Utaetus

Utaetus; Temporal range: Mid-Late Eocene; (Bartonian-Priabonian); ~40–36 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Cingulata
Family:	Dasypodidae
Genus:	†Utaetus; Ameghino 1902
Type species
	Utaetus buccatus; Ameghino 1902
Species
	U. buccatus Ameghino 1902 (type); U. deustus Ameghino 1902 (nomen dubium); U. laevus Ameghino 1902 (nomen dubium); U. lenis Ameghino 1902 (nomen dubium);
Synonyms
	Anteutatus Ameghino 1902; Orthutaetus Ameghino 1902; Posteutatus Ameghino 1902;

Utaetus is an extinct genus of mammal in the order Cingulata, related to the modern armadillos. The genus contains a single species, Utaetus buccatus. It lived in the Middle to Late Eocene (about 40 to 36 million years ago) and its fossil remains were found in Argentina and Brazil in South America.

Description

This animal, about 1 metre (3.3 ft) long, was very similar to a modern armadillo. In particular, the appearance likely recalled that of the modern Euphractus, and it already had the typical well-developed xenarthral joints on the vertebrae. Among the other characteristics in common with the modern armadillos, Utaetus possessed a bony connection between the ischium and the sacrum (this structure was constituted by caudal vertebrae known as pseudosacral) and continuous-growth cylindrical teeth similar to chisels, with wear in the occlusal part. There were ten lower teeth on each side of the jaw; the first two were much smaller and are interpreted as incisors. Unlike later armadillos, however, Utaetus still had a varying amount of enamel on the lingual and buccal surfaces of the teeth, and the cervical vertebrae were separated (and not co-ossified). The skeleton shows that this animal was suitable for digging, as evidenced by the presence of a large acromion on the scapula and a prominent olecranon on the ulna. The posterior margin of the scapula was thickened, and formed a secondary incipient spine.

Classification

The genus Utaetus was first described in 1902 by Florentino Ameghino, based on fossil remains initially thought to date back to the Cretaceous. The type species is Utaetus buccatus, also known for cranial material, but Ameghino described further species based on fragmentary remains (U. deustus, U. lenis, U. laevus, U. laxus), now considered nomina dubia.Utaetus is considered a primitive member of the Dasipodidae, the family that includes the modern armadillos and their extinct relatives. In particular,Utaetus and its close relatives (such as Parutaetus) could be closely related to the modern genus Euphractus. The name Utaetus is an anagram of Eutatus, another extinct armadillo.

Distribution

Fossils of Utaetus have been found in:[1]

Argentina
- Geste Formation (Divisaderan)
- Quebrada de los Colorados Formation (Barrancan)[2]
- Sarmiento Formation (Barrancan)
Brazil
- Guabirotuba Formation (Divisaderan)[3]

gollark: Pastebin loves randomly blocking people.

gollark: I actually had to mirror potatOS onto my server because of some technical errors™ on pastebin's end.

gollark: creativity 1010101010

gollark: haha yes

gollark: osmarks.tk is coping fine.

References

Utaetus at Fossilworks.org
Powell et al., 2011, p.361
Sedor et al., 2017

Bibliography

Powell, J.E.; M.J. Babot; D.A. García López; M.V. Deraco, and C. Herrera. 2011. Eocene vertebrates of northwest Argentina: annotated list in J. Salfity, R. A. Marquillas (eds.), Cenozoic Geology of the Central Andes of Argentina, 349–370. SCS Publisher. Accessed 2019-04-13.
Sedor, F.A.; É.V. Oliveira; D.D. Silva; L.A. Fernandes; R.F. Cunha; A.M. Ribeiro, and E.V. Dias. 2017. A New South American Paleogene Land Mammal Fauna, Guabirotuba Formation (Southern Brazil). Journal of Mammalian Evolution 24. 39–55. Accessed 2019-04-13.