Geotrupidae

Geotrupidae (from Greek geos, earth, and trypetes, borer) is a family of beetles in the order Coleoptera. They are commonly called earth-boring dung beetles. Most excavate burrows in which to lay their eggs. They are typically detritivores, provisioning their nests with leaf litter (often moldy), but are occasionally coprophagous, similar to dung beetles. The eggs are laid in or upon the provision mass and buried, and the developing larvae feed upon the provisions. The burrows of some species can exceed 2 metres in depth.

Geotrupidae
Geotrupes egeriei
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Polyphaga
Infraorder: Scarabaeiformia
Superfamily: Scarabaeoidea
Family: Geotrupidae
Latreille, 1802
Genera

25, see text

A few species communicate by stridulation (rubbing body parts together to make sounds).

Classification

They were originally classified as the subfamily Geotrupinae in the family Scarabaeidae before being elevated to a family. Traditionally the dor beetle family Bolboceratidae was included (as the subfamily Bolboceratinae) on the basis of the number of antenna segments, but examination of a different set of characteristics prompted Scholtz & Browne (1995) to elevate Bolboceratidae to a family, a result supported by recent phylogenetic research.[1]

The family has more than 600 species in about 30 genera in two subfamilies; recent phylogenetic studies indicate that Taurocerastinae is not related to Geotrupinae, and is instead more closely related to Lucanidae and Diphyllostomatidae.[1]

  • Geotrupinae
    • Allotrupes Boucomont, 1912
    • Anoplotrupes Jekel, 1866
    • Baraudia López-Colón, 1996
    • Ceratophyus Fischer von Waldheim, 1823
    • Ceratotrupes Jekel, 1865
    • Chelotrupes Jekel, 1866
    • Cnemotrupes Jekel, 1866
    • Cretogeotrupes Nikolajev, 1992
    • Enoplotrupes Lucas, 1869
    • Geohowdenius Zunino, 1984
    • Geotrupes Latreille, 1796
    • Halffterius Zunino, 1984
    • Haplogeotrupes Nikolaev, 1979
    • Jekelius López-Colón, 1989
    • Lethrus Scopoli, 1777
    • Megatrupes Zunino, 1984
    • Mycotrupes LeConte, 1866
    • Odontotrypes Fairmaire, 1887
    • Onthotrupes Howden, 1964
    • Phelotrupes Jekel, 1866
    • Peltotrupes Blanchard, 1888
    • Pseudotrypocopris Miksic, 1954
    • Sericotrupes Zunino, 1984
    • Silphotrupes Jekel, 1866
    • Thorectes Mulsant, 1842
    • Trypocopris Motschulsky, 1860
    • Typhaeus Leach, 1815
    • Zuninoeus López-Colón, 1989
  • Taurocerastinae
    • Frickius Germain, 1897
    • Taurocerastes Philippi, 1866
gollark: > would you like it if i said @random_person please vote for trump or @random_person please vote for hillaryyes.
gollark: AT LEAST TWO!!!!!!
gollark: Which Lyric did VARIOUS TIMES.
gollark: It's better than pinging @​everyone.
gollark: Ye§.

References
  1. MCKENNA, D. D., WILD, A. L., KANDA, K., BELLAMY, C. L., BEUTEL, R. G., CATERINO, M. S., FARNUM, C. W., HAWKS, D. C., IVIE, M. A., JAMESON, M. L., LESCHEN, R. A. B., MARVALDI, A. E., MCHUGH, J. V., NEWTON, A. F., ROBERTSON, J. A., THAYER, M. K., WHITING, M. F., LAWRENCE, J. F., ŚLIPIŃSKI, A., MADDISON, D. R. and FARRELL, B. D. (2015), The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Syst Entomol, 40: 835–880. doi:10.1111/syen.12132


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