Protista taxonomy

A protist (/ˈprtɪst/) is any eukaryotic organism (one with cells containing a nucleus) that is not an animal, plant, or fungus. The protists do not form a natural group, or clade, since they exclude certain eukaryotes with whom they share a common ancestor;[lower-alpha 1] but, like algae or invertebrates, the grouping is used for convenience. In some systems of biological classification, such as the popular five-kingdom scheme proposed by Robert Whittaker in 1969, the protists make up a kingdom called Protista, composed of "organisms which are unicellular or unicellular-colonial and which form no tissues".[1][2][lower-alpha 2]

The following groups contain protists. The clade Opisthokonta also contains the animals and the fungi, and the kingdom Archaeplastida also contains algae and plants.

Unikonta (Sarcomastigota) Cavalier-Smith, 1987

Phylum Amoebozoa Cavalier-Smith, 1998

Clade Opisthokonta Cavalier-Smith 1981

Holozoa
Holomycota

Bikonta (Biciliata) Cavalier-Smith, 2002

Kingdom Archaeplastida (Primoplantae) Sina Adl, 2005

Division Glaucophyta Heinrich L. Skuja, 1954
Division Rhodophyta Wettstein, 1922
Subkingdom Plant

Kingdom Chromista Cavalier-Smith, 1981

Subkingdom SAR Cavalier-Smith, 2010

Infrakingdom Rhizaria Cavalier-Smith, 2002
Phylum Cercozoa Cavalier-Smith, 1998
Phylum Retaria Cavalier-Smith, 2002
  • Subphylum Foraminifera (Granuloreticulosa) d'Orbigny, 1826
    • Class Athalamea
      • Family Reticulomyxidae
    • Class Polythalamea
      • Order †Fusulinida
        • Superfamily †Archaediscacea Cushman, 1928
          • Family †Archaediscidae Cushman, 1928
          • Family †Lasiodiscidae Reytlinger, 1956
        • Superfamily †Colaniellacea Fursenko, 1959
          • Family †Colaniellidae Fursenko, 1959
        • Superfamily †Earlandiacea Cummings, 1955
          • Family †Earlandiidae Cummings, 1955
          • Family †Pseudoammodiscidae Conil & Lys, 1970
          • Family †Pseudolituotubidae Conil & Longerstaey, 1980
        • Superfamily †Endothyracea
          • Family †Endothyridae
        • Superfamily †Fusulinacea Leoblich & Tappan, 1988
        • Superfamily †Geinitzinacea
        • Superfamily †Moravamminacea Loeblich & Tappan, 1988
          • Family †Caligellidae
          • Family †Moravamminidae
          • Family †Paratickenellidae
        • Superfamily †Nodosinellacea Loeblich & Tappan, 1988
          • Family †Earlandinitidae
          • Family †Nodosinellidae
        • Superfamily †Palaeotextulariacea Galloway, 1933
          • Family †Biseriamminidae Chernysheva, 1941
          • Family †Palaeotextulariidae Galloway, 1933
          • Family †Semitextulariidae Pokorný, 1956
        • Superfamily †Parathuramminacea E.V. Bykova, 1955
          • Family †Archaesphaeridae Malakhova, 1956
          • Family †Auroriidae Loeblich & Tappan, 1986
          • Family †Chrysothuramminidae Loeblich & Tappan, 1986
          • Family †Eovolutinidae Loeblich & Tappan, 1986
          • Family †Ivanovellidae Chuvashov & Yuferev, 1984
          • Family †Marginaridae Loeblich & Tappan, 1986
          • Family †Parathuramminidae E.V. Bykova, 1955
          • Family †Tuberitinidae A.D. Miklukho-Maklay, 1958
          • Family †Uralinellidae Chuvashov, Yuferev & Zadorozhnyy, 1984
          • Family †Usloniidae A.D. Miklukho-Maklay, 1963
        • Superfamily †Ptychocladiacea Elias, 1950
          • Family †Ptychocladiidae Elias, 1950
        • Superfamily †Tetrataxacea Galloway, 1933
          • Family †Abadehellidae Loeblich & Tappan, 1984
          • Family †Pseudotaxidae Mamet, 1974
          • Family †Tetrataxidae Galloway, 1933
          • Family †Valvulinellidae Loeblich & Tappan, 1984
        • Superfamily †Tournayellacea Dain, 1953
          • Family †Palaeospiroplectamminidae Loeblich & Tappan, 1984
          • Family †Tournayellidae Dain, 1953
      • Order †Involutinida
      • Order Allogromiida
        • Family Allogromiidae
        • Family Hospitellidae
        • Family Lagynidae
        • Family Maylisoriidae
        • Family Phthanotrochidae
      • Order Carterinida
      • Order Globigerinida Delage & Hérouard, 1896
        • Superfamily †Globotruncanacea Brotzen, 1942
          • Family †Globotruncanidae Brotzen, 1942
          • Family †Rugoglobigerinidae Subbotina, 1959
        • Superfamily †Hantkeninacea Cushman, 1927
          • Family †Cassigerinellidae Bolli, Loeblich & Tappan, 1957
          • Family †Globanomalinidae Loeblich & Tappan, 1984
          • Family †Hantkeninidae Cushman, 1927
        • Superfamily †Planomalinacea Bolli, Loeblich & Tappan, 1957
          • Family †Globigerinelloididae Longoria, 1974
          • Family †Planomalinidae Bolli, Loeblich & Tappan, 1957
          • Family †Schackoinidae Pokorný, 1958
        • Superfamily †Rotaliporacea Sigal, 1958
          • Family †Favusellidae Longoria, 1974
          • Family †Globuligerinidae Loeblich & Tappan, 1984
          • Family †Hedbergellidae Loeblich & Tappan, 1961
          • Family †Rotaliporidae Sigal, 1958
        • Superfamily Globigerinacea Carpenter, Parker & Jones, 1862
          • Family Globigerinidae Carpenter, Parker & Jones, 1862
          • Family Hastigerinidae Bolli, Loeblich & Tappan, 1957
        • Superfamily Globorotaliacea Cushman, 1927
          • Family †Catapsydracidae Bolli, Loeblich & Tappan, 1957
          • Family †Eoglobigerinidae Blow, 1979
          • Family †Truncorotaloididae Loeblich & Tappan, 1961
          • Family Candeinidae Cushman, 1927
          • Family Globorotaliidae Cushman, 1927
          • Family Pulleniatinidae Cushman, 1927
        • Superfamily Heterohelicacea Cushman, 1927
          • Family †Heterohelicidae Cushman, 1927
          • Family Chiloguembelinidae Reiss, 1963
          • Family Guembelitriidae Montanaro Gallitelli, 1957
      • Order Lagenida
        • Superfamily Nodosariacea
          • Family Nodosariidae
          • Family Ellipsolagenidae
          • Family Glandulinidae
          • Family Lagenidae
          • Family Polymorphinidae
          • Family Vaginulinidae
        • Superfamily Robuloidacea
          • Family Robuloididae
          • Family Ichthyolariidae
          • Family Partisaniidae
          • Family Syzraniidae
      • Order Lituolida
        • Superfamily Ammodiscoidea
          • Family Ammodiscidae
        • Superfamily Ataxophragmioidea
          • Family †Ataxophragmiidae
          • Family †Cuneolinidae
          • Family Globotextulariidae
          • Family Textulariellidae
        • Superfamily Haplophragmioidea
          • Family Ammobaculinidae
          • Family Ammosphaeroidinidae
          • Family Discamminidae
          • Family Haplophragmoididae
        • Superfamily Hormosinelloidea
          • Family Ammolagenidae
          • Family Hormosinellidae
        • Superfamily Hormosinoidea
          • Family Aschemocellidae
          • Family Dusenburyinidae
          • Family Hormosinidae
          • Family Sphaeramminidae
          • Family Telamminidae
        • Superfamily Lituoloidea
          • Family Lituolidae
          • Family Placopsilinidae
        • Superfamily Lituotuboidea
          • Family Lituotubidae
          • Family Trochamminoidae
        • Superfamily Loftusioidea
        • Superfamily Orbitolinoidea
          • Family Orbitolinidae
        • Superfamily Rzehakinoidea
        • Superfamily Spiroplectamminoidea
          • Family Duquepsammiidae
          • Family Nouriidae
          • Family Pseudobolivinidae
          • Family Spiroplectamminidae
          • Family Textulariopsidae
        • Superfamily Verneuilinoidea
          • Family Prolixoplectidae
          • Family Reophacellidae
          • Family Tritaxiidae
          • Family Verneuilinidae
      • Order Miliolida
        • Superfamily Cornuspiracea Schultze, 1854
          • Family †Baisalinidae Loeblich & Tappan, 1986
          • Family Cornuspiridae Schultze, 1854
          • Family Discospirinidae Wiesner, 1931
          • Family Fischerinidae Millett, 1898
          • Family Hemigordiopsidae A. Nikitina, 1969
          • Family Nubeculariidae Jones, 1875
          • Family Ophthalmidiidae Wiesner, 1920
        • Superfamily Miliolacea
        • Superfamily Soritacea Ehrenberg, 1839
        • Superfamily Squamulinacea Reuss & Fritsch, 1861
          • Family Squamulinidae Reuss & Fritsch, 1861
      • Order Robertinida Loeblich & Tappan, 1984
        • Superfamily †Conorboiodacea Thalmann, 1952
          • Family †Conorboididae Thalmann, 1952
        • Superfamily †Duostominacea Brotzen, 1963
          • Family †Asymmetrinidae Brotzen, 1964
          • Family †Duostominidae Brotzen, 1963
          • Family †Oberhauserellidae Fuchs, 1970
        • Superfamily Ceratobuliminacea Cushman, 1927
          • Family Ceratobuliminidae Cushman, 1927
          • Family Epistominidae Wedekind, 1937
        • Superfamily Robertinacea Reuss, 1850
          • Family Robertinidae Reuss, 1850
      • Order Rotaliida Delage & Hérouard, 1896
        • Superfamily †Eouvigerinacea Cushman, 1927
          • Family †Eouvigerinidae Cushman, 1927
          • Family †Lacosteinidae Sigal, 1952
        • Superfamily †Orbitoidacea Schwager, 1876
          • Family †Lepidorbitoididae Vaughan, 1933
          • Family †Linderinidae Loeblich & Tappan, 1984
          • Family †Orbitoididae Schwager, 1876
        • Superfamily Acervulinacea Schultze, 1854
          • Family Acervulinidae Schultze, 1854
          • Family Homotrematidae Cushman, 1927
        • Superfamily Annulopatellinacea Loeblich & Tappan, 1964
          • Family Annulopatellinidae Loeblich & Tappan, 1965
        • Superfamily Asterigerinacea d'Orbigny, 1839
          • Family †Boreloididae Reiss, 1963
          • Family †Lepidocyclinidae Scheffen, 1932
          • Family Alfredinidae S.N. Singh & Kalia, 1972
          • Family Amphisteginidae Cushman, 1927
          • Family Asterigerinatidae Reiss, 1963
          • Family Asterigerinidae d'Orbigny, 1839
          • Family Epistomariidae Hofker, 1954
        • Superfamily Bolivinacea Glaessner, 1937
          • Family †Bolivinoididae Loeblich & Tappan, 1984
          • Family Bolivinidae Glaessner, 1937
        • Superfamily Bolivinitacea Cushman, 1927
          • Family Bolivinitidae Cushman, 1927
        • Superfamily Buliminacea Jones, 1875
          • Family Buliminellidae Hofker, 1951
          • Family Buliminidae Jones, 1875
          • Family Millettiidae Saidova, 1981
          • Family Pavoninidae Eimer & Fickert, 1899
          • Family Reussellidae Cushman, 1933
          • Family Siphogenerinoididae Saidova, 1981
          • Family Trimosinidae Saidova, 1981
          • Family Uvigerinidae Haeckel, 1894
        • Superfamily Cassidulinacea d'Orbigny, 1839
          • Family †Cassidulinidae d'Orbigny, 1839
          • Family Cassidulinitidae Saidova, 1981
        • Superfamily Chilostomellacea Brady, 1881
          • Family †Coleitidae Loeblich & Tappan, 1984
          • Family †Globorotalitidae Loeblich & Tappan, 1984
          • Family Alabaminidae Hofker, 1951
          • Family Chilostomellidae Brady, 1881
          • Family Gavelinellidae Hofker, 1956
          • Family Heterolepidae González-Donoso, 1969
          • Family Karreriidae Saidova, 1981
          • Family Oridorsalidae Loeblich & Tappan, 1984
          • Family Osangulariidae Loeblich & Tappan, 1964
          • Family Quadrimorphinidae Saidova, 1981
          • Family Trichohyalidae Saidova, 1981
        • Superfamily Delosinacea Parr, 1950
          • Family †Tremachoridae Lipps & K.L. Lipps, 1967
          • Family Caucasinidae Parr, 1950
          • Family Delosinidae Parr, 1950
        • Superfamily Discorbacea Ehrenberg, 1838
          • Family †Conorbinidae Reiss, 1963
          • Family Bagginidae Cushman, 1927
          • Family Bueningiidae Saidova, 1981
          • Family Eponididae Hofker, 1951
          • Family Heleninidae Loeblich & Tappan, 1988
          • Family Mississippinidae Saidova, 1981
          • Family Pannellainidae Loeblich & Tappan, 1967
          • Family Pegidiidae Heron-Allen & Earland, 1928
          • Family Placentulinidae G.K. Kasimova, Poroshina & Geodakchan, 1980
          • Family Rosalinidae Reiss, 1963
          • Family Rotaliellidae Loeblich & Tappan, 1964
          • Family Sphaeroidinidae Cushman, 1927
          • Family Ungulatellidae Seiglie, 1964
        • Superfamily Discorbinellacea Sigal, 1952
          • Family Parrelloididae Hofker, 1956
          • Family Planulinoididae Saidova, 1981
          • Family Pseudoparrellidae Voloshinova, 1952
        • Superfamily Fursenkoinacea Loeblich & Tappan, 1961
          • Family Fursenkoinidae Loeblich & Tappan, 1962
          • Family Virgulinellidae Loeblich & Tappan, 1984
        • Superfamily Glabratellacea Loeblich & Tappan, 1964
          • Family Buliminoididae Seiglie, 1970
          • Family Glabratellidae Loeblich & Tappan, 1964
          • Family Heronalleniidae Loeblich & Tappan, 1986
        • Superfamily Loxostomatacea Loeblich & Tappan, 1962
          • Family †Loxostomatidae Loeblich & Tappan, 1962
          • Family Bolivinellidae Hayward, 1980
          • Family Tortoplectellidae Loeblich & Tappan, 1985
        • Superfamily Nonionacea Schultze, 1854
          • Family Almaenidae Myatlyuk, 1959
          • Family Nonionidae Schultze, 1854
          • Family Spirotectinidae Saidova, 1981
        • Superfamily Nummulitacea de Blainville, 1827
          • Family †Asterocyclinidae Brönnimann, 1951
          • Family †Discocyclinidae Galloway, 1928
          • Family †Pellatispiridae Hanzawa, 1937
          • Family Nummulitidae de Blainville, 1827
        • Superfamily Planorbulinacea Schwager, 1877
          • Family Bisacciidae Loeblich & Tappan, 1988
          • Family Cibicididae Cushman, 1927
          • Family Cymbaloporidae Cushman, 1927
          • Family Planorbulinidae Schwager, 1877
          • Family Planulinidae Bermúdez, 1952
          • Family Victoriellidae Chapman & Crespin, 1930
        • Superfamily Pleurostomellacea Reuss, 1860
          • Family Pleurostomellidae Reuss, 1860
        • Superfamily Rotaliacea Ehrenberg, 1839
          • Family †Chapmaninidae Thalmann, 1938
          • Family †Miogypsinidae Vaughan, 1928
          • Family †Pseudorbitoididae M.G. Rutten, 1935
          • Family Calcarinidae Schwager, 1876
          • Family Elphidiidae Galloway, 1933
          • Family Rotaliidae Ehrenberg, 1839
        • Superfamily Siphoninacea Cushman, 1927
          • Family Siphoninidae Cushman, 1927
        • Superfamily Stilostomellacea Finlay, 1947
          • Family Stilostomellidae Finlay, 1947
        • Superfamily Turrilinacea Cushman, 1927
          • Family †Turrilinidae Cushman, 1927
          • Family Stainforthiidae Reiss, 1963
          • Family Tosaiidae Saidova, 1981
      • Order Silicoloculinida
      • Order Spirillinida
        • Family Patellinidae
        • Family Spirillinidae
      • Order Textulariida Delage & Herouard, 1896
        • Superfamily †Orbitolinacea Martin, 1890
          • Family †Orbitolinidae Martin, 1890
        • Superfamily †Pavonitinacea Loeblich & Tappan, 1961
          • Family †Marieitidae Loeblich & Tappan, 1986
        • Superfamily †Biokovinacea Gusiæ, 1977
          • Family †Biokovinidae Gusiæ, 1977
          • Family †Charentiidae Loeblich & Tappan, 1984
          • Family †Lituoliporidae Gusiæ & Veliæ, 1978
        • Superfamily †Cyclolinacea Loeblich & Tappan, 1964
          • Family †Cyclolinidae Loeblich & Tappan, 1964
          • Family †Orbitopsellidae Hottinger & Caus, 1982
        • Superfamily Ataxophragmiacea Schwager, 1877
          • Family †Ataxophragmiidae Schwager, 1877
          • Family †Coskinolinidae Moullade, 1965
          • Family †Cuneolinidae Saidova, 1981
          • Family †Dictyopsellidae Bröniman, Zaninetti & Whittaker, 1983
          • Family †Dicyclinidae Loeblich & Tappan, 1964
          • Family †Pfenderinidae Smout & Smugden, 1962
          • Family Globotextulariidae Cushman, 1927
          • Family Textulariellidae Grönhagen & Luterbacher, 1966
        • Superfamily Coscinophragmatacea Thalmann, 1951
          • Family Coscinophragmatidae Thalmann, 1951
          • Family Haddoniidae Saidova, 1981
        • Superfamily Haplophragmiacea Eimer & Fickert, 1899
          • Family †Barkerinidae Smout, 1956
          • Family †Haplophragmiidae Eimer & Fickert, 1899
          • Family †Labyrinthidomatidae Loeblich & Tappan, 1987
          • Family †Nezzazatidae Hamaoui & Saint-Marc, 1970
          • Family Acupeinidae Brnimann & Zaninetti, 1984
          • Family Ammobaculinidae Saidova, 1981
          • Family Ammosphaeroidinidae Cushman, 1927
        • Superfamily Hormosinacea Haeckel, 1894
          • Family †Cribratinidae Loeblich & Tappan, 1964
          • Family Ascheomocellidae Vyalov, 1966
          • Family Dusenburyinidae Loeblich & Tappan, 1984
          • Family Hormosinidae Haeckel, 1894
          • Family Telamminidae Loeblich & Tappan, 1985
          • Family Thomasinellidae Loeblich & Tappan, 1984
        • Superfamily Lituolacea de Blainville, 1827
          • Family †Mayncinidae Loeblich & Tappan, 1985
          • Family †Nautiloculinidae Loeblich & Tappan, 1985
          • Family †Oxinoxisidae Vyalov, 1968
          • Family Discamminidae Mikhalevich, 1980
          • Family Haplophragmoididae Maync, 1952
          • Family Lituolidae de Blainville, 1827
          • Family Lituotubidae Loeblich & Tappan, 1984
          • Family Placopsilinidae Rhumbler, 1913
          • Family Sphaeramminidae Cushman, 1933
        • Superfamily Loftusiacea Brady, 1884
          • Family †Ecougellidae Loeblich & Tappan, 1985
          • Family †Hottingeritidae Loeblich & Tappan, 1985
          • Family †Loftusiidae Brady, 1884
          • Family †Mesoendothyridae Voloshinova, 1958
          • Family †Spirocyclinidae Munier-Chalmas, 1887
          • Family Cyclamminidae Marie, 1941
        • Superfamily Rzehakinacea Cushman, 1933
        • Superfamily Spiroplectamminacea Cushman, 1927
          • Family †Plectrorecurvoididae Loeblich & Tappan, 1964
          • Family †Textulariopsidae Loeblich & Tappan, 1982
          • Family Nouriidae Chapman & Parr, 1936
          • Family Pseudobolivinidae Wiesner, 1931
          • Family Spiroplectamminidae Cushman, 1927
        • Superfamily Textulariacea Ehrenberg, 1838
          • Family †Chrysalidinidae Neagu, 1968
          • Family †Valvulamminidae Loeblich & Tappan, 1986
          • Family Eggerellidae Cushman, 1937
          • Family Glaucoamminidae Saidova, 1981
          • Family Pseudogaudryinidae Loeblich & Tappan, 1985
          • Family Textulariidae Ehrenberg, 1838
          • Family Valvulinidae Berthelin, 1880
        • Superfamily Trochamminacea Schwager, 1877
          • Family Remaneicidae Loeblich & Tappan, 1964
          • Family Trochamminidae Schwager, 1877
        • Superfamily Verneuilinacea Cushman, 1911
          • Family †Conorbinellidae Brönnimann, Zaninetti & Whittaker, 1983
          • Family †Conotrochamminidae Saidova, 1981
          • Family †Tritaxiidae Plotnikova, 1979
          • Family Prolixoplectidae Loeblich & Tappan, 1985
          • Family Verneuilinidae Cushman, 1911
    • Class Schizocladea
      • Family Schizocladidae
    • Class Xenophyophorea
      • Order Psamminida
        • Family Cerelasmidae
        • Family Psammettidae
        • Family Psamminidae
        • Family Syringamminidae
      • Order Stannomida
        • Family Pelosinidae
        • Family Stannomidae
  • Subphylum Radiozoa
    • Superclass Spasmaria
      • Class Acantharea Haeckel, 1881
        • Order Arthracanthida
          • Suborder Phyllacanthina
            • Family Dictyacanthidae
            • Family Phyllostauridae
            • Family Stauracanthidae
          • Suborder Sphaenacanthina
            • Family Acanthometridae
            • Family Aspidommatidae
            • Family Diploconidae
            • Family Dorataspididae
            • Family Hexalaspididae
            • Family Lithopteridae
            • Family Nivaliidae
        • Order Chaunacanthida
          • Family Conacontidae
          • Family Gigartacontidae
          • Family Stauracontidae
        • Order Holacanthida
          • Family Acanthochiasmatidae
          • Family Acanthoplegmatidae
          • Family Chiastolidae
        • Order Symphyacanthida
          • Family Amphilithiidae
          • Family Astrolithiidae
          • Family Pseudolithiidae
    • Superclass Radiolaria Müller, 1858
      • Class Polycystinea
        • Order Albaillellaria Deflandre, 1953
          • Family Albaillellidae Deflandre, 1953 sensu Holdsworth (1977)
          • Family Ceratoikiscidae Holdsworth, 1969
          • Family Corythoecidae
          • Family Follicucullidae Ormiston & Babcock, 1979
          • Family Palacantholithidae
        • Order Archaeospicularia Dumitrica, Caridroit & De Wever, 2000
            • Family Archeoentactiniidae Won & Below, 1999
          • Superfamily Echidninacea
            • Family Echidninidae Kozur, Mostler & Repetski, 1996
          • Superfamily Secuicollactacea Nazarov & Ormiston, 1984
            • Family Secuicollactidae
            • Family Pseudorotasphaeridae
        • Order Collodaria Haeckel 1881
          • Family Collosphaeridae
          • Family Sphaerozoidae
          • Family Thalassosphaeridae
        • Order Entactinaria Kozur & Mostler, 1982
          • Family Capnuchosphaeridae
          • Family Entactiniidae
          • Family Eptingiidae
          • Family Heptacladidae
          • Family Hexalonchidae
          • Family Hexastylidae
          • Family Hindeosphaeridae
          • Family Inaniguttidae
          • Family Kungalariidae
          • Family Multiarcusellidae
          • Family Palaeolithocycliidae
          • Family Palaeoscenidiidae
          • Family Polyentactiniidae
          • Family Proventocitidae
          • Family Pylentonemidae
          • Family Quinquecapsulariidae
          • Family Rhizosphaeridae
          • Family Saturnalidae
          • Family Spongentactiniidae
          • Family Spongosaturnaloididae
          • Family Thalassothamnidae
        • Order Latentifistularia Caridroit, 1999
          • Family Cauletellidae
          • Family Latentifistulidae
          • Family Ormistonellidae
        • Order Nassellaria
          • Superfamily Acanthodesmiacea
            • Family Acanthodesmiidae
            • Family Artostrobiidae
            • Family Bulbocyrtiidae
            • Family Cannobotryidae
            • Family Carpocaniidae
            • Family Cuniculiformidae
            • Family Monicastericidae
            • Family Planispinocyrtiidae
            • Family Pterocorythidae
            • Family Rotaformidae
            • Family Sethophormididae
            • Family Spongolophophenidae
            • Family Spongosilicarmigeridae
            • Family Stephaniidae
            • Family Theopiliidae
            • Family Triospyridae
          • Superfamily Amphipyndacea
            • Family Amphipyndacidae
            • Family Canoptidae
            • Family Parvicingulidae
            • Family Spongocapsulidae
            • Family Syringocapsidae
          • Superfamily Archaeodictyomitracea
            • Family Archaeodictyomitridae
            • Family Hsuidae
            • Family Unumidae
          • Superfamily Eucyrtidiacea
            • Family Bekomidae
            • Family Eucyrtidiidae
            • Family Eucyrtiidae
            • Family Lophocyrtiidae
            • Family Obeliscoitidae
            • Family Pseudodictyomitridae
            • Family Theocotylidae
            • Family Xitidae
        • Order Spumellaria Ehrenberg, 1875
          • Family Actinommidae
          • Family Anakrusidae
          • Family Angulobracchiidae
          • Family Archaeospongoprunidae
          • Family Astrosphaeridae
          • Family Bolenidae
          • Family Catenopylidae
          • Family Cavaspongiidae
          • Family Coccodiscidae
          • Family Conocaryommidae
          • Family Dactyliosphaeridae
          • Family Emiluviidae
          • Family Entapiidae
          • Family Ethmosphaeridae
          • Family Gomberellidae
          • Family Hagiastridae
          • Family Heliodiscidae
          • Family Hexaporobrachiidae
          • Family Larnacillidae
          • Family Leugeonidae
          • Family Litheliidae
          • Family Miropylidae
          • Family Myelastridae
          • Family Oertlispongidae
          • Family Orbiculiformidae
          • Family Pantanelliidae
          • Family Parasaturnalidae
          • Family Parvivaccidae
          • Family Patulibracchiidae
          • Family Patruliidae
          • Family Phacodiscidae
          • Family Phaseliformidae
          • Family Praeconocaryommidae
          • Family Pseudoacanthocircidae
          • Family Pseudoaulophacidae
          • Family Pyloniidae
          • Family Pyramispongiidae
          • Family Relindellidae
          • Family Spongodiscidae
          • Family Sponguridae
          • Family Stylosphaeridae
          • Family Thalassicollidae
          • Family Tholoniidae
          • Family Tritrabidae
          • Family Veghicycliidae
          • Family Xiphostylidae
Clade Halvaria
Infrakingdom Heterokontophyta (Heterokonta) Cavalier-Smith, 1986
Infrakingdom Alveolata Cavalier-Smith, 1991

Phlyum Ciliophora Doflein, 1901

  • Subphylum Postciliodesmatophora
    • Class Heterotrichea Stein, 1859
      • Order Armophorida
        • Family Caenomorphidae
        • Family Metopidae
      • Order Clevelandellida
        • Family Nyctotheridae
      • Order Heterotrichida
      • Order Licnophorida
        • Family Licnophoridae
      • Order Odontostomatida Sawaya, 1940
        • Family Epalxellidae Corliss, 1960
        • Family Discomorphellidae Corliss, 1960
        • Family Mylestomatidae Kahl, Doflein & Reichenow, 1929
      • Order Phacodiniida
        • Family Phacodiniidae
      • Order Plagiotomida
        • Family Plagiotomidae
    • Class Karyorelictea Corliss, 1974
  • Subphylum Intramacronucleata
    • Class Spirotrichea
      • Subclass Choreotrichia
        • Order Tintinnida
          • Family Ascampbelliellidae
          • Family Codonellidae
          • Family Codonellopsidae
          • Family Cyttarocylididae
          • Family Dictyocystidae
          • Family Epiplocylididae
          • Family Metacylididae
          • Family Nolaclusiliidae
          • Family Petalotrichidae
          • Family Ptychocylididae
          • Family Rhabdonellidae
          • Family Tintinnidae
          • Family Tintinnidiidae
          • Family Undellidae
          • Family Xystonellidae
      • Subclass Hypotrichia
        • Order Euplotida
          • Suborder Discocephalina
            • Family Discocephalidae
          • Suborder Euplotina
            • Family Aspidiscidae
            • Family Certesiidae
            • Family Euplotidae
            • Family Gastrocirrhidae
            • Family Uronychiidae
        • Order Kiitrichida
          • Family Kiitrichidae
      • Subclass Stichotrichia
    • Class Litostomatea
      • Subclass Haptoria Corliss, 1974
        • Order Cyclotrichiida
          • Family Mesodiniidae
        • Order Haptorida
          • Family Acropisthiidae
          • Family Didiniidae
          • Family Enchelyidae
          • Family Helicoprorodontidae
          • Family Homalozoonidae
          • Family Lacrymariidae
          • Family Pseudotrachelocercidae
          • Family Spathidiidae
          • Family Tracheliidae
          • Family Trachelophyllidae
        • Order Pleurostomatida
          • Family Amphileptidae
          • Family Litonotidae
      • Subclass Trichostomatia
        • Order Entodiniomorphida
          • Suborder Archistomatina de Puytorac, 1974
            • Family Buetschliidae Poche, 1913
          • Suborder Blepharocorythina Wolska, 1971
            • Family Blepharocorythidae Hsiung, 1930
          • Suborder Entodiniomorphina Reichenow, Doflein & Reichenow, 1929
            • Family Cycloposthiidae Poche, 1913
            • Family Ophryoscolecidae Stein, 1859
            • Family Spirodiniidae Strelkow, 1939
            • Family Troglodytellidae Corliss, 1979
        • Order Vestibuliferida
    • Class Phyllopharyngea
      • Subclass Chonotrichia
        • Order Cryptogemmida
          • Family Actinichonidae
          • Family Echinichonidae
          • Family Inversochonidae
          • Family Isochonidae
          • Family Isochonopsidae
          • Family Stylochonidae
        • Order Exogemmida
          • Family Chilodochonidae
          • Family Filichonidae
          • Family Heliochonidae
          • Family Lobochonidae
          • Family Phyllochonidae
          • Family Spirochonidae
      • Subclass Phyllopharyngia
        • Order Chlamydodontida
          • Family Chilodonellidae
          • Family Chitonellidae
          • Family Chlamydodontidae
          • Family Kryoprorodontidae
          • Family Lynchellidae
        • Order Dysteriida
          • Family Dysteriidae
          • Family Hartmannulidae
          • Family Kyaroikeidae
          • Family Plesiotrichopidae
      • Subclass Suctoria
        • Order Endogenida
          • Family Acinetidae
          • Family Acinetopsidae
          • Family Corynophryidae
          • Family Dendrosomatidae
          • Family Enchelyomorphidae
          • Family Endosphaeridae
          • Family Pseudogemmidae
          • Family Tokophryidae
          • Family Trichophryidae
        • Order Evaginogenida
          • Family Discophryidae
          • Family Rhynchophryidae
        • Order Exogenida
          • Family Dendrosomididae
          • Family Dentacinetidae
          • Family Ephelotidae
          • Family Lecanophryidae
          • Family Manuelophryidae
          • Family Metacinetidae
          • Family Ophryodendridae
          • Family Paracinetidae
          • Family Phalacrocleptidae
          • Family Podophryidae
          • Family Praethecacinetidae
          • Family Rhabdophryidae
          • Family Severonidae
          • Family Spelaeophryidae
          • Family Tachyblastonidae
          • Family Thecacinetidae
    • Class Colpodea
      • Order Colpodida
        • Family Colpodidae
    • Class Nassophorea
      • Order Nassulida
        • Family Nassulidae
        • Family Paranassulidae
      • Order Microthoracida
        • Family Discotrichidae
        • Family Leptopharyngidae
        • Family Microthoracidae
      • Order Synhymeniida
        • Family Nassulopsidae
        • Family Orthodonellidae
        • Family Scaphidiodontidae
    • Class Prostomatea
      • Order Prostomatida
        • Family Metacystidae
      • Order Prorodontida
        • Family Balanionidae
        • Family Colepidae
        • Family Holophryidae
        • Family Placidae
        • Family Plagiocampidae
        • Family Prorodontidae
        • Family Urotrichidae
    • Class Plagiopylea
      • Order Plagiopylida Small & Lynn, 1985
        • Family Plagiopylidae
        • Family Sonderidae
    • Class Oligohymenophorea
      • Subclass Peniculia
        • Order Peniculida
          • Suborder Frontoniina
            • Family Clathrostomatidae
            • Family Frontoniidae
            • Family Lembadionidae
            • Family Maritujidae
            • Family Stokesiidae
          • Suborder Parameciina
            • Family Neobursaridiidae
            • Family Parameciidae
            • Family Urocentridae
      • Subclass Hymenostomatia
        • Family Glaucomidae
        • Family Ichthyopthiriidae
        • Family Ophryoglenidae
        • Family Tetrahymenidae Corliss, 1952
        • Family Turaniellidae
      • Subclass Scuticociliatia
        • Order Philasterida
          • Family Cinetochilidae
          • Family Cohnilembidae
          • Family Cryptochilidae
          • Family Entodiscidae
          • Family Entorhipidiidae
          • Family Loxocephalidae
          • Family Orchitophryidae
          • Family Paralembidae
          • Family Parauronematidae
          • Family Philasteridae
          • Family Pseudocohnilembidae
          • Family Schizocaryidae
          • Family Thigmophryidae
          • Family Thyrophylacidae
          • Family Uronematidae
        • Order Pleuronematida
          • Family Calyptotrichidae
          • Family Ctedoctematidae
          • Family Cyclidiidae
          • Family Dragescoidae
          • Family Histiobalantiidae
          • Family Peniculistomatidae
          • Family Pleuronematidae
        • Order Thigmotrichida
          • Family Ancistridae
          • Family Hemispeiridae
          • Family Hysterocinetidae
          • Family Nucleocorbulidae
      • Subclass Astomatia
        • Order Astomatida
          • Family Anoplophryidae
          • Family Archiastomatidae
          • Family Clausilocolidae
          • Family Contophryidae
          • Family Haptophryidae
          • Family Hoplitophryidae
          • Family Intoshellinidae
          • Family Maupasellidae
          • Family Radiophryidae
      • Subclass Apostomatia
      • Subclass Peritrichia
        • Order Mobilida
          • Family Leiotrochidae
          • Family Polycyclidae
          • Family Trichodinidae
          • Family Urceolariidae
        • Order Sessilida
          • Family Ellobiophryidae
          • Family Epistylididae
          • Family Lagenophryidae
          • Family Operculariidae
          • Family Rovinjellidae
          • Family Scyphidiidae
          • Family Vaginicolidae
          • Family Vorticellidae
          • Family Zoothamniidae

Phylum Myzozoa T. Cavalier-Smith & E.E. Chao, 2004

Division Haptophyta (Prymnesiophyta) Hibberd & Cavalier-Smith, 1986

Division Cryptophyta Pascher, 1914

Excavata (Cavalier-Smith) Simpson, 2003

Eozoa Cavalier-Smith, 2003

Phylum Loukozoa Cavalier-Smith, 1999
Phylum Metamonada Grassé 1952
  • Subphylum Anaeromonada
    • Class Anaeromonadea
      • Order Trimastigida
        • Family Trimastigidae
      • Order Oxymonadida
        • Family Oxymonadidae
        • Family Polymastigidae
        • Family Pyrsonymphidae
        • Family Saccinobaculidae
        • Family Streblomastigidae
  • Subphylum Trichozoa
    • Superclass Parabasalia
      • Class Trichomonadea
        • Order Trichomonadida
          • Family Calonymphidae
          • Family Cochlosomatidae
          • Family Devescovinidae
          • Family Monocercomonadidae
          • Family Trichomonadidae
        • Order Lophomonadida
          • Family Deltotrichonymphidae
          • Family Joeniidae
          • Family Kofoidiidae
          • Family Lophomonadidae
          • Family Microjoeniidae
          • Family Rhizonymphidae
        • Order Spirotrichonymphida
          • Family Holomastigotidae
          • Family Holomastigotoididae
          • Family Spirotrichonymphidae
      • Class Trichonymphea
        • Order Trichonymphida
          • Family Eucomonymphidae
          • Family Hoplonymphidae
          • Family Spirotrichosomidae
          • Family Staurojoeninidae
          • Family Teranymphidae
          • Family Trichonymphidae
    • Superclass Carpediemonadia
    • Superclass Eopharyngia
      • Class Trepomonadea
        • Subclass Diplozoa
          • Order Distomatida Klebs, 1892
            • Family Hexamitidae Kent, 1880
          • Order Giardiida Kunstler, 1882
            • Family Giardiidae Cavalier-Smith, 1996
        • Subclass Enteromonadia
          • Order Enteromonadida
            • Family Enteromonadidae Kulda & Nohýnková, 1976
      • Class Retortamonadea

Discicristata Cavalier-Smith, 2003

Phylum Percolozoa Cavalier-Smith 1991

Phylum Euglenozoa (Euglenophyta) Cavalier-Smith, 1981[3]

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See also

Notes

  1. The first eukaryotes were “neither plants, animals, nor fungi”, hence as defined, the Protista would include the earliest common ancestor of all eukaryotes.
  2. In the original 4-kingdom model proposed in 1959, Protista included all unicellular microorganisms such as bacteria.

References

  1. Whittaker RH (January 1969). "New concepts of kingdoms or organisms. Evolutionary relations are better represented by new classifications than by the traditional two kingdoms". Science. 163 (3863): 150–160. Bibcode:1969Sci...163..150W. CiteSeerX 10.1.1.403.5430. doi:10.1126/science.163.3863.150. PMID 5762760.
  2. "whittaker new concepts of kingdoms – Google Scholar". scholar.google.ca. Retrieved 2016-02-28.
  3. T. Cavalier-Smith (1981). "Eukaryote kingdoms: seven or nine". BioSystems. 14 (3–4): 461–481. doi:10.1016/0303-2647(81)90050-2. PMID 7337818.
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