Sanjuansaurus

Sanjuansaurus ("San Juan Province lizard") is a genus of herrerasaurid dinosaur from the Late Triassic (Carnian) Cancha de Bochas and La Peña Members of the Ischigualasto Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina.

Sanjuansaurus Temporal range: Carnian ~231.4 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Holotype
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	Saurischia
Family:	†Herrerasauridae
Genus:	†Sanjuansaurus Alcober & Martinez, 2010
Species:	†S. gordilloi
Binomial name
†Sanjuansaurus gordilloi Alcober & Martinez, 2010

Description and classification

Restoration

Sanjuansaurus was comparable in size to a medium-sized Herrerasaurus, with a thigh bone that was 395 millimetres (15.6 in) long and a tibia that is 260 millimetres (10 in) in length. Alcober and Martinez performed a phylogenetic analysis and found Sanjuansaurus to be a herrerasaurid.[1] It was determined that Sanjuansaurus and Herrerasaurus share many similarities in the morphology of the skull, neck vertebrae, back vertebrae, hip vertebrae, scapula, and the hip bones. Alcober and Martinez observed that Sanjuansaurus and Staurikosaurus share many similarities in the morphology of the hip bones, and the tibia.[1] The pubis of Sanjuansaurus, unlike in other herrerasaurids, points toward the cranium.

Distinguishing anatomical features

A diagnosis is a statement of the anatomical features of an organism (or group) that collectively distinguish it from all other organisms. Some, but not all, of the features in a diagnosis are also autapomorphies. An autapomorphy is a distinctive anatomical feature that is unique to a given organism or group.

According to Alcober and Martinez (2010), Sanjuansaurus can be distinguished based on the following characteristics:[1]

• the cervical vertebrae have shelf-like, posterolaterally directed transverse processes
• the neural spines of the sixth to eighth dorsal vertebrae, at least, bear acute anterior and posterior processes
• the glenoid has everted lateral margins
• the pubis is relatively short, measuring 63% of the length of the femur
• a pronounced, rugose scar is present on the medial surface of the femur at the level of the fourth trochanter

Discovery

Maxilla of PVSJ 605

Sanjuansaurus was named and described in 2010 by Oscar Alcober and Ricardo Martinez. The type species was named S. gordilloi after Raul Gordillo, the head fossil preparator and artist of the San Juan Museum. It is known from and based on an associated and partially articulated partial skeleton (PVSJ 605) consisting of a jaw fragment, most of the vertebral column from the axis to the anterior tail, the shoulder blades, an ulna, part of the pelvis, most of the long bones of the legs, and a few other bones.

Paleoecology

Provenance and occurrence

Sanjuansaurus holotype specimen PVSJ 605 was discovered in 1994, in gray-green sandstone of the Cancha de Bocas Member near the base of the Ischigualasto Formation, in Ischigualasto Provincial Park in San Juan, Argentina. This horizon dates to approximately 231.4 Ma, during the late Carnian Stage of the Late Triassic.[1]

Fauna and habitat

Skeletal restoration

In the Ischigualasto Formation, dinosaurs constituted only about 6% of the total number of fossils,[2] but by the end of the Triassic Period, dinosaurs were becoming the dominant large land animals, and the other archosaurs and synapsids declined in variety and number.[3]

Studies suggest that the paleoenvironment of the Ischigualasto Formation was a volcanically active floodplain covered by forests and subject to strong seasonal rainfalls. The climate was moist and warm,[4] though subject to seasonal variations.[5] Vegetation consisted of ferns (Cladophlebis), horsetails, and giant conifers (Protojuniperoxylon). These plants formed highland forests along the banks of rivers.[6] Sanjuansaurus lived in the jungles of Late Triassic South America alongside early dinosaurs, Eoraptor, Herrerasaurus, Chromogisaurus, and Panphagia, as well as Saurosuchus,[7] a giant land-living rauisuchian (a quadrupedal meat eater with a theropod-like skull); the broadly similar but smaller Venaticosuchus, an ornithosuchid; and the predatory chiniquodontids. Herbivores were much more abundant than carnivores and were represented by rhynchosaurs such as Hyperodapedon (a beaked reptile); aetosaurs (spiny armored reptiles); kannemeyeriid dicynodonts (stocky, front-heavy beaked quadrupedal animals) such as Ischigualastia; and therapsid traversodontids (somewhat similar in overall form to dicynodonts, but lacking beaks) such as Exaeretodon. These non-dinosaurian herbivores were much more abundant than early ornithischian dinosaurs like Pisanosaurus.[8]

References

1. Alcober, Oscar A.; Martinez, Ricardo N. (2010). "A new herrerasaurid (Dinosauria, Saurischia) from the Upper Triassic Ischigualasto Formation of northwestern Argentina". ZooKeys. 63: 55–81. doi:10.3897/zookeys.63.550. PMC 3088398. PMID 21594020.
2. Benton, Michael J. (1999). "Origin and early evolution of dinosaurs". In Farlow, James O.; Brett-Surman, M.K. (eds.). The Complete Dinosaur. Indiana University Press. pp. 204–215. ISBN 0-253-21313-4.
3. Parrish, J. Michael (1999). "Evolution of the archosaurs". In Farlow, James O.; Brett-Surman, M.K. (eds.). The Complete Dinosaur. Indiana University Press. pp. 191–203. ISBN 0-253-21313-4.
4. Tucker, Maurice E.; Benton, Michael J. (1982). "Triassic environments, climates, and reptile evolution" (PDF). Palaeogeography, Palaeoclimatology, Palaeoecology. 40 (4): 361–379. doi:10.1016/0031-0182(82)90034-7. Retrieved 2009-07-23.
5. Columbi, Carina E. (2008-10-05). Stable isotope analysis of fossil plants from the Upper Triassic Ischigualasto Formation in the northwest of Argentina. Houston, TX: The Geological Society of America. Retrieved 2009-07-23.
6. Sereno, P.C. & Novas, F.E. (1992). The complete Skull and Skeleton of an Early dinosaur. Science. 258: 1137-1140. [Herrerasaurus ischigualastensis]
7. Sill, W.D. (1974). "The anatomy of Saurosuchus galilei and the relationships of the rauisuchid thecodonts". Bulletin of the Museum of Comparative Zoology. 146: 317–362.
8. Bonaparte, J.F. (1970). "Annotated list of the South American Triassic tetrapods". Gondwana Symposium Proceedings and Papers. 2: 665–682.