Pancheloniidae

Pancheloniidae is a clade of sea turtles[1] It is defined as all turtles more closely related to cheloniid sea turtles than to dermochelyid ("leatherback") sea turtles.[2]

Pancheloniids Temporal range: Late Cretaceous-Holocene, 73–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Ctenochelys stenoporus skeletons
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Clade:	Panchelonioidea
Clade:	Pancheloniidae
Subgroups
See text.

Genera

The following genera are placed here:

• †Argillochelys[3]
• †Catapleura[4]
• †Ctenochelyidae[4][5]
• †Erquelinnesia[4]
• †Glyptochelone[4]
• †Itilochelys[6]
• †Lytoloma[4]
• †Osteopygis[4]
• †Pampaemys[4]
• †Peritresius[4]
• †Porthochelys[4]
• †Prionochelys[4]
• †Procolpochelys[7]
• †Retechelys[4]
• †Thinochelys[4]

Historic taxon placement

The following list of non-cheloniid pancheloniid species was published by Hirayama and Tong in 2003[4]

• †Allopleuron hoffmani[4], later assigned to Cheloniidae
• †Catapleura arcansaw[4]
• †Catapleura repanda[4]
• †Ctenochelys stenoporus[4]
• †Erquelinnesia gosseleti[4]
• †Glyptochelone suyckerbuykii[4]
• †Lytoloma cantabrigiens[4]
• †Osteopygis emarginatus[4]
• †Pampaemys meridionalis[4]
• †Peritresius ornatus[4]
• †Porthochelys laticeps[4]
• †Prionochelys nauta[4]
• †Retechelys platyops[4]
• †Tasbacka aldabergeni[4], later assigned to Cheloniidae
• †Thinochelys lapisossea[4]
• †Toxochelys latiremis[4], later assigned to clade Durocryptodira.[8]

Cladogram

Below is a cladogram showing the phylogenetic relationships of living and extinct sea turtles in the clade Pancheloniidae based on Lynch and Parham (2003)[9] and Parham and Pyenson (2010).[10]

Lytoloma fossil

Pancheloniidae	†Toxochelys       †Mexichelys       †Lophochleyinae       †Euclastes       †Argillochelys     †Eochelone         †Erquelinnesla     †Pacifichelys         †Puppigerus     Cheloniidae

References

1. Kear BP, Lee MS (March 2006). "A primitive protostegid from Australia and early sea turtle evolution". Biol. Lett. 2 (1): 116–9. doi:10.1098/rsbl.2005.0406. PMC 1617175. PMID 17148342.
2. Joyce W.G, Parham J.F, Gauthier J.A. Developing a protocol for the conversion of rank-based taxon names to phylogenetically defined clade names, as exemplified by turtles. J. Paleont. 2004;78:989–1013.
3. E.A. Zvonok, I.G. Danilov (2017). "A revision of fossil turtles from the Kiev clays (Ukraine, middle Eocene) with comments on the history of the collection of fossil vertebrates of A.S. Rogovich" (PDF). Proceedings of the Zoological Institute of the Russian Academy of Sciences. 321 (4): 485–516.CS1 maint: uses authors parameter (link)
4. Hirayama, Ren; Tong, Haiyan (2003). "Osteopygis (Testudines: Cheloniidae) from the Lower Tertiary of the Ouled Abdoun phosphate basin, Morocco". Palaeontology. 46 (5): 845–856. doi:10.1111/1475-4983.00322.
5. Andrew D. Gentry (2018). "Prionochelys matutina Zangerl, 1953 (Testudines: Pan-Cheloniidae) from the Late Cretaceous of the United States and the evolution of epithecal ossifications in marine turtles". PeerJ. 6: e5876. doi:10.7717/peerj.5876. PMC 6215699. PMID 30402356.
6. Danilov, I.G.; Averianov, A.O; Yarkov, A.A. (2010). "Itilochelys rasstrigin gen. et sp. nov., a new hard-shelled sea turtle (Cheloniidae sensu lato) from the Lower Palaeocene of Volgograd Province, Russia" (PDF). Proceedings of the Zoological Institute RSA. 314 (1): 24–41.
7. "Pancheloniidae". Fossilworks. Retrieved 19 October 2019.
8. "Toxochelys". FossilWorks. Retrieved 19 October 2019.
9. Lynch, S.C.; Parham, J.F. (2003). "The first report of hard-shelled sea turtles (Cheloniidae sensu lato) from the Miocene of California, including a new species (Euclastes hutchisoni) with unusually plesiomorphic characters" (PDF). PaleoBios. 23 (3): 21–35.
10. James F. Parham; Nicholas D. Pyenson (2010). "New Sea Turtle from the Miocene of Peru and the Iterative Evolution of Feeding Ecomorphologies since the Cretaceous". Journal of Paleontology. 84 (2): 231–247. doi:10.1666/09-077R.1.