Alcobaça Formation

The Alcobaça Formation is a geological formation in Portugal. It dates back to the Oxfordian stage of the Late Jurassic.[1] It is an important source of information on the diversity of Late Jurassic mammals.[2][3] Many of the fossils were collected from the now disused and flooded Camadas de Guimarota coal mine.

Alcobaça Formation Stratigraphic range: Oxfordian-Tithonian ~160–145 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Type	Geological formation
Sub-units	Camadas de Guimarota
Underlies	Lourinhã Formation
Overlies	Montejunto Formation
Lithology
Primary	Marl
Other	Mudstone, coal, siltstone, limestone, sandstone
Location
Coordinates	39.6°N 9.0°W / 39.6; -9.0
Approximate paleocoordinates	33.7°N 1.6°E / 33.7; 1.6
Region	Estremadura, Leiria, Lisboa
Country	Portugal
Extent	Lusitanian Basin
Type section
Named for	Alcobaça, Portugal
Named by	Choffat
Year defined	1885
Alcobaça Formation (Portugal)

Vertebrate paleofauna

Dinosaur eggs are geographically located in Lisbon District, Portugal.[4] Dinosaur tracks are geographically located in Leiria District, Portugal.[5]

Amphibians

Amphibians of the Alcobaça Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Celtedens	Indeterminate	Porto Pinheiro

Ornithischians

Indeterminate euornithopod remains located in Lisbon District.[4] Indeterminate stegosaurid remains present in Lisbon District.[4]

Ornithischians of the Alcobaça Formation
Genus	Species	District	Stratigraphic position	Abundance	Notes	Images
Dacentrurus[4]	D. armatus[4]	Lisboa[4]
Ornithopoda[4]	Indeterminate[4]	Lisboa[4]
Trimucrodon[4]	T. cuneatus[4]	Lisboa[4]			Later determined to be indeterminate ornithischian remains.[4]

Saurischians

Indeterminate sauropod remains located in Leiria and Lisboa.[6]

Saurischians of the Alcobaça Formation
Genus	Species	District	Stratigraphic position	Abundance	Notes	Images
Allosaurus[5]	Indeterminate[5]	Leiria[5]
Lusotitan[4]	L. atalaiensis[4]	Lisboa[4]
Ceratosaurus[4]	Indeterminate[4]	Lisboa[4]
Dinheirosaurus[4]	D. lourinhanensis[4]	Lisboa[4]		"[Two] cervicals, [nine] dorsals, ribs, limb elements."[7]
Lourinhasaurus[4]	L. alenquerensis[4]	Lisboa[4]
Megalosaurus[4]	M. insignis[4]	Lisboa[4]			Later determined to be indeterminate theropod remains.[4]

Turtles

Turtles of the Alcobaça Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Peltochelys	P. duchastelli	Guimarota
Platychelys	Indeterminate	Guimarota
Pleurosternidae	Indeterminate	Guimarota

Lepidosauromorphs

Lepidosauromorphs of the Alcobaça Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Becklesius	B. hoffstetteri	Guimarota
Dorsetisaurus	D. purbeckensis	Guimarota
Marmoretta	Indeterminate	Guimarota			Basal Lepidosauromorph
Portugalophis	P. lignites	Guimarota			Stem-snake
Saurillodon	S. proraformis	Guimarota
Saurillus	S. henkeli, cf. obtusus	Guimarota

Crocodyliformes

Crocodyliformes of the Alcobaça Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Goniopholis	G. baryglyphaeus	Guimarota
Knoetschkesuchus	K. guimarotae	Guimarota
Lisboasaurus	L. mitracostatus	Guimarota
Machimosaurus	M. hugii	Guimarota

Mammals

Mammals of the Alcobaça Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Bathmochoffatia	B. hapax	Guimarota
Bernardodon	B. sp, B. atlanticus	Porto Pinheiro
Drescheratherium	D. acutum	Guimarota
Dryolestes	D. leirensis	Guimarota
Ecprepaulax	E. anomala	Porto Pinheiro
Guimarotodon	G. leiriensis	Guimarota
Guimarotodus	G. inflatus	Guimarota			Dryolestidae
Haldanodon	H. exspectatus	Guimarota
Henkelodon	H. naias, H. sp	Guimarota
Henkelotherium	H. guimarotae	Guimarota
Iberodon	I. quadrituberculatus	Porto Pinheiro
Kielanodon	K. hopsoni	Guimarota
Krebsotherium	K. lusitanicum	Guimarota
Kuehneodon	K. guimarotensis, K. dietrichi, K. simpsoni, K. uniradiculatus, K. sp	Guimarota
Laolestes	L. andresi	Porto Pinheiro
Meketibolodon	M. robustus	Guimarota
Meketichoffatia	M. krausei, M. sp	Guimarota
Nanolestes	N. krusati	Porto Pinheiro
	N. drescherae	Guimarota
Paulchoffatia	P. delgadoi	Guimarota
Pinheirodon	P. sp, P. vastus, P. pygmaeus	Porto Pinheiro
Plesiochoffatia	P. thoas, P. peperethos	Guimarota
Portopinheirodon	P. asymmetricus	Porto Pinheiro			A dryolestid
Priacodon	Indeterminate	Porto Pinheiro
Proalbionbaatar	P. plagiocyrtus	Guimarota
Pseudobolodon	P. dryas, P krebsi, P. oreas	Guimarota
Renatodon	R. amalthea	Guimarota			A paulchoffatiid
? Tinodon	Indeterminate	Porto Pinheiro
Xenachoffatia	X. oinopion	Guimarota
Paulchoffatiidae	Indeterminate	Guimarota			4 distinct genera with 7 species
Pinheirodontidae	Indeterminate	Porto Pinheiro

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon		Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

See also

• List of dinosaur-bearing rock formations
• List of fossiliferous stratigraphic units in Portugal

References

1. Weishampel et al., 2004, pp.545–549
2. "Guimarota". Paleobiology Database. Retrieved 31 August 2018.
3. "Porto Pinheiro, Lourinha". Paleobiology Database. Retrieved 31 August 2018.
4. "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
5. "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
6. "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," and "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Pages 548-549.
7. "Table 13.1," in Weishampel, et al. (2004). Page 265.

Bibliography

• Weishampel, David B.; Peter Dodson, and Halszka (eds.) Osmólska. 2004. The Dinosauria, 2nd edition, 1–880. Berkeley: University of California Press. Accessed 2019-02-21. ISBN 0-520-24209-2