Esox

Esox is a genus of freshwater fish, the only living genus in the family Esocidae—the esocids which were endemic to North America and Eurasia during the Paleogene through present.[2]

Esox Temporal range: Upper Cretaceous–recent[1] PreꞒ Ꞓ O S D C P T J K Pg N
Northern pike (E. lucius)
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Esociformes
Family:	Esocidae G. Cuvier, 1817
Genus:	Esox Linnaeus, 1758
Type species
Esox lucius Linnaeus, 1758

The species of this genus are known as pike and pickerel.[3][4] The type species is E. lucius, the northern pike.

The big pike species are native to the Palearctic and Nearctic ecozones, ranging across northern North America and from Western Europe to Siberia in Eurasia.

Pikes have the elongated, torpedo-like form of predatory fishes, with sharply pointed heads and sharp teeth. Their coloration is typically grey-green with a mottled or spotted appearance with stripes along their backs, providing camouflage among weeds. Individual pike marking patterns are unique, like fingerprints. Pike can grow to a maximum recorded length of 1.83 m (6 ft), reaching a maximum recorded weight of 35 kg (77 lb).

Species

Currently, seven recognized species are placed in this genus:

Image	Scientific name	Common Name	Subspecies	Distribution
	Esox aquitanicus Denys, Dettai, Persat, Hautecœur & Keith, 2014	Aquitanian pike[5]		Charente to the Adour drainages in France
	Esox americanus J. F. Gmelin, 1789	American pickerel	Esox americanus americanus J. F. Gmelin, 1789 (Redfin pickerel) Esox americanus vermiculatus Lesueur, 1846 (Grass pickerel)	the Saint Lawrence drainage in Quebec down to the Gulf Coast, from Mississippi to Florida,the Great Lakes Basin, from Ontario to Michigan, down to the western Gulf Coast, from eastern Texas to Mississippi.
	Esox cisalpinus Bianco & Delmastro, 2011	Southern pike[6]		central and northern Italy, southeastern France and Switzerland, and it might also occur in western Balkan.
	Esox lucius Linnaeus, 1758	Northern pike		Britain, Ireland, northern Europe, most of Canada, and most parts of the United States
	Esox masquinongy Mitchill, 1824	Muskellunge	Esox masquinongy masquinongy (Great Lakes muskellunge or Spotted muskellunge ) Esox masquinongy ohioensis (Chautauqua muskellunge or Barred muskellunge) Esox masquinongy immaculatus (Clear muskellunge)	mesotrophic lakes and large rivers from northern Michigan, northern Wisconsin, and northern Minnesota through the Great Lakes region, north into Canada
	Esox niger Lesueur, 1818	Chain pickerel		southern Canada to Florida, and west to Texas. On the Atlantic Coast, in Maine, New Hampshire, New Brunswick, and Nova Scotia
	Esox reichertii Dybowski, 1869	Amur pike		the Amur River system in east Asia

Hybrids between Esox masquinongy and Esox lucius are well-known and referred to as the tiger muskellunge.

Fossil species

One fossil species, Esox kronneri, is known from the Eocene of the Green River formation,[7] and a second, Esox nogaicus, is known from the Pleistocene of the Ukraine. Additional species are found in Miocene (Esox sibiricus) and Pliocene (Esox moldavicus) deposits from the Ukraine, Poland, Kazakhstan, Mongolia, and Moldavia.[8] Two additional fossil species, both from the Cretaceous of Alberta, Canada, are placed in their own genera: Estesesox foxi[9] (Santonian to Campanian[10]), and Oldmanesox canadensis[9][7] (Campanian to Maastrichtian[11]).

Etymology

The generic name Esox (pike fish) derives from the Greek ἴσοξ (ee-soks, a large fish) and appears to be both profoundly archaic and cognate with Celtic, Welsh eog and Irish Gaelic iasc (fish), as well as alpine Gaulic *esosk which is consistent with the original indoeuropean root for the common word for fish, *pei(k)sk. Pliny uses the Latin form Esox in reference to a large fish in the Rhine normally identified with Salmonidae (lax or salmon). Carolus Linnæus attributes Esox to the pike fish which is of similar form and appearance but taxonomically different from the salmoniformes, whereas the first mention of Esox as a marine animal appears in the writings of Hesych(ius).

A young E. lucius specimen—a "Chain pickerel" in the original sense—in an aquarium.

The English common name "pike" is an apparent shortening of "pike-fish", in reference to its pointed head, Old English píc originally referring to a pickaxe. The plural of pike is pike.

A northern English and Lowland Scots name for the pike, ged, similarly derives from Old Norse gaddr (spike) (cf. the modern Swedish name for the pike, gädda, the Danish "gedde", the Norwegian "gjedde" and Scottish Gaelic: geadais). The Dutch name for the pike (snoek) has been given to a wide variety of fish reminding sailors of the pike (see snoek, snook).

The English "pike" originally referred specifically to the adult fish, the diminutive form "pickerel" (now used to name some of the smaller pike, E. americanus and E. niger) referring to the young. The walleye (Sander vitreus) is sometimes called a pickerel, but it is unrelated to the pike, being a member of the perch family (Percidae). Pike are not to be confused with the unrelated pikeminnows (traditionally known as squawfish) of genus Ptychocheilus (family Cyprinidae) or pikeperch (Sander lucioperca) which is more akin to walleye than to pike. Pike are also called "jackfish" in North America and informally "slough shark" in Western Canada.

Diet

Pike in Haus des Meeres, Vienna

Pike feed on a wide range of food sources, predominantly smaller shoal fish. Pike are also cannibalistic, sometimes preying upon smaller members of their own species. This can be seen clearly in the northern pike.

They will also prey on insects and amphibians such as newts or frogs in times when their usual food is scarce, and occasionally on small mammals like moles or mice when caught water-borne. Small birds such as ducklings may become a target for hungry pike. Pike are also known to prey on swimming snakes.

They are, however, undeserving of their reputation for being overly vicious predators. There are few substantiated incidents of pike "attacks" on people. Pike's further reputation as a pest seems to lie predominantly amongst a small handful of anglers and fishery managers who think, perhaps unfairly, that pike are a threat to native rough fish and also other sport fish.

Angling

Sign on the River Shannon, Ireland

A caught pike

Effective methods for catching this hard-fighting fish include dead baits, live baits, and lure fishing. Pike can easily be damaged when handled since they are not as robust as their reputation would suggest. Colour of lure can be influenced by water clarity and weather conditions. Since pike have numerous sharp teeth it is wise to take extreme care when unhooking them. The use of a wet leather gauntlet and surgical forceps to remove hooks is highly recommended on safety grounds. If practicing catch and release fishing, care for the pike should be the pike angler's utmost concern. The formerly recommended practice of grasping a pike by its eye sockets (misinterpreted as "its eyes") resulted in numerous released pike that quickly died from inability to see prey any longer.

The current recommended method of grasping pike is to close the hand firmly over the gill covers, and to make the period of handling as short as possible before release. Grabbing a pike by the gill covers is not feasible when a pike is very big, but it is easy to handle a pike by inserting the fingers at the bottom of the gill opening and grabbing the lower jaw. Big pike should also be supported at the belly. When a pike is held this way it is also easier to keep the mouth open to remove a hook. Some anglers now use special grips to grab the pike's front lower jaw, which can add to the safety of an anglers because of the danger imposed by the hooks of the lure or tackle and the pike's teeth. However these can cause serious damage to a pike's lower jaw. The Pike Anglers Club was formed in 1977 to campaign for the preservation of pike and the sport of pike fishing.

Pike are susceptible to gut hooking when fished for with natural bait. Upon taking the bait, the pike will hold it for a short time in its mouth as it moves off. The pike will then, usually, turn the bait in its mouth, so that it sits in alignment with its throat to ease swallowing. It is recommended that when pike fishing the process is not allowed to go this far and a strike is recommended as soon as a bite is indicated. Otherwise, what is known as gut hooking will result, which will normally kill or seriously injure the fish. Dutch research shows that cutting the line immediately when the fish is gut hooked will still give low mortality (14%). The hooks in the gut or stomach were either encapsulated or removed from the body.[12] Placing hooks near the rear of the bait reduces the risk of deep hooking.[13]

Other methods of catching and handing pike that are now frowned upon are the gaff and the gag. The gaff is a metal hook on the end of a pole used to hook through the fish's body in place of a more humane landing net. A gag is a device for holding open the pike's mouth whilst unhooking. These are now illegal in Scotland, as they put a huge amount of pressure on a pike's jaw, thus causing irreparable damage.

Cuisine

The taste of pike and pickerel is highly esteemed, but the "multitude of long, fine, forked bones" are problematic.[14] [15] The dish of quenelles de brochet (pike dumplings), which puts the meat through a sieve, was invented to deal with this.[16] Indeed, Escoffier believed, falsely, that quenelles had completely displaced the whole fish from the menu.[14]

In mythology

Fossil Esox

Russian mythology holds that the pike is one of several forms assumed by evil water spirits called vodyanoy, and a ravenous mythical pike is traditionally blamed for decimating the fish population in the Sheksna River. Russian fairy tales, on the other hand, also tell about an old wise pike that can fulfil wishes of the one who catches it, if its catcher releases it back into its habitat.[17]

In the Finnish Kalevala, Väinämöinen creates a kantele (string instrument) from the jawbone of a pike.

Submarines and tanks

Two United States Navy submarines have been named Pike – SS-6 of 1903 and SS-173 of 1935 – and three – SS-22 of 1912, SS-177 of 1936, and SS-524 of 1944 – named Pickerel. In addition, the Soviet submarines known to NATO as the Victor III class and Akula class are called the Shchuka (Щука, "pike") class in Russian. The Soviet Iosif Stalin tank (IS-3) was also nicknamed Shchuka, in reference to its sharply pointed hull front.

Heraldry

In heraldry, the pike is called a lucy (English heraldry) or a ged (Scottish heraldry).[3] It is usually blazoned either naiant (swimming), embowed (bowed) or hauriant (jumping), though pairs of lucies may appear addorsed (back to back), as in the arms of the Finnish town of Uusikaupunki (Argent, two lucies addorsed azure).

Uusikaupunki coat of arms

The canting arms of Lucie de Cockermouth: Gules, three lucies in pale argent, (2 and 1).

Coat of arms of Gimte, in Lower Saxony, Germany.

References

1. Froese, Rainer, and Daniel Pauly, eds. (2008). "Esocidae" in FishBase. December 2008 version.
2. PaleoBiology Database: Esox, basic info
3. Fox-Davies, Arthur Charles (1909). A Complete Guide to Heraldry. London: T.C. & E.C. Jack. p. 255. LCCN 09023803.
4. Woodward, John (1892). A treatise on heraldry, British and foreign. Edinburgh: W. & A.K. Johnston. p. 694. LCCN 02020303. Archived from the original on 2 November 2007. Retrieved 29 May 2009.
5. Denys, Gaël Pierre Julien; Dettai, Agnès; Persat, Henri; Hautecœur, Mélyne; Keith, Philippe (2014). "Morphological and Molecular Evidence of Three Species of Pikes Esox spp. (Actinopterygii, Esocidae) in France, including the Description of A New Species". Comptes Rendus Biologies. 337 (9): 521–34. doi:10.1016/j.crvi.2014.07.002. PMID 25242691.
6. Lucentini, Puletti, Ricciolini, Gigliarelli, Fontaneto, Lanfaloni, Bilò, Natali, Panara (2011). Molecular and Phenotypic Evidence of a New Species of Genus Esox (Esocidae, Esociformes, Actinopterygii): The Southern Pike, Esox flaviae. PLoS ONE 6(12): e25218. doi:10.1371/journal.pone.0025218
7. Grande, L. (1999). "The First Esox (Esocidae: Teleostei) from the Eocene Green River Formation, and a Brief Review of Esocid Fishes". Journal of Vertebrate Paleontology. 19 (2): 271–292. doi:10.1080/02724634.1999.10011141.
8. Kovalchuk, Oleksandr M.; Wilson, Mark V.H.; Grande, Terry (2017). "A review of Neogene and Quaternary pikes of southeastern Europe and a new species from the early Pleistocene of Nogaisk, Ukraine". Acta Palaeontologica Polonica. 62. doi:10.4202/app.00311.2016.
9. Wilson, Brinkman & Neuman, 1992
10. "Fossilworks: Estesesox".
11. "Fossilworks: Oldmanesox".
12. "Archived copy" (PDF). Archived from the original (PDF) on 27 March 2009. Retrieved 7 February 2009.CS1 maint: archived copy as title (link)
13. "Archived copy". Archived from the original on 13 September 2013. Retrieved 10 October 2013.CS1 maint: archived copy as title (link)
14. Waverley Root, Food, 1996, p. 353
15. 'Piscator', "Pike Fishing", The Sportsman (Second Series), 2:3:139 (March 1840)
16. Marthe Daudet, Shirley King, translator and adaptor, Pampille's Table: Recipes and Writings from the French Countryside from Marthe Daudet's Les Bons Plats de France [1934], p. 153
17. "Emelya the Simpleton", russian folklore fairy tale. At the Pike's Behest

External links

• "Esox". Integrated Taxonomic Information System. Retrieved 5 December 2004.
• Oxford English Dictionary, s.vv. "Esox", "Ged¹", and "Pike, n.⁴".
• Pike in Your Waters, the Pike Anglers Club, pub 2003
• Pike Preservation - Fishing Equipment, Pike Fishing Scotland
• Float Tube Fishing In Ireland Pike Fishing from a Float Tube
• Pike Fishing in Denmark Coarse Fishing in Denmark
• Pike Fishing in Scotland Coarse Fishing in the Highlands
• River Swimming Water Safety mentions Pike attack as a risk of open water swimming. Accessed 21 October 2007