Emuellidae

Emuellidae are a small family of trilobites, a group of extinct marine arthropods, that lived during the late Lower Cambrian (late Botomian) of the East Gondwana supercontinent, in what are today South-Australia and Antarctica. Emuellidae can be recognized among trilobites in having a set of unique features. The headshield or cephalon has large genal spines reaching back as far as the 3rd to 6th segment of the thorax. The eye-ridges contact the back of the frontal lobe of the glabella and extend laterally and backwards, roughly parallel to the frontal and lateral rim of the cephalon. There are small, clearly incised pits at the junction between the eye-ridge and the frontal lobe of the cephalic axis (or glabella). The thorax reaches its greatest width at the 6th segment. The frontal part or prothorax consists of 6 segments, with number 5 and 6 fused, and the 6th carrying very large trailing spines. The rear part or opistothorax consists of a variable but extremely large number of seqments (up to 97).[1]

Emuellidae Temporal range: 517 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓ late Botomian
Balcoracania dailyi  of the family Emuellidae Lower Cambrian Emu Shale Kangaroo Island, South Australia © Dave Simpson
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Trilobita
Order:	Redlichiida
Suborder:	Redlichiina
Superfamily:	Emuelloidea
Family:	Emuellidae Pocock, 1970
Genera
Emuella Pocock, 1970 Balcoracania Pocock, 1970

Description

(See the Trilobite article for a definition of morphological terms)

Cephalon: Cranidium subquadrate, glabella cylindrical, slightly contracted at S3, three pairs of glabellar furrows, preglabellar field short or absent, eye ridge wide, long, directed slightly postero-laterally, palpebral lobe cresentic, posterior area of fixigena with fulcrum, free cheeks (or librigenae) with long spines; hypostome conterminant, attached to a narrow rostral plate.

The thorax is divided into a prothorax of six segments (the 6th carrying very large, trailing, pleural spines and extremely long opisthothorax of up to 97 segments (Balcoraciana dailyi holds the record for greatest number of thoracic segments in a trilobite species).

Pygidium: A minute, segmented disc.

Taxonomic history

Fossils now assigned to the Emuellidae were first discovered by Dr. B. Daily, of the Geology Department, University of Adelaide in 1956.[2]

Position of the Emuellidae within the Redlichiida

Originally, the Emuellidae were described as part of the Redlichiina. The primitive features prompted the theory that the Emuellids actually represented the stem group of all trilobites, with the Olenellina having secondary fused facial sutures.[2] Later, the Emuellidae were placed in their own superfamily (Emuelloidea), recognizing that trilobites lacking facial sutures (i.e. the Olenellina) are the stem group.[3] This was followed by the clustering of the Emuellidae in a new superfamily Ellipsocephaloidea.[4] Most recently, the Emuellidae are regarded an early branch of the Redlichiina suborder, the closest relatives being the genera Bigotina, Abadiella, and the close-knit group of the families Estaingiidae, Ichangiidae and Ellipsocephalidae.[1]

Genera assigned to the Emuellidae

Holyoakia has previously been assigned to the Emuellidae.[5] The tailshiel (or pygidium) in Holyoakia is about the same size as its cranidium, with a well-defined axis, eight axial rings, well-developed pleural ribs and furrows, and a spiny margin. The pygidia of Emuella and Balcoracania however are poorly differentiated, minute, and have a smooth margin. Later scholars therefore placed Holyoakia in the Dorypygidae.[1]

Distribution

Balcoracania dailyi occurs in the late Lower Cambrian (late Botomian) of South Australia (White Point conglomerate, Cape d' Estaing and Emu Bay sections, Kangaroo Island;[6] Warragee Member, Billy Creek Formation, Flinders Range;[6][7] Coads Hill Member, Billy Creek Formation, Reaphook Hill).[8] Balcoracania sp. has been collected from the Lower Cambrian of Antarctica (Shackleton Limestone, central Transantarctic Mountains).[5]

Emuella dalgarnoi is found in the late Lower Cambrian (late Botomian) of South Australia (Emu Bay Shale, Kangaroo Island).[6]

Emuella polymera has been collected from the late Lower Cambrian (late Botomian) of South Australia (Cape d' Estaing section, Kangaroo Island).[6]

Key to the species

1	Cephalon subpentagonal. Glabella contacts the frontal margin of the cephalon. Axis at the 3rd thorax segment much wider than each of the pleural zones. Up to 58 thorax segments. → 2
-	Cephalon semicircular. There is a short distance between the glabella and the frontal margin of the cephalon. Axis at the 3rd thorax segment almost as wide as each of the pleural zones. Up to 103 thorax segments. South Australia and Antarctica. → Balcoracania dailyi Pocock, 1970
2	Rear border of the cephalon narrower between the midline and the intergenal angle than between the intergenal angle and the genal angle. South Australia. → Emuella polymera Pocock, 1970
-	Rear border of the cephalon wider between the midline and the intergenal angle than between the intergenal angle and the genal angle. South Australia. → Emuella dalgarnoi Pocock, 1970

References

1. Paterson, J.R.; Edgecombe, G.D. (2006). "The Early Cambrian trilobite Family Emuellidae Popock, 1970: Systematic position and revision of Australian Species". Journal of Paleontology. 85: 496–513.
2. Pocock, K.J. (1970). "The Emuellidae, a new family of trilobites from the Lower Cambrian of South Australia". Palaeontology. 13: 522–562.
3. Zhang, W.T.; Lu, Y.H.; Zhu, Z.L.; Qian, Y.Y.; Li, H.L.; Zhou, Z.Y.; Zhang, S.G.; Yuan, J.L. (1980). "Cambrian trilobite faunas of Southwestern China". Palaeontologica Sinica. 159: 1–497.
4. Lin, T.R. (1990). "Application of custer analysis to taxonomy of superfamilies and families of superorder Redlichiina (Trilobita)". Acta Palaeontologica Sinica. 29: 94–105.
5. Palmer, A.R.; Rowell, A.J. (1995). "Early Cambrian trilobites from the Shackleton Limestone of the Central Transantarctic Mountains". Paleontological Society Memoir. 45: 1–28.
6. Popock, K.J. (1970). "The Emuellidae, a new family of trilobites from the Lower Cambrian of South Australia". Palaeontology. 13: 522–562.
7. Moore, P.S. (1979). "Stratigraphy and depositional environments of the Billy Creek Formation (Cambrian), central and northern Flinders Range, South Australia". Transactions of the Royal Society of South Australia. 103: 197–211.
8. Moore, P.S. (1980). "Stratigraphy and depositional environments of the Billy Creek Formation (Cambrian), east of the Flinders Range, South Australia". Transactions of the Royal Society of South Australia. 104: 117–132.

External links

• Hagadorn, J.W. (2002). "Burgess Shale-type Localities: The global picture". In Bottjer, D.J.; W. Etter; J.W. Hagadorn; C.M. Tang (eds.). Exceptional Fossil Preservation -- A Unique View on the Evolution of Marine Life. Columbia University Press. ISBN 0231102542.
• Greg Edgecombe & the Australian Museum. "Australian Trilobites: A Species List and Bibliography". Retrieved August 23, 2005.
• Sam Gon III. "A guide to the Orders of Trilobites". Retrieved August 23, 2005.
• Nedin, C. (1995). "The Emu Bay Shale, a Lower Cambrian fossil Lagerstätte, Kangaroo Island, South Australia". Memoirs of the Association of Australasian Palaeontologists. 18: 31–40.
• Simpson, Dave. "Trilobites of South Australia". Archived from the original on February 19, 2006. Retrieved August 23, 2005.

• photographs of Balcoracania dailyi
• photographs and line-drawing of Emuella polymera