Harmonia (beetle)

Harmonia
	Asian lady beetle, Harmonia axyridis
	Harmonia testudinaria
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Coccinellidae
Subfamily:	Coccinellinae
Genus:	Harmonia; Mulsant, 1846
Synonyms
	Ballia Mulsant, 1853; Callineda Crotch, 1871; Leis Mulsant, 1850; Ptychanatis Crotch, 1874; Rhopaloneda Timberlake, 1943; Stictoleis Crotch, 1874;

Harmonia is a genus of lady beetles belonging to the subfamily Coccinellinae.

Description

The distinction of the individual species within the genus is often very difficult, and in some cases requires examination of the reproductive system. The coloring and drawing in the genus Harmonia are very variable, not only between species, but also within each species. The basic color of the elytra is usually red to orange with black spots. The patches can be linked together to form black bars. The species can reach a length of 3 to 7 mm. The rear edge of the pronotum is slightly curved. The antennae are a little shorter than the head and are composed of eleven segments. The three last segments are thickened at the tip and form a club.

List of species

Harmonia axyridis (Pallas, 1771) – Asian lady beetle, originally from Nepal, China, Taiwan, Japan, introduced to Western Europe and America
Harmonia antipoda (Boisduval, 1835) – antipodean ladybird, endemic to New Zealand
Harmonia basinotata Bielawski, 1964 – New Guinea
Harmonia bicolor (Blackburn, 1892) – Australia
Harmonia conformis (Boisduval, 1835) – common spotted lady beetle, Australia, introduced in New Zealand
Harmonia dimidiata (Fabricius, 1781) – India, Pakistan, Nepal, Bhutan, China, Taiwan, Japan; introduced in North America
Harmonia dunlopi (Crotch, 1874) – India, Malaysia, Philippines
Harmonia eucharis (Mulsant, 1850) – India, Pakistan, Myanmar, Nepal, Himalayan Region, South China
Harmonia expallida (Weise, 1907) – India
Harmonia octomaculata (Fabricius, 1781) – India, Pakistan, Nepal, Bangladesh, Sri Lanka, Micronesia, Australia
Harmonia quadripunctata (Pontoppidan, 1763) – cream-streaked lady beetle, Europe
Harmonia sedecimnotata (Fabricius, 1801) – India, Nepal, Southeast Asia
Harmonia testudinaria (Mulsant, 1850) – Indonesia, New Guinea, North Australia
Harmonia uninotata Bielawski, 1964 – New Guinea
Harmonia yedoensis (Takizawa, 1917) – Japan

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References

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