Hasle Formation

The Hasle Formation is a geologic formation on the island on Bornholm, Denmark. It is of early to late Pliensbachian age. Vertebrate fossils have been uncovered from this formation.[1][2]

Hasle Formation Stratigraphic range: Late Pliensbachian ~190 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Underlies	Sorthat Formation
Overlies	Rønne Formation
Thickness	80–140 m (260–460 ft)
Lithology
Primary	Siltstone, sandstone
Location
Coordinates	55.2°N 14.7°E / 55.2; 14.7
Approximate paleocoordinates	45.0°N 18.5°E / 45.0; 18.5
Region	Bornholm
Country	Denmark
Hasle Formation (Denmark)

Fossil content

Fish

Unidentified fin spines are known from this formation.[1]

Genus	Species	Material	Notes	Images
Hybodus[1]	H. reticulatus[1]	unknown
	H. delabechei[1]	unknown
Lissodus	L. hasleensis sp. n.[1]	unknown
Synechodus[1]	S. occultidens[1]	unknown
Paraorthacodus[1]	P. sp[1]	unknown
Agaleus[1]	A. dorsetensis[1]	unknown
Oblidens[3]	O. bornholmensis[1]	Upper posterior (palatine) and lower posterior (mandibular) tooth plates.

Reptiles

Theropods

Genus	Species	Material	Notes	Images
Stenonyx[4]	Indeterminate	25 mm footprints referred to a non-avian Theropod.[4]	Footprints of an Small Theropod Dinosaur, probably a member of Neotheropoda, related to genera such as Tachiraptor or Dracoraptor.

Sauropodomorpha

Genus	Species	Material	Notes	Images
Massopoda[5]	Massopoda Indeterminate	DK966, an upper part of a femur from a juvenile animal	A Massopoda Sauropodomorph. DK966 has been compared with various taxa from the lower jurassic: "Aristosaurus erectus", a young Massospondylus, various Massospondylus specimens, Lufengosaurus huenei holotype, Seitaad ruessi, Thecodontosaurus, Pantydraco, etc. DK966 can be probably a young specimen of Massopoda, related to Massospondylus, more than a Plateosaurid or an Anchisaurid. We find it related to "Hortalotarsus", "Aristosaurus" and Massospondylus more than Seitaad for example. Based on the several taxa we had compared probably presented a body similar to Pantydraco being a young specimen. The complete bone is estimated on only 6 cm, making the animal no more than 70 cm length.

Ammonites

Genus	Species	Material	Notes	Images
Aegoceras[2]	A. centaurus[2]	specimens[2]	Later renamed Beaniceras centaurus, senior synonym of Aegoceras centaurus[2]
	A. armatum[2]	specimens[2]
	A. capraius[2]	specimens[2]
	A. mangenesti[2]	specimens[2]
	A. pettos[2]	specimens[2]	Junior synonym of Coeloceras pettos, possible synonym of Coeloceras grenouillouxi[2]
Uncertain	? Acanthopleuroceras sp. indet.[2]	specimens[2]
Apderoceras[2]	A. aculeatum[2]	specimens[2]
Arnioceras[2]	A. falcaries[2]	specimens[2]
Beaniceras[2]	B. centaurus[2]	specimens[2]	Senior synonym of Aegoceras centaurus[2]
Coeloceras[2]	C. pettos[2]	specimens[2]	Senior synonym of Aegoceras pettos. Possible junior synonym of Coeloceras grenouillouxi[2]
	C. grenouillouxi[2]	specimens[2]	Possible senior synonym of Coeloceras pettos[2]
Paramicroderoceras[2]	P. fila[2]	specimens[2]
	P.? sp indet[2]	specimens[2]
Phylloceras[2]	P. losconnbii[2]	specimens[2]
Phricedoceras[2]	P. taylori[2]	specimens[2]
Platypleuroceras[2]	P. brevispina[2]	specimens[2]
	P. submulticum[2]	specimens[2]
	P. caprarium[2]	specimens[2]
	P. spp. indet[2]	specimens[2]
Radstockiceras[2]	R. hechingense[2]	specimens[2]
Tragophyloceras[2]	T. numismale[2]	specimens[2]
	T. sp[2]	specimens[2]
Uptonia[2]	U. lata[2]	specimens[2]
	U. sp indet[2]	specimens[2]

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon		Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

See also

• List of fossiliferous stratigraphic units in Denmark

References

1. Rees, J. (1998). "Early Jurassic selachians from the Hasle Formation on Bornholm, Denmark" (PDF). Acta Palaeontologica Polonica. 43 (3): 43–439.
2. Donovan, Desmond T. & Surlyk, Finn. "Lower Jurassic (Pliensbachian) ammonites from Bornholm, Baltic Sea, Denmark" (PDF). Geological Survey of Denmark and Greenland Bulletin. 1: 555–583.CS1 maint: uses authors parameter (link)
3. Christopher J. Duffin; Jesper Milàn (2017). "A new myriacanthid holocephalian from the Early Jurassic of Denmark" (PDF). Bulletin of the Geological Society of Denmark. 65: 161–170.
4. Jesper Milàn, Finn Surlyk. 2014: An enigmatic, diminutive theropod footprint in the shallow marine Pliensbachian Hasle Formation, Bornholm, Denmark
5. Milàn, J., Cuny, G. (2019):Knogle fra mini sauropod fundet ved Hasle. Geomuseum Faxe Danmark. University of Lyon Frankrig. https://www.researchgate.net/publication/332291953_Knogle_fra_mini_sauropod_fundet_ved_Hasle