107][108][110][111][112]

  • Biharisporites scaber
  • Biharisporites scabrus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Lycopodiopsida inside Lycophyta. Lycophyte spores of rather uncertain affinities, are more common on fluvial deposits.

Paxillitriletes[113]

  • Paxillitriletes phyllicus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Isoetales inside Lycophyta. Spores related with modern Isoetes, representing Small plants related with water bodies.[114] It comprises the main Megaspore zonation of the Toarcian of Poland, being the most abundant spore found.

Minerisporites[113]

  • Minerisporites institus
  • Minerisporites volucris
  • Minerisporites richardsoni[107]
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Isoetaceae inside Lycopsida. Related to plant similar to Isoetes lacustris, present on flooded basins, and other ecosystems with relative abundant water supply.[115]

Echitriletes[116][110]

  • Echitriletes hispidus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Isoetaceae inside Lycopsida.

Erlansonisporites[113][107]

  • Erlansonisporites sparassis
  • Erlansonisporites excavatus
  • Erlansonisporites retieulatus
  • Erlansonisporites sparassis
  • Erlansonisporites reticulatus
  • Erlansonisporites tegimentus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the genus Selaginella inside Selaginellaceae. Herbaceous Moss.[117] Relatively abundant Pollen Genera

Horstisporites[107]

  • Horstisporites areolatus
  • Horstisporites planatus
  • Horstisporites puteus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Selaginellaceae inside Lycopsida. Less abundant than othe Lycophite Spores, its abundance increases near Deltaic deposits.

Trileites[107][110]

  • Trileites turbanaeformis
  • Trileites murray
  • Trileites calvus
  • Trileites candoris
  • Trileites pyramidalis
  • Trileites solitus
  • Trileites usitatus
  • Triletes reticulatus
  • Triletes datura
  • Triletes litchi
  • Triletes phyllicus
  • Triletes retiarius
  • Triletes richardsoni
  • Triletes sparassis
  • Triletes utilis
  • Triletes wicheri
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Selaginellaceae inside Lycopsida. Among the most abundant genera of Pollen present on the Formation and the south of Fennoscandia

Valvisisporites[110]

  • Valvisisporites auritus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Selaginellaceae inside Lycopsida. This herbaceous Lycophyte flora appears more on the southern boreholes, linked to large wood debris.

Maexisporites[110]

  • Maexisporites planatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Selaginellaceae inside Lycopsida. Herbaceous flora spores linked to deltaic facies.

Striatriletes[110]

  • Striatriletes reticulatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Selaginellaceae inside Lycopsida.

Foveosporites[109]

  • Foveosporites canalis
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Selaginellaceae inside Lycopsida. Less abundant than other similar spores, found on a few samples on all the boreholes.

Klukisporites[109]

  • Klukisporites pseudoreticulatus
  • Klukisporites variegatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Selaginellaceae inside Lycopsida. Represents large spores from herbaceous lycophytes, that are more abundant on the south of the Toarcian polish basin than on the north.

Nathorstisporites[107][108][110]

  • Nathorstisporites hopliticus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Pleuromeiales inside Lycophyta. The Plueromeiales where tall Lycophites (2 to 6 m) common on the Trassic. Probably come from a relict genus.

Carnisporites[107][108][110][111][112]

  • Carnisporites granulatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Cynepteridaceae inside Schizaeaceae. Related to Ferns similar to Cynepteris, from the late triassic of North America.

Reticulatisporites[106]

  • Reticulatisporites arcuatus
  • Reticulatisporites ornatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with Lygodiaceae inside Schizaeales. Spores nearly identical to that one found associated with the modern moss genus Lygodium. Arboreal moss related to high humid environments, being major fuel for peat fires.

Perinopollenites[109]

  • Perinopollenites elatoides
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with Gymnospermopsida inside Tracheophyta. Pollen that resemble those of paleozoic genera such as Gangamopteris, with arboreal built.

Chasmatosporites[109]

  • Chasmatosporites apertus
  • Chasmatosporites major
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with Gymnospermopsida inside Tracheophyta. Pollen that resemble those of paleozoic genera such as Gangamopteris, with arboreal built.

Corrugatisporites[107][108][110][111][112]

  • Corrugatisporites amplectiformis
  • Corrugatisporites scanicus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Filicopsida inside Monilophyta. Fern spores of uncertain placement

Ischyosporites[107][108][110][111][112]

  • Ischyosporites variegatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Polypodiidae inside Filicopsida. Fern spores of uncertain placement.

Leiotriletes[107][108][110][111][112]

  • Leiotriletes mecklenburgensis
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Polypodiidae inside Filicopsida. Fern spores of uncertain placement

Osmundacidites[6]

  • Osmundacidites wellmanii
  • Blanowice Coals

Spores

Affinities with Osmundaceae inside Pteridophyta. Spores nearly identical to that one found associated with the modern fern genus Osmunda. Members of the genus Osmunda have been found on coeval age strata on Sweden.

Crybelosporites[107][108][110][111][112]

  • Crybelosporites stylosus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Marsileaceae inside Salviniales. Represents spore from fully acuatic ferns, found associated with fluvial or deltaic deposits, where probably formed large underwater colonies.

Gleicheniidites[107][108][110][111][112]

  • Gleicheniidites carinatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Gleicheniaceae inside Polypodiidae. Resemble the modern Gleichenia Spores, and proably represent a similar genus or a member of it. Fern related to large colonies, found mostly on humid environments.

Marattisporites[107][108][110][111][112]

  • Marattisporites scabratus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Marattiaceae inside Polypodiidae. Resemble the modern Marattia spores, probably belonging to a similar genus, related with large sized herbaceous ferns of humid environments.

Camptotriletes[109]

  • Camptotriletes labirynthus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Marattiaceae inside Polypodiidae.

Matonisporites[107][108][110][111][112]

  • Matonisporites phlebopteroides
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores

Affinities with the Matoniaceae inside Polypodiidae. The so-called comb-ferns, found forming large colonies on humid settings.

Cyathidites[6][106]

  • Cyathidites minor
  • Blanowice Coals

Spores

Affinities with Cyatheaceae inside Cyatheales. Cyathidites minor almost certainly belong to well known Mesozoic species Coniopteris hymenophylloides and to other fossil cyatheaceous or dicksoniaceous ferns such as Eboracia lobifolia and Dicksonia mariopteri.

Cerebropollenites[107][108][110][111][112]

  • Cerebropollenites macroverrucosus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Mechowo borehole
  • Gorzow Wielkopolski borehole
  • Pollen

Affinities with the Gymnospermophyta inside Spermatophyta. Pollen from uncertain Gimnosperm affinities.

Eucommiidites[118]

  • Eucommiidites troedssonii
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with the Cycadales inside Cycadopsida. Originally was trought to come from Angiosperms.

Psophosphaera[109]

  • Psophosphaera coniferoides
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with the Cycadales inside Cycadopsida. Cycad Pollen, coming from arbustive to low arboreal flora.

Chasmatosporites[6][106]

  • Chasmatosporites apertus
  • Blanowice Coals

Pollen

Named originally Pollenites apertus, they resemble pollen grains of the genus Cycas. Later works refer them to modern pollen grains that proper of wrote that resemble ?Cycadopsida (?Cycadales), concretely the genus Encephalartos. Alternatively, can be Pollen from members of Ginkgopsida (?Gnetales).[119]

Bennettistemon[107][108][110][111][112][120]

  • Bennettistemon bursigerum
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with the inside Cycadopsida.[120] Very abundant Pollen.

Pseudowalchia[109]

  • Pseudowalchia biangulina
  • Helenow Beds
  • Pollen

Affinities with Voltziales inside Coniferae. Primitive Conifer and possible relict taxon

Inaperturopollenites[107][108][110][111][112]

  • Inaperturopollenites laevis
  • Inaperturopollenites turbatus
  • Inaperturopollenites elatoides
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with the Pinidae inside Coniferae.

Rotundipollenites[109]

  • Rotundipollenites laevis
  • Rotundipollenites lineatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with Pinaceae inside Coniferae.

Pityosporites[107][108][110][111][112]

  • Pityosporites haploxylon
  • Pityosporites minimus
  • Pityosporites alata
  • Pityosporites alatipollenites
  • Pityosporites silvestris
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with Pinaceae inside Coniferae. Resemble modern Pinus Pollen, probably belonging to a similar Genus.

Quadraeculina[106]

  • Quadraeculina anellaeformis
  • Blanowice Coals
  • Pollen

Affinities with Pinaceae inside Coniferae. Pollen From arbustive to arboreal plants, resembling the modern genus Picea

Tsugapollenites[6][106]

  • Tsugapollenites macroserratus
  • Tsugaepollenites macroverrucosus
  • Blanowice Coals

Pollen

Affinities with Abietoideae inside Coniferae. Pollen From arbustive to arboreal plants, resembling the pollen of the modern genus Tsuga

Sciadopityspollenites[6][106]

  • Sciadopityspollenites serratus
  • Blanowice Coals

Pollen

Affinities with Sciadopityaceae inside Coniferae. This pollen resembles the present on the modern Cupressaceae Sciadopitys

Pseudopodocarpus[107][108][110][111][112]

  • Pseudopodocarpus sp.
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with Podocarpaceae inside Pinopsida. Pollen From arbustive to arboreal plants

Pristinuspollenites[6][106]

  • Pristinuspollenites sulcatus
  • Blanowice Coals

Pollen

Affinities with Podocarpaceae inside Pinopsida. Pollen From arbustive to arboreal plants

Classopollis[107][108][110][111][112]

  • Classopollis torosus
  • Classopollis classoides
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with the Cheirolepidiaceae inside Coniferae. Is very abundant on hot and dry settings, found specially on the Brody-Lubienia Borehole.

Cupressacites[107][108][110][111][112]

  • Cupressacites subgranulatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with the Cupressaceae inside Coniferae. The Pollen from this genus is similar to the present on the modern Fitzroya.

Applanopsipollenites[107][108][110][111][112]

  • Applanopsipollenites dampieri
  • Applanopsipollenites trilobatus
  • Applanopsipollenites segmentatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with the Araucariaceae inside Coniferae. Resemble the pollen from the modern genus Agathis.

Araucariacites[6][106]

  • Araucariacites australis
  • Blanowice Coals

Pollen

Affinities with Araucariaceae inside Pinopsida. Pollen From arbustive to arboreal plants

Tricolpites[116]

  • Tricolpites troedssonii
  • Tricolpites gorzowiensis
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with Angiospermae and probably Magnoliophyta. Possible pollen from primigenial Flowering plants or relatives

Clavatipollenites[109]

  • Clavatipollenites hughesii
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Pollen

Affinities with Angiospermae and probably Magnoliophyta. Possible pollen from primigenial Flowering plants or relatives

Megaflora

The Lublin Upland fluvial sandstones contain diverse types of fossil flora, associated genera and species only with Lower Jurassic sediments. Originally, while studying the Carboniferous flora from the boreholes in the area of the planned "Bogdanka" mine, appeared typical flora in similar to Jurassic formations.[121] The age of the plant material was not determined concretely until 2020, where was recovered as being Lower Toarcian in age (with some parts probably recovering Latest Pliensbachian strata).[122] Lublin lias is dominated by cycads and Bennetites Ginkgoales and pinnate, ferns occur sporadically, all on a conglomerate with numerous species occurs in the bottom of the Toarcian, where the deposits are filled with of coal, mudstone, sandstone and clay siderite (reworked from the Carboniferous), as well as pebbles from Devonian limestones.[123] Due to the fact that similar Boreholes and deposits nearby show similar characters it should be assumed that we have to dealing with the end of a river, that eroded the nearest Devonian-Carboniferous deposits (At the northeast), being brought by flowing waters and deposited in the aquatic environment-inland.[123] Vegetation mostly grew outside the sedimentation area, as well as on shores and shallows.[124]

Genus Species Stratigraphic position Material Notes Images

Equisetites[124][125]

  • Equisetites beani
  • Lublin Coal Basin

Stems

Affinities with the Equisetales inside Equisetopsida. Very rare. Local Equisetales obtained a considerable size, comparable to modern subtropical bamboos, close to lakes and in the wettest environments

Pachypteris[121][124][125]

  • Pachypteris lanceolata
  • Pachypteris rhomboidalis
  • Lublin Coal Basin

Pinnae

Affinities with the Peltaspermales inside Pteridospermatophyta. Bipinnate leaves, rachis longitudinally striated, with a long petiole and secondary rachises inserted oppositely to suboppositely on the upper side of the primary rachis. This Leaves belong to large Aboreal Ferns related with dry environments.

Stenopteris[121][124][125]

  • Stenopteris (Rhaphidopteris?) duetschii
  • Lublin Coal Basin

Pinnae

Affinities with the Umkomasiaceae inside Pteridospermatophyta. Is very similar to Rhaphidopteris, characterized by usually bi-tripinnate, coriaceous leaves with narrow segments. It belongs to Large (up to 25 m tall) arboreal Fern-like plants.

Caytonia[121][125]

  • Caytonia sp
  • Lublin Coal Basin

Ovulate Structure

Affinities with the Caytoniaceae inside Caytoniales. Represents the Ovulate organs or large Tree ferns, and is related with the middle Jurassic flora of United Kingdom.

Sagenopteris[124][125]

  • Sagenopteris sp
  • Borehole L-95 near Łęczna (Lublin Coal Basin)

Pinnae

Affinities with the Caytoniaceae inside Caytoniales. Sagenopteris is most likely the Pinnae of the Tree Fern that also contain Caytonia.

Anomozamites[124][125]

  • Anomozamites cf. gracilis
  • Borehole L-95 near Łęczna (Lublin Coal Basin)

Pinnae

Affinities with the Bennettitales inside Bennettitopsida. Anomozamites is characterised by slender, (almost) completely and regularly segmented leaves whose leaflets are generally as long as broad or, at maximum, two times as long as broad. This genus is related with more arboreal Bennetitalean flora. Shows coriaceous leaves and is a genus linked more with dry climates.

Otozamites[124][125]

  • Otozamites beani
  • Otozamites graphicus
  • Borehole L-95 near Łęczna (Lublin Coal Basin)

Pinnae

Affinities with the Bennettitales inside Bennettitopsida. A more arbustive type of Bennetite, abundant on the Pliensbachian-Toarcian strata along Eurasia, related to rather dry climates.

Nilssoniopteris[121][124][125]

  • Nilssoniopteris major
  • Nilssoniopteris vittata
  • Borehole L-95 near Łęczna (Lublin Coal Basin)

Pinnae

Affinities with the Bennettitales inside Bennettitopsida. This Bennetite has a leaf similar to Nilssonia, large, lanceolate in outline with coriaceous structure, like the modern angiosperm Philodendron. Is related with rather Dry-hot environments, with less Fern flora.

Nilssonia[121][124][125]

  • Nilssonia compta
  • Nilssonia orientalis
  • Nilssonia polymorpha
  • Nilssonia sp.
  • Borehole L-95 near Łęczna (Lublin Coal Basin)

Pinnae

Affinities with the Cycadeoidaceae inside Bennettitales. The most abundant plant macrofossil locally, and the most diverse found on the Lublin coals. It represents a Cycad Like plant with Coriaceous leaves. Is related by some botanists with the Bennetite branch that led to Angiosperms.

Weltrichia[121][125]

  • Weltrichia spectabilis
  • Lublin Coal Basin

Bennetite "Flower"

Affinities with the Cycadeoidaceae inside Bennettitales. The Bennetite flowers are the main organ that links this relatives with the Cycas with modern Angiosperms. Probably the Pollen assigned to Angiosperms come from plants with this "Flowers".

Pterophyllum[121][124][125]

  • Pterophyllum sp.
  • Borehole L-95 near Łęczna (Lublin Coal Basin)

Pinnae

Affinities with the Williamsoniaceae inside Bennettitopsida. Leaves from Arboreal Bennetites, similar to the modern Cyca Encephalartos woodii, with robust Trunks, built for Dry and hot climates.

Baiera[124][125]

  • Baiera digitata
  • Borehole L-95 near Łęczna (Lublin Coal Basin)

Leaves

Affinities with the Ginkgoaceae inside Ginkgoidae. Linked to the Hettangian-Sinemurian flora from Greenland and Skane, but also with coeval flora from the Sorthat Formation. Is the main Tree flora recovered locally.

Hirmeriella[124][125]

  • Hirmeriella sp.
  • Borehole L-95 near Łęczna (Lublin Coal Basin)

Leaves

Affinities with the Cheirolepidiaceae inside Pinales. The type Leave from the Cheirolepidaceae family (as Cheirolepis, the genus that give name to the family, is a junior synonym), appears to be linked with Hot climates, able to survive in dry, extreme conditions, and been fire tolerant.

Footnotes

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