List of Neanderthal fossils

This is a list of Neanderthal fossils.

Some important European Neanderthals

Remains of more than 300 European Neanderthals have been found. This is a list of the most notable.

	Name	Age	Cranial capacity (cm^3)	Year discovered	Country	Discovered by	Now located at
	Ehringsdorf skull	150k–120k	1450 [1]	1908–1925	Germany
	Engis 2	Undated	? (child)	1829	Belgium	Philippe-Charles Schmerling	University of Liège
	Gibraltar 1	40k	1200 [1]	1848	Gibraltar	Edmund Flint	Natural History Museum
	Gibraltar 2	Undated	? (child)	1926	Gibraltar	Dorothy Garrod	Natural History Museum
	Krijn	100-40 ka	(Not a full skull)	2001	Netherlands	Luc Anthonis	Rijksmuseum van Oudheden
	La Chapelle-aux-Saints 1	60k	1600[1]	1908	France	L. Bardon, A. Bouyssonie and J. Bouyssonie
	La Ferrassie 1	70k–50k	1641 [1]	1909	France	Louis Capitan and Denis Peyrony	Musée de l'Homme
	Neanderthal 1	40k	1452 [1]	1856	Germany	Kleine Feldhofer Grotte	Rheinisches Landesmuseum Bonn
	Saccopastore 1	250k	1200 [2]	1929	Italy
	Saccopastore 2	250k	1300 [3]	1935	Italy	Alberto Blanc and Henri Breuil
	Altamura Man	170k		1993	Italy

Southwest Asian Neanderthals

As of 2017, this list of Southwest Asian Neanderthals may be considered essentially complete.

List of Southwest Asian Neanderthals

Present-day country (country of discovery)	Site	Principal Neanderthal finds	MNI	Geological age (ka)	Descriptions	Notes
Turkey	Karain	Four teeth	1	—	Senyürek (1949)[4][5] Yalçınkaya (1988)[6][5]
Lebanon	Ksâr 'Akil	K2: Teeth and partial maxilla	1	—	Ewing (1963)[7]	Ewing lost this specimen while transferring Ksar Akil material from Boston College to Fordham University.[8]
Lebanon	El Masloukh	Upper second molar[9]	(1)	—	?	Neanderthal attribution is stratigraphic, not morphological.[10]
Israel	Kebara	KMH1: 7-9 mo. old partial skel. KMH2: Post-cranial adult ♂ Various fragments KMH3: Milk tooth (m^1-r)[11] KMH4: 9 milk teeth, germ of 1 permanent tooth [12][11] KMH5: Child mandibular symphysis fragment, no teeth[11] KMH6: Right maxillary fragment with M^1 and M^2[11] KMH7: Milk tooth (m?-r)[11] KMH8: Milk tooth (m^2-l)[11] KMH9: Foot bone (4th right metatarsal)[11] KMH10: Foot bone (1st toe distal phalanx)[11] KMH11: Right clavicle fragment[11] KMH12: Milk tooth (m^?-r)[11] KMH13: Milk tooth germ (m^1-l)[11] KMH14: Tooth (M2-l)[11] KMH15: Milk tooth (m^1-r)[11] KMH16: Milk tooth (left i1)[11] KMH17: Clavicle fragment[11] (KMH18: Mandibular fragment with tooth [M^2-r])[11] (KMH19: Fragmentary tooth crown [M?-r])[11] (KMH20: Parietal bone fragment)[11] (KMH21: Germ of tooth [M^1-l])[11] (KMH22: Milk tooth [upper c-l])[11] (KMH23: Milk tooth [i2-r])[11] KMH24: Tooth (M^3-l)[11] (KMH25: 3 milk teeth germs [upper c-l, m^1, m^2])[11] (KMH26: Tooth germ [i^2-r])[11] KMH27: Tooth (I^2)[13][13] KMH28: Tooth (I2)[13][13] (KMH29: Milk tooth [i2-l])[11] KMH30: Milk tooth (m^1)[11][13] (KMH31: Tooth [lower c])[13]	21 + (10)	64-59[14][15]	KMH1: Smith et al. (1977)[16] KMH2: Arensburg et al. (1985)[17] KMH5-17, 24-31 : Tillier et al. (2003) [11]	Neanderthal attribution uncertain in KMH18-23, 25, 29, and 31[11][13]
West Bank (Mandatory Palestine)	Shuqba	S-D1: Tooth and cranial frags.[9]	1	—	Keith (1931)[18]
Israel (Mandatory Palestine)	Tabun	T C1: Nearly complete adult ♀ T C2: Toothed mandible missing I1 (♂) Various fragments T E1: Right femur shaft (♂?) T E2: Tooth (M1 or M2, ♀?) T C3: Right femur shaft (♀) T C4: Distal right radius frag. (♀) T C5: Right hamate bone T C6: Right pisiform bone T C7: Distal thumb phalanx T B1: 10-11 year-old maxilla (♂?) with I^2-r, M^2-r T BC2: Four teeth (I^2-l, M^1-l, P^3-r, M^1-r) T B3: One tooth (I2-r) T B4: Four teeth (I^1-l, I^2-l, M1-l, M3-r) T B5: Two teeth (M^2-l, M^2-r) T BC6: Two teeth (I^1-l, M2-l)	15	≈170-90 [19][20][21]	McCown (1936) McCown and Keith (1939)	T C1: Neanderthal attribution is not universally accepted.[22] As of 1975, the whereabouts of T BC2, B3, and BC6 are unknown.[9]^:146
Israel	Ein Qashish	(EQH-2: Third molar) EQH-3: Adult lower limbs	1 + (1)[23]	70-60[23]	Been et al. (2017)[23]	Discovered in 2013, these were the first diagnostically Neanderthal remains in Southwest Asia not found in a cave.[23] EQH-2: 70% posterior probability that Neanderthal attribution is correct.[23]
Israel	Shovakh	(Tooth, M(3)-l[24][Note 1])	(1)	—	S. Binford (1966)[25] Trinkaus (1987)[24]	"[A]lthough within archaic and modern human ranges of variation, this complex occlusal morphology may suggest that it is more likely to have derived from a Neandertal than an early modern human". (Trinkaus 1987)[24]
Israel	Amud	A1: Adult full skeleton ♂ A2: Maxillary fragment A7: 10-mo.-old partial skel.	3[Note 2][26]	61-53[26]	A1: Suzuki et al. (1970)[27] A7: Rak et al. (1994)[28]
Syria	Dederiyeh	D1: 19-30-month-old full skel. D2: 21-30-month-old full skel.	17	—	D1: Akazawa et al. (1993)[29] D2: Akazawa et al. (1999) [30]
Iraq	Shanidar	S1: Adult partial skel. ♂ S2: Adult crushed skel. ♂ S3: Post-cranial adult ♂ S4: Adult partial skel. (♂) S5: Adult partial skel. (♂) S6: Adult partial skel. (♀) S7: 6-9-mo.-old crushed skel. S8: Adult skeletal fragments (♀) S9: 6-12-month-old vertebrae S10: 17-25-month-old skel.	10	S2, S4: > 100 Others: 60	S1: Stewart (1959)[31] S2: Stewart (1961)[32] S3: Solecki (1960)[33] S4: Stewart (1963)[34] S5: Trinkaus (1977)[35] Pomeroy et al. (2017)[36] S6: Same as S4 S7: Senyürek (1957)[37][38] S8: Same as S4 S9: ? S10: Cowgill et al. (2007)[39]	Shanidar 2 and 4 are sometimes not treated as Neanderthals. All but Shanidar 3 and 10 (and fragments of 5 excavated in 2015-2016)[36] may have been destroyed in the 2003 invasion of Iraq.[40]
Iran	Wezmeh	maxillary right premolar tooth	1	70-40[41]	Zanolli et al (2019)[41]
Iran	Bisitun	Adult radius shaft	1	—	Trinkaus and Biglari (2006)[42]
Total			71 + (13)

1. M_?-r according to S. Binford (1966)[25].
2. Valladas, Mercier, and Froget (1999) write that "[o]f the remains of 18 individuals recovered at the Amud Cave, 15 were derived from unambiguous Middle Palaeolithic contexts, all of them located in the northern area of the excavation (Hovers et al., 1995). The stratigraphic distribution of these remains encompasses the layers B₁ and B₂, with only a single specimen derived from layer B₄ (Sakura, 1970). Three individuals bear diagnostic characteristics which define them as Neanderthals. Amud I, the skeleton of an adult male was found at the top of layer B1, while the partial skeleton of the baby Amud 7 (Rak, Kimbet & Hovers, 1994) was recovered from the top of layer B₂, just under the contact with the base of layer B₁. Amud II, represented by a fragment of the right maxilla, was excavated from layer B₂."

Central and North Asian Neanderthals

Central Asian Neanderthals were found in Uzbekistan and North Asian Neanderthals in Asian Russia.

List of Central and North Asian Neanderthals

Country	Site	Principal Neanderthal finds	MNI	Geological age (ka)	Initial descriptions	Notes
Uzbekistan	Teshik-Tash	8-11-yr-old skeleton	1	—	Okladnikov (1949)
Uzbekistan	Obi-Rakhmat	Subadult skull frag. and teeth	1	74[43]	Glantz et al. (2008)[44]
Asian Russia	Chagyrskaya	Partial mandible	1	—	(Announced in Viola 2012)
Asian Russia	Okladnikov	Sub-adult humerus and femur	1		(Announced in Krause et al. 2007)[45]	mtDNA sampled
Asian Russia	Denisova	Altai 1: Toe phalanx♀ D11: Bone fragment	2	—	Mednikova (2011) Brown, et al. (2016)[46]	Altai 1: Full genome sequenced[47] D11: mtDNA sampled
Total			6

See also

• List of Southwest Asian Neanderthals
• Neanderthal Museum
• List of fossil sites
• List of human fossils

Notes

References

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Further reading

• Arsuaga, Juan Luis (2009). The Neanderthal's Necklace: In Search of the First Thinkers. New York: Four Walls Eight Windows. ISBN 0786740736.
• Gooch, Stan (2008). The Neanderthal Legacy: Reawakening Our Genetic and Cultural Origins. Rochester, Vermont: Inner Traditions. ISBN 159477742X.
• Muller, Stephanie Muller; Shrenk, Friedemann (2008). The Neanderthals. New York: Routledge. ISBN 1134095163.
• Silberman, Neil Asher, ed. (2012). The Oxford Companion to Archaeology (2 ed.). Oxford: Oxford University Press. ISBN 0199735786.
• Stringer, Chris (2012). Lone Survivors: How We Came to Be the Only Humans on Earth. New York: Macmillan. ISBN 1429973447.

External links

• "Homo neanderthalensis". Smithsonian Institution.