Flatidae

Flatidae are a family of fulgoroid planthoppers. They are cosmopolitan in distribution and are distinguished from others in the superfamily by a combination of characters. Like all other planthoppers, they suck phloem sap of plants. Some species are known to communicate with vibrations through the plant stems.[1] Communication may be with mates, or with ants that tend the nymphs, protecting them and gathering honeydew secretions.[2] Adults of some species have brightly coloured forewings which are tougher and known as tegmina unlike the membranous hindwings which are used for flight. Although a few can be identified by their coloration, most species requires dissection and examination under a microscope with access to literature on already described species.[3]

Flatidae
adult bugs and nymphs (Phromnia rosea)
Ankarana Reserve, Madagascar
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hemiptera
Infraorder: Fulgoromorpha
Superfamily: Fulgoroidea
Family: Flatidae
Subfamilies
  • Flatoidinae
  • Flatinae
Flatid nymph from Assam, India
Flatid adult from St. Louis, Missouri
Ormenoides venusta nymph

There are two sub-families within the family. In the subfamily Flatinae, the body of adults is flattened laterally and the tegmina are tent-like. In the Flatoidinae, the body is not laterally compressed and the tegmina are not as tent-like and sometimes held horizontally. The wing venation is distinctive in that the veins in the anal region are nodose, and the costal area has numerous cross veins.[4] The antennae are small and the first segment is collar-like and small. The second segment is longer and ends in a bulge and a flagellum arises from it. They have two ocelli. Nymphs have a tail of waxy filaments.

Genera

Genera within the family Flatidae include:[5]

  • Acanthoflata Fennah, 1947
  • Acrophaea Melichar, 1901
  • Acutisha Medler, 1991
  • Adelidoria Metcalf, 1952
  • Adexia Melichar, 1901
  • Aflata Melichar, 1901
  • Afrexoma Fennah, 1976
  • Afrocyarda Fennah, 1965
  • Afrodascalia Fennah, 1958
  • Afrophantia Fennah, 1958
  • Afrormenis Fennah, 1958
  • Afroseliza Fennah, 1961
  • Alcaxor Fennah, 1947
  • Amasha Medler, 1992
  • Anadascalia Melichar, 1923
  • Anatracis Fennah, 1958
  • Anaya Distant, 1906
  • Anggira Distant, 1906
  • Anidora Melichar, 1901
  • Anormenis Melichar, 1923
  • Anthoflata Fennah, 1947
  • Antillormenis Fennah, 1942
  • Anzora Medler, 1986
  • Aphanophantia Kirkaldy, 1906
  • Apolexis Jacobi, 1936
  • Arelata Stål, 1862
  • Atella Stål, 1866
  • Atracis Stål, 1866
  • Atracodes Melichar, 1902
  • Aulophorina Strand, 1928
  • Austrodascalia Fletcher, 1988
  • Bahuflata Dlabola, 1979
  • Barsac Fletcher, 1988
  • Betracis Medler, 1988
  • Bochara Distant, 1906
  • Boretsis Medler, 1996
  • Brysora Medler, 2000
  • Budginmaya Fletcher & Moir, 2009
  • Burnix Medler, 1988
  • Byllis Stål, 1866
  • Byllisana Metcalf & Bruner, 1948
  • Bythopsyrna Melichar, 1901
  • Caesonia Stål, 1866
  • Calauria Stål, 1866
  • Cameruniola Strand, 1928
  • Capistra Fennah, 1947
  • Carthaeomorpha Melichar, 1901
  • Catracis Medler, 1988
  • Cenestra Stål, 1862
  • Cerfennia Stål, 1870
  • Cerynia Stål, 1862
  • Chaetormenis Melichar, 1923
  • Chaturbuja Distant, 1906
  • Circumdaksha Distant, 1910
  • Cisatra Melichar, 1923
  • Colgar Kirkaldy, 1900
  • Colgaroides Distant, 1910
  • Colobesthes Amyot & Serville, 1843
  • Comnar Medler, 1988
  • Conflata Schmidt, 1912
  • Copsyrna Stål, 1862
  • Cromgar Medler, 2000
  • Cromna Walker, 1857
  • Cromnella Fennah, 1969
  • Cryomna Medler, 2000
  • Cryptobarsac Fletcher & Moir, 2002
  • Cryptoflata Melichar, 1901
  • Cyarda Walker, 1858
  • Cyphopterum Melichar, 1905
  • Daeda Banks, 1910
  • Daksha Distant, 1906
  • Dakshiana Metcalf & Bruner, 1948
  • Dalapax Amyot & Serville, 1843
  • Danavara Distant, 1906
  • Dascalia Stål, 1862
  • Dascalina Melichar, 1902
  • Dascaliomorpha Melichar, 1923
  • Dascanga Medler, 2000
  • Decipha Medler, 1988
  • Delostenopium Jacobi, 1928
  • Demina Medler, 2000
  • Dendrona Melichar, 1923
  • Deocerus Metcalf & Bruner, 1948
  • Derisa Melichar, 1901
  • Dermoflata Melichar, 1901
  • Desanta Medler, 2000
  • Diastracis Medler, 1988
  • Doria Melichar, 1901
  • Doriana Metcalf, 1952
  • Dworena Medler, 1986
  • Epormenis Fennah, 1945
  • Erotana Medler, 2000
  • Eudascalia Melichar, 1923
  • Eugyaria Synave, 1962
  • Euhyloptera Fennah, 1945
  • Eumelicharia Kirkaldy, 1906
  • Euphanta Melichar, 1901
  • Eurima Melichar, 1901
  • Eurocalia Van Duzee, 1907
  • Eurocerus Metcalf, 1945
  • Euryphantia Kirkaldy, 1906
  • Euryprosthius Karsch, 1890
  • Exoma Melichar, 1902
  • Falcophantis Fletcher, 1988
  • Farona Melichar, 1901
  • Flata Fabricius, 1798
  • Flataloides 1938
  • Flatarina Metcalf & Bruner, 1948
  • Flatarissa Metcalf & Bruner, 1948
  • Flatida White, 1846
  • Flatidissa Metcalf, 1952
  • Flatina Melichar, 1901
  • Flatiris Fennah, 1947
  • Flatoides Guérin-Méneville, 1844
  • Flatoidessa Melichar, 1923
  • Flatoidinus Melichar, 1923
  • Flatomorpha Melichar, 1901
  • Flatopsis Melichar, 1902
  • Flatoptera Melichar, 1901
  • Flatormenis Melichar, 1923
  • Flatosaria Melichar, 1923
  • Flatosoma Melichar, 1901
  • Flatula Melichar, 1901
  • Franciscus Distant, 1910
  • Gaja Distant, 1906
  • Garanta Medler, 2000
  • Geisha Kirkaldy, 1900
  • Geraldtonia Distant, 1910
  • GiseliaHaupt, 1956
  • Gomeda Distant, 1906
  • Grapaldus Distant, 1914
  • Gyaria Stål, 1862
  • Gyariella Schmidt, 1924
  • Gyarina Melichar, 1901
  • Hameishara Linnavuori, 1973
  • Hansenia Melichar, 1901
  • Hesperophantia Kirkaldy, 1904
  • Hilavrita Distant, 1906
  • Humgar Medler, 2000
  • Hyphancylus Fowler, 1904
  • Hypsiphanta Jacobi, 1928
  • Idume Stål, 1866
  • Ijagar Medler, 2000
  • Ilesia Fennah, 1942
  • Insulume Medler, 1999
  • Ityraea Stål, 1866
  • Jamella Kirkaldy, 1906
  • Juba Jacobi, 1901
  • Karrama Medler, 1988
  • Kayania Distant, 1910
  • Ketumala Distant, 1906
  • Lasura Medler, 1992
  • Latois Stål, 1866
  • Lawana Distant, 1906
  • Lechaea Stål, 1866
  • Lecopia Medler, 1991
  • Lemaria Medler, 1988
  • Leptodascalia Melichar, 1923
  • Leptormenis Melichar, 1923
  • Lesabes Medler, 1988
  • Lichena Melichar, 1901
  • Lichenopsis Schmidt, 1912
  • Locrona Fennah, 1945
  • Madoxychara Stroinski & Swierczewski, 2013
  • Malleja Medler, 1990
  • Massila Walker, 1862
  • Melicharia Kirkaldy, 1900
  • Melichitona Metcalf, 1952
  • Melormenis Metcalf, 1938
  • Melormenoides Metcalf, 1954
  • Menora Medler, 1999
  • Mesophantia Melichar, 1901
  • Metcalfa Caldwell & Martorell, 1951
  • Meulona Zia, 1935
  • Microflata Melichar, 1901
  • Mimophantia Matsumura, 1900
  • Miniscia Medler, 1991
  • Mistaria Medler, 1988
  • Mistharnophantia Kirkaldy, 1907
  • Monoflata Melichar, 1923
  • Mosiona Melichar, 1923
  • Nakta Distant, 1906
  • Narowalenus Shakila, 1991
  • Neocalauria Synave, 1957
  • Neocerus Melichar, 1901
  • Neocromna Distant, 1910
  • Neodaksha Distant, 1910
  • Neoflata Melichar, 1923
  • Neomelicharia Kirkaldy, 1903
  • Neosalurnis Distant, 1910
  • Neosephena Medler, 2000
  • Neovariata Shakila, 1984
  • Nephesa Amyot & Serville, 1843
  • Nivalios Zhang, Peng & Wang, 2011
  • Nullina Medler, 1991
  • Ormenana Metcalf & Bruner, 1948
  • Ormenaria Metcalf & Bruner, 1948
  • Ormenina Fennah, 1947
  • Ormenis Stål, 1862
  • Ormenoflata Melichar, 1923
  • Ormenoides Melichar, 1923
  • Ortracis Medler, 1996
  • Oryxa Melichar, 1901
  • Panormenis Melichar, 1923
  • Papuanella Distant, 1914
  • Paracalauria Synave, 1962
  • Paracromna Melichar, 1901
  • Paradaksha Distant, 1910
  • Paradascalia 1938
  • Paraflata Melichar, 1901
  • Paraflatoides Melichar, 1923
  • Paraflatoptera Lallemand, 1939
  • Paragomeda Distant, 1914
  • Paraketumala Distant, 1912
  • Paranotus Karsch, 1890
  • Paraseliza Melichar, 1923
  • Parasiphanta Fletcher, 1988
  • Paratella Melichar, 1901
  • Paroxychara Lallemand & Synave, 1951
  • Parthenormenis Fennah, 1949
  • Pauliana Lallemand, 1950
  • Perinetella Synave, 1956
  • Persepolia Dlabola & Safavi, 1972
  • Petrusa Stål, 1862
  • Phaedolus Karsch, 1890
  • Phaiophantia Lindberg, 1958
  • Phalaenomorpha 1843
  • Phantia Fieber, 1866
  • Phantiopsis Melichar, 1905
  • Phlebopterum Stål, 1854
  • Phromnia Stål, 1862
  • Phylliana Metcalf, 1952
  • Phyllodryas Kirkaldy, 1913
  • Phyllyphanta Amyot & Serville, 1843
  • Phymoides Distant, 1910
  • Planata Medler, 1999
  • Planodascalia Metcalf & Bruner, 1948
  • Poeciloflata Melichar, 1901
  • Poeciloptera Latreille, 1804
  • Poekilloptera Latreille, 1796
  • Porophloeus Melichar, 1902
  • Psenoflata Fennah, 1947
  • Pseudodascalia Melichar, 1923
  • Pseudoflatoides Metcalf, 1938
  • Pseudoryxa Schmidt, 1904
  • Pseudoseliza Peng, Wang & Zhang, 2010
  • Pulaha Distant, 1906
  • Pulastya Distant, 1906
  • Rabocha Melichar, 1923
  • Rhinophantia Melichar, 1901
  • Rhynchopteryx Van Duzee, 1914
  • Ricanoflata Melichar, 1923
  • Riculiflata Fennah, 1947
  • Riodeorolix Lindberg, 1956
  • Sabaethis Jacobi, 1916
  • Salurnis Stål, 1870
  • Samcerus Medler, 1993
  • Sanurus Melichar, 1901
  • Satapa Distant, 1906
  • Saurana Medler, 1992
  • Scarpanta Stål, 1862
  • Scarpantina Melichar, 1901
  • Scarposa Uhler, 1895
  • Scarpuna Medler, 2006
  • Seliza Stål, 1862
  • Sephena Melichar, 1901
  • Shadaka Medler, 2000
  • Siphanta Stål, 1862
  • Siscia Stål, 1870
  • Soares Stroinski & Swierczewski, 2012
  • Somisha Medler, 1991
  • Sosephena Medler, 1990
  • Staliana Medler, 1988
  • Stenocyarda Fennah, 1965
  • Stenume Medler, 1999
  • Summanus Distant, 1916
  • Talopsus Medler, 1989
  • Taparella Medler, 1989
  • Tejasa Distant, 1906
  • Tetraceratium Muir, 1924
  • Tisia Dlabola, 1981
  • Tormenis Medler, 1999
  • Trisephena Medler, 1990
  • Ulundia Distant, 1910
  • Umidena Medler, 1992
  • Unnata Distant, 1906
  • Urana Melichar, 1901
  • Utakwana Distant, 1914
  • Uxantis Stål, 1870
  • Uysanus 1908
  • Walena Medler, 1999
  • Zarudnya Melichar, 1901
  • Zecheuna Zia, 1935
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References

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