Salterella

Salterella is an enigmatic Cambrian genus with a small, conical, calcareous shell that appears to be septate,[3] but is rather filled with stratified laminar deposits.[4] The shell contains grains of sediment, which are obtained selectively (with a preference for denser grains) by a manner also observed in foramanifera.[5] The genus was established by Elkanah Billings in 1861, and was named after the English palaeontologist John William Salter.[2]

Salterella Temporal range: Early Cambrian PreꞒ Ꞓ O S D C P T J K Pg N
S. pulchella[1]
Scientific classification
Kingdom:	Animalia (?)
Phylum:	†Agmata
Family:	†Salterellidae
Genus:	†Salterella Billings, 1861[2]
Type species
†Salterella rugosa (= Serpulites maccullochi Murchison, 1859) Billings, 1861
Species
†S. conulata Clark, 1924 †S. maccullochi (Murchison, 1859)

The genus is known from multiple locations worldwide, such as Newfoundland and Labrador and Quebec in Canada,[6][7] Svalbard,[8] the Scottish Highlands[9] and Argentina.[10]

The related fossil genus Volborthella was formerly placed in synonymy with Salterella by Ellis L. Yochelson in 1983, due to the similarities between the two genera (though Volborthella notably lacks an outer calcareous shell).[9] However, Volborthella was later accepted as a separate genus again by Yochelson & Kisselev in 2003.[11] Both genera are currently placed in the Salterellidae family in the phylum Agmata.

Species

At least two species of Salterella are known:

• †Salterella conulata Clark, 1924[6]
• †Salterella maccullochi (Murchison, 1859) (Synonyms: S. mexicana Lochman, 1952; S. expansa Poulsen, 1927; S. rugosa Billings, 1861, Serpulites maccullochi Murchison, 1859)[9][10]

The following species placed in the genus are poorly known:[12]

• †Salterella pulchella Billings, 1861[2]
• †Salterella acervulosa Resser & Howell, 1938: A species that is abundant in the Kinzers Formation in Pennsylvania, but specimens of it are not well preserved.
• †Salterella curvata Shaler & Foerste, 1888: A species known only from the Hoppin Formation in Massachusetts. It was described from a single specimen, which has been lost.

Other:

• Salterella (?) orientalis Kobayashi, 1935:[13] Found in South Korea
• Salterella howitti Chapman, 1923: Found in Phosphate Hill Formation, Victoria, Australia

Former species

The following species were formerly placed in Salterella, and have since been moved to other genera:

• †"Salterella" obtusa Billings, 1861:[2] This Middle Cambrian species was moved to the genus Hyolithes d'Eichwald, 1840, and renamed Hyolithes billingsi Walcott, 1886.[1] It was later moved again to Linevitus Syssoiev, 1958.
• †"Salterella" billingsi Safford, 1869: A Middle Ordovician species of Salterella from the Murfreesboro Limestone formation in Tennessee described by James M. Safford in 1869. However, T. H. Clark considered it distinct from other Salterella fossils, creating the genus Polylopia Clark, 1925 to encompass it.[14]
• †"Salterella" hardmani Etheridge in Foord, 1890:[15] This Middle Cambrian species was described from the Ord Basin of Northern Territory, Australia. It was moved to the genus Biconulites Teilhard, 1931.[16] Much later, the genus Guduguwan Kruse, 1990 was created to contain it.[17]
• †"Salterella" planoconvexa Tate, 1892: A species described from South Australia. It belongs in Hyolitha.

See also

• Volborthella

References

1. Charles Doolittle Walcott (1886). Second contribution to the studies on the Cambrian faunas of North America. 30 of Geological Survey bulletin. Govt. Print. Off. pp. 1–369.
2. Billings, E.H. (1861). "On some new or little known species of lower Silurian fossils from the Potsdam Group (Primordial zone)". Palaeozoic Fossils. 1 (1): 1–18.
3. Lipps, J. H.; Sylvester, A. G. (1968). "The Enigmatic Cambrian Fossil Volborthella and Its Occurrence in California". Journal of Paleontology. 42 (2): 329–336. JSTOR 1302218.
4. Yochelson, E. L.; Flower, R. H.; Webers, G. F. (1973). "The bearing of the new Late Cambrian monoplacophoran genus Knightoconus upon the origin of the Cephalopoda". Lethaia. 6 (3): 275–309. doi:10.1111/j.1502-3931.1973.tb01199.x.
5. Peel, John S. (2016). "Anatase and Hadimopanella selection by Salterella from the Kap Troedsson formation (Cambrian Series 2) of North Greenland". GFF. 139 (1): 70–74. doi:10.1080/11035897.2016.1227365.
6. Yochelson, Ellis L. (1970). "On The Early Cambrian Fossil Salterella conulata Clark in Eastern North America" (PDF). United States Geological Survey Professional Paper. 683-B: 1–10.
7. Skovsted, Christian B. (2003). "Unusually preserved Salterella from the Lower Cambrian Forteau Formation of Newfoundland". GFF. 125 (1): 17–22. doi:10.1080/11035890301251017.
8. Lauritzen, Ørnulf; Yochelson, Ellis L. (1982). "Salterella rugosa (Early Cambrian: Agmata) on Nordaustlandet and Spitsbergen, Svalbard". Polar Research. 1982 (1): 5–16. doi:10.3402/polar.v1982i1.6996.
9. Yochelson, E.L. (1983). "Salterella (Early Cambrian: Agmata) from the Scottish Highlands" (PDF). Palaeontology. 26: 253–260.
10. Astini, Ricardo A.; Thomas, William A.; Yochelson, Ellis L. (2004). "Salterella in the Argentine Precordillera: an Early Cambrian palaeobiogeographic indicator of Laurentian affinity". Palaeogeography, Palaeoclimatology, Palaeoecology. 213 (1–2): 125–132. doi:10.1016/s0031-0182(04)00369-4.
11. Yochelson, Ellis L.; Kisselev, Gennadii N. (2003). "Early Cambrian Salterella and Volborthella (Phylum Agmata) re‐evaluated". Lethaia. 36 (1): 8–20. doi:10.1080/00241160310001254.
12. Yochelson, Ellis L. (1981). "A Survey of Salterella (Phylum Agmata)". In Taylor, Michael E. (ed.). Short papers for the Second International Symposium on the Cambrian System, 1981 (PDF) (Report). U.S. Geological Survey. pp. 244–248. doi:10.3133/ofr81743.
13. Kobayashi, T. (1935). "The Cambro-Ordovician formations and fauna of South Chosen. Palaeontology. Part III. Cambrian faunas of South Chosen with a special study on the Cambrian trilobite genera and families" (PDF). Journal of the Faculty of Science (Imperial University of Tokyo), Section II. 4: 49–344.
14. Yochelson, Ellis L. (1968). "On the nature of Polylopia" (PDF). United States Geological Survey Professional Paper. 593-F: 1–7.
15. Foord, A. H. (1890). "Description of Fossils from the Kimberley District, Western Australia". Geological Magazine. 7: 98–106. doi:10.1017/S0016756800189939.
16. Spath, L. F. (1936). "So-called Salterella from the Cambrian of Australia". Geological Magazine. 73 (10): 433–440. doi:10.1017/S0016756800095273.
17. Kruse, P. D. (1996). "Hyolith guts in the Cambrian of northern Australia – turning hyolithomorphs upside down". Lethaia. 29 (3): 213–217. doi:10.1111/j.1502-3931.1996.tb01651.x.