Iris sambucina

Iris sambucina, the elder scented iris, is a plant species in the genus Iris, it is also in the subgenus Iris. It is a rhizomatous perennial, from southern and central Europe (Germany, GB and Ireland, Italy, the Balkans, Croatia), and Spain. It has green, curved or sword-like leaves, tall round stem (with branches), multiple flowers in shades from brown violet, or brown-purple, to purple-violet, blue violet, mauve, and to purple. The large flowers are fragrant, with the scent of elderflowers, hence the name. It was first considered a separate species (when found), then it was classified as a synonym of Iris germanica, before being classified (in some sources) as a separate species again, but with a hybrid origin from Iris pallida and Iris variegata. It is sometimes cultivated as an ornamental plant in temperate regions.

Iris sambucina
Iris sambucina flower in bud
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Monocots
Order: Asparagales
Family: Iridaceae
Genus: Iris
Subgenus: Iris subg. Iris
Section: Iris sect. Iris
Species:
I. sambucina
Binomial name
Iris sambucina
Synonyms[1][2]
  • Iris × lurida Aiton
  • Iris × squalens L.

Description

It is similar in form to Iris germanica, but it has darker colour flowers and fragranced flowers.[2]

It has thick, fleshy or fibrous rhizomes, that are well branched.[3][4]

It has herbaceous,[4][5] glaucous,[6] sword-like,[3] or curving,[6] green leaves.[3] They can grow (in spring,[4]) up to between 20–80 cm (8–31 in) long,[4] and between 1.3 and 2.5 cm wide.[3] They also smell strongly of elder (Sambucus ebulus).[7]

It has a round stem,[3] or peduncle, that can grow up to between 40–100 cm (16–39 in) tall.[8][9][10]:135 The stem is normally taller than the leaves,[11] and they are usually branched.[2][3]

The stem has large, spathes (leaves of the flower bud), that are green at the base but (scarious) or membranous on the top half.[3]

The stems (and the many branches) hold 2,[12] or more flowers,[11] blooming in summer,[6] between April,[4] or May,[8] to June.[11][13][14]

The large,[6][12] around 4–7 cm (2–3 in) long flowers,[3] are fragrant (with the scent of elderflowers),[12][15][16] and come in shades of violet,[8] from brown violet,[2][4][6] or brown-purple,[10]:135 purple-violet,[14] blue violet,[16] mauve,[5] to purple.[3][4][17] They are normally darker than Iris germanica.[7][17] They can also vary in colour from region to region.[4]

Like other irises, it has 2 pairs of petals, 3 large sepals (outer petals), known as the 'falls' and 3 inner, smaller petals (or tepals), known as the 'standards'.[10]:17 The deflexed falls are spathulate (spoon-like) or oblong shaped,[6][15] and can be veined,[2] brown-purple on yellow-white background,[10]:135 or with claret (dark red).[15] In the middle of the falls, is a row of short hairs called the 'beard', which is yellow,[2][3][6] golden yellow,[9] or orange.[2][4][10]:135 The upright standards are elliptical or oblong shaped, although the tips are inclined to each other.[4][6] They are also serrated,[11][17] or jagged.[6]

The style arms are light purple and 3–4 cm long, the perianth tube is 2–2.5 cm long.[3]

After the iris has flowered, most irises produces a seed capsule, but due to the hybrid origin, the capsules are not usually developed.[3]

Biochemistry

As most irises are diploid, having two sets of chromosomes, this can be used to identify hybrids and classification of groupings.[10]:18 It is published, that it a chromosome count: 2n=24,[9][15][18] or 2n=40.[4]

As Iris germanica has a count of 2n=44 or 48 and Iris variegata is counted as 2n=24.[19] Iris variegata is more likely to be the parent species.

Taxonomy
Painted illustration from Deutschlands Flora in Abbildungen by Jacob Sturm

It is commonly known as 'elder scented iris',[20][21][22] or 'Iris elderflower'.[3]

It is known as 'fläderiris' in Swedish,[1] or 'holunder schwertlilie' in Germany,[14] (meaning 'elderberry iris').[8][16]

The Latin specific epithet sambucina refers to sambuci-, sambucinus, -a, or -um elder-like or resembling Sambucus.[23] This refers to the smell of the flowers resembling the scent of elder-flowers.[2][24]

It was first published and described by Carl Linnaeus in his book Systema Naturae (Syst. Nat) Edition 10, Volume 2 on page 863, on 7 June 1759.[1][15][25]

It was later named Iris glauca by Richard Anthony Salisbury in the Transactions of the Horticultural Society of London Vol.1 on page 302 in 1812, but this was rejected by all other authors, due to Salisbury's plagiarism history.[25]

It was also published in Curtis's Botanical Magazine Vol.6 on tab.187 (with a colour illustration) in 1794, The Gardeners' Chronicle in 1883, the 'Handbook of the Irideae' by J. P. Baker, on page 86 in 1892 and the Journal of RHS Vol.15 on page 3 on October 1899.[15]

William Rickatson Dykes notes that Linnaeus had given the names of Iris sambucina and Iris squalens, to 2 similar irises, whose parents were Iris pallida and Iris variegata.[24][26]

It was later classified as a hybrid,[3][27] but it is unsure which species (of iris) are the parents.[4] It probably originated in the Mediterranean as a diploid species.[3] Many authors think it is a hybrid of Iris variegata L.[28] and Iris pallida Lam.[1][9][15][29]

Others think it is a synonym of Iris germanica L.,[12][30] including The Plant List.[31] It is listed in the Encyclopedia of Life and Catalogue of Life,[32] as a synonym of Iris germanica.[33]

It was verified as Iris sambucina by United States Department of Agriculture and the Agricultural Research Service on 9 January 2003, then updated on 12 September 2005.[1]Iris sambucina is a tentatively accepted name by the RHS and listed in the RHS Plant Finder book.[21]

Distribution and habitat

It is native to southern [6][7][11] or central Europe.[9]

Range

It is found in Europe, within Germany,[10]:135 GB and Ireland,[7] Italy, Balkans (former Yugoslavia),[1] and Spain.[29]

Habitat

It grows on sunny, well drained rock slopes,[3][4] in meadows of mountains, and on walls of ancient castles.[13] Within the South Tyrol (of Italy), the plants can be found at an altitude of 950 m (3,120 ft) above sea level.[13]

Conservation

It is considered rare, even though it was a wide distribution range.[4]

Cultivation

It is hardy to between USDA Zone 3a (from −39.9 °C (−40 °F)) to Zone 8b (up to −9.4 °C (15 °F)),[5] including Zone 5.[14]

It prefers to grow in calcareous,[8] well drained,[4][34] or rich soils,[2][6] in full sun.[3][5][8]

It can often been cultivated,[3][17] and can be grown in mixed flower borders, on the margins of shrubberies or being naturalized in the garden.[6][8] It was introduced to Britain in 1658,[2] and was then cultivated in 1748 in the UK, by Mr Philip Miller.[11] Although, it is not often found in bulb nurseries,[4] but a few cultivars can be found in German nurseries.[16]

It can though be attacked by slugs.[4]

Propagation

It can be propagated by division,[3][5][6] due to its hybrid origin, it is sterile and does not produce seed.[5]

Hybrids and cultivars

It can form hybrids that have flowers in combinations of yellow, red and brown hues.[3]

Known cultivars include; Adb-el-Kaker, Abel-Kaver, Cheron, Cherion, Gypsy Queen, Leopold, Leopoldine,[15] Mägdeberg,[16] Nemesis, President Lincoln and Virgil.[15]

Toxicity

Like many other irises, most parts of the plant are poisonous (rhizome and leaves), and if mistakenly ingested can cause stomach pains and vomiting. Handling the plant may cause skin irritation or an allergic reaction.[5]

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References
  1. "Taxon: Iris sambucina L." ars-grin.gov (Germplasm Resources Information Network). Retrieved 8 December 2015.
  2. Mills, Colin. "Iris sambucina L." hortuscamden.com. Retrieved 8 December 2015.
  3. Grulich, Vit (10 February 2014). "Iris x sambucina l. – iris elderflower" (in Czech). botany.cz. Retrieved 7 December 2015.
  4. "Chapter I Rhizomatous Iris (Part 3)". irisbotanique.over-blog.com. Retrieved 8 December 2015.
  5. "Tall Bearded Iris 'Sambucina's". davesgarden.com. Retrieved 7 December 2015.
  6. William Robinson Hardy Flowers (1878), p. 152, at Google Books
  7. Fitter, Richard; Fitter, Alastair (1974). The Wild Flowers of Britain and Northern Europe (3rd, Revised ed.). London: Collins (published 1978). p. 272. ISBN 0-00-219069-9.
  8. "Iris sambucina bio" (in German). shop.hof-berggarten.de. Archived from the original on 4 March 2016. Retrieved 12 January 2016.
  9. Cortopassi, Lucia (20 May 2014). "Iris sambucina". signa.org (Species Iris Group of North America). Retrieved 8 December 2015.
  10. Austin, Claire (2005). Irises; A Garden Encyclopedia. Timber Press. ISBN 0-88192-730-9.
  11. Lady Charlotte MurrayThe British Garden: A Descriptive Catalogue of Hardy Plants, Indigenous Or cultivated in the climate of Great Britain Vol.1 (1808) , p. 48, at Google Books
  12. William Prince Catalogue of Fruit and Ornamental Trees and Plants, Bulbous Flower Roots, Green-house plants. Cultivated at the Linnean Botanic Garden. (1822), p. 51, at Google Books
  13. Hegi, Gustav (1926). "Hochspringen nach". Illustrated Flora of Central Europe (in German). Part 2. Munich: JF Lehmann's publishing house. p. 290.
  14. "Holunder Schwertlilie" (in German). allgaeustauden.de. Retrieved 12 January 2016.
  15. Laurin, Terry (10 May 2014). "(SPEC) Iris Sambucina Fos". wiki.irises.org (American Iris Society). Retrieved 7 December 2015.
  16. "Iris sambucina" (in German). syringa-pflanzen.de. Retrieved 12 January 2016.
  17. Daniel Chambers Macreight Manual of British Botany: In which the Orders and Genera are Arranged and described according to the natural system of De Candolle, (1837), p. 230, at Google Books
  18. Oberdorfer, Erich; Schwabe, Angelika; Müller, Theo (2001). Phytosociological Exkursionsflora for Germany and adjacent areas (in German). 8. Stuttgart (Hohenheim): Eugen Ulmer. p. 141. ISBN 3-8001-3131-5.
  19. "Iris summary" (PDF). pacificbulbsociety.org. 14 April 2014. Retrieved 23 November 2014.
  20. John MackieA catalogue of forest trees, fruit trees, evergreen and flowering shrubs, hot-house, green-house and herbaceous plants, kitchen garden and flower seeds, p. 47, at Google Books
  21. "Iris × sambucina". www.rhs.org.uk. Retrieved 8 December 2015.
  22. Thomas Ignatius M. Forster Circle of the seasons, and perpetual key to the calendar and almanack (1828) , p. 159, at Google Books
  23. D. Gledhill The Names of Plants, p. 339, at Google Books
  24. Dykes, William. "Dykes on Iris" (PDF). beardlessiris.org (The Group for Beardless Irises). Retrieved 21 November 2014.
  25. "Iridaceae Iris sambucina L." ipni.org (International Plant Names Index). Retrieved 8 December 2015.
  26. Dykes, William (2009). "Handbook of Garden Irises" (PDF). beardlessiris.org (The Group for Beardless Irises). Retrieved 1 November 2014.
  27. Cassidy, George E.; Linnegar, Sidney (1987). Growing Irises (Revised ed.). Bromley: Christopher Helm. p. 123. ISBN 0-88192-089-4.
  28. British Iris Society (1997)A Guide to Species Irises: Their Identification and Cultivation, p. 57, at Google Books
  29. Stebbings, Geoff (1997). The Gardener's Guide to Growing Irises. Newton Abbot: David and Charles. p. 26. ISBN 0-7153-0539-5.
  30. Kelly Norris A Guide to Bearded Irises: Cultivating the Rainbow for Beginners and Enthusiasts (2012), p. 128, at Google Books
  31. "Iris × sambucina". theplantlist.org. 23 March 2012. Retrieved 8 December 2015.
  32. "Iris germanica L. (accepted name)". catalogueoflife.org. Retrieved 6 January 2015.
  33. "Iris germanica". eol.org. Retrieved 6 January 2015.
  34. Meyer, Chris J.; Seago Jr., James L.; Peterson, Carol A. (2009). "Environmental effects on the maturation of the endodermis and multiseriate exodermis of Iris germanica roots". Annals of Botany. Botanical Society of America. 103 (5): 687–702. doi:10.1093/aob/mcn255. PMC 2707867. PMID 19151041. Retrieved 7 January 2015.

Sources
  • Aldén, B., S. Ryman & M. Hjertson Våra kulturväxters namn – ursprung och användning. Formas, Stockholm (Handbook on Swedish cultivated and utility plants, their names and origin). 2009 (Vara kulturvaxt namn)
  • Czerepanov, S. K. Vascular plants of Russia and adjacent states (the former USSR). 1995 (L USSR)
  • Huxley, A., ed. The new Royal Horticultural Society dictionary of gardening. 1992 (Dict Gard)
  • Mathew, B. The Iris. 1981 (Iris)
  • Tutin, T. G. et al., eds. Flora europaea. 1964–1980 (F Eur)
  • Walters, S. M. et al., eds. European garden flora. 1986– (Eur Gard F)

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