Eviota

Eviota is a genus of fish in the family Gobiidae, commonly as dwarfgobies found in the Indo-Pacific region, where it is distributed from Japan to Australia and from Africa to Pitcairn Island.[1][2] Species are mainly associated with coral reefs.[2] Many of these fish are short-lived, with life cycles as brief as 3.5 weeks in the tropics. Some species are hermaphrodites and some representatives live symbiotically among the tentacles of the mushroom coral.[2][3][4]

Eviota
Eviota albolineata
Eviota epiphanes
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Gobiiformes
Family:	Gobiidae
Genus:	Eviota O. P. Jenkins, 1903
Type species
Eviota epiphanes O. P. Jenkins, 1903
Synonyms
Allogobius Waite, 1904 Eviotops Smith, 1957

Species

There are currently 113 recognized species in this genus:[5]

• Eviota abax (D. S. Jordan & Snyder, 1901)
• Eviota afelei D. S. Jordan & Seale, 1906 (Afele's dwarfgoby)
• Eviota albolineata S. L. Jewett & Lachner, 1983 (Spotted fringe-fin dwarfgoby) [6]
• Eviota algida D. W. Greenfield & Erdmann, 2014 (Upwelling dwarfgoby) [7]
• Eviota ancora D. W. Greenfield & T. Suzuki, 2011 [8]
• Eviota aquila D. W. Greenfield & S. L. Jewett, 2014 (Dark dwarfgoby) [9]
• Eviota asymbasia D. W. Greenfield & S. L. Jewett, 2016 (Inconsistent dwarfgoby) [10]
• Eviota atriventris D. W. Greenfield & T. Suzuki, 2012 (Black-belly dwarfgoby) [11]
• Eviota bifasciata Lachner & Karnella, 1980 (Two-stripe dwarfgoby)
• Eviota bilunula D. W. Greenfield & T. Suzuki, 2016 (Crescent dwarfgoby) [12]
• Eviota bimaculata Lachner & Karnella, 1980 (Two-spot fringe-fin dwarfgoby)
• Eviota bipunctata D. W. Greenfield & S. L. Jewett, 2016 [10]
• Eviota brahmi D. W. Greenfield & Tornabene, 2014 (Brahm's dwarfgoby) [13]
• Eviota cometa S. L. Jewett & Lachner, 1983 (Comet dwarfgoby)
• Eviota deminuta Tornabene, Ahmadia & J. T. Williams, 2013 (Diminutive dwarfgoby) [1]
• Eviota disrupta Karnella & Lachner, 1981
• Eviota distigma D. S. Jordan & Seale, 1906 (Distigma dwarfgoby)
• Eviota dorsimaculata Tornabene, Ahmadia & J. T. Williams, 2013 (Dorsal-spot dwarfgoby) [1]
• Eviota dorsogilva D. W. Greenfield & J. E. Randall, 2011 (Cream-back dwarfgoby) [14]
• Eviota dorsopurpurea D. W. Greenfield & J. E. Randall, 2011 (Purple dwarfgoby) [14]
• Eviota epiphanes O. P. Jenkins, 1903 (Divine dwarfgoby)
• Eviota epistigmata D. W. Greenfield & S. L. Jewett, 2014 (Twin-spot dwarfgoby) [9]
• Eviota erdmanni Tornabene & D. W. Greenfield, 2016 (Erdmann's dwarfgoby) [15]
• Eviota eyreae D. W. Greenfield & J. E. Randall, 2016 (Eyre's dwarfgoby) [16]
• Eviota fallax D. W. Greenfield & G. R. Allen, 2012 (Twin dwarfgoby) [17]
• Eviota fasciola Karnella & Lachner, 1981 (Barred dwarfgoby)
• Eviota filamentosa T. Suzuki & D. W. Greenfield, 2014 (Thread-fin dwarfgoby) [18]
• Eviota flavipinnata T. Suzuki, D. W. Greenfield & Motomura, 2015 (Yellow-fin dwarfgoby) [19]
• Eviota flebilis D. W. Greenfield, T. Suzuki & Shibukawa, 2014 (Tearful dwarfgoby) [20]
• Eviota geminata D. W. Greenfield & Bogorodsky, 2014 (Geminate dwarfgoby) [21]
• Eviota guttata Lachner & Karnella, 1978 (Spotted dwarfgoby) [6]
• Eviota herrei D. S. Jordan & Seale, 1906 (Herre's dwarfgoby)
• Eviota hinanoae Tornabene, Ahmadia & J. T. Williams, 2013 (Hinano's dwarfgoby) [1]
• Eviota hoesei A. C. Gill & S. L. Jewett, 2004 (Doug's dwarfgoby)
• Eviota indica Lachner & Karnella, 1980
• Eviota infulata (J. L. B. Smith, 1957) (Epaulet dwarfgoby)
• Eviota inutilis Whitley, 1943 (Chest-spot dwarfgoby)
• Eviota irrasa Karnella & Lachner, 1981 (Unpolished dwarfgoby)
• Eviota japonica S. L. Jewett & Lachner, 1983
• Eviota jewettae D. W. Greenfield & R. Winterbottom, 2012 (Jewett's dwarfgoby) [22]
• Eviota karaspila D. W. Greenfield & J. E. Randall, 2010 (Eastern head-spot dwarfgoby) [23]
• Eviota kermadecensis Hoese & A. L. Stewart, 2012 [2]
• Eviota korechika Shibukawa & T. Suzuki, 2005 (Korechika's dwarfgoby)
• Eviota lachdeberei Giltay, 1933 (Red-light dwarfgoby)
• Eviota lacrimae Sunobe, 1988
• Eviota lacrimosa Tornabene, Ahmadia & J. T. Williams, 2013 (Weeping dwarfgoby) [1]
• Eviota lateritea D. W. Greenfield & R. Winterbottom, 2016 (Laterite dwarfgoby) [24]
• Eviota latifasciata S. L. Jewett & Lachner, 1983 (Brown-banded dwarfgoby)
• Eviota maculibotella D. W. Greenfield & R. Winterbottom, 2016 (Spotted-dick dwarfgoby) [24]
• Eviota masudai Matsuura & Senou, 2006
• Eviota melanosphena D. W. Greenfield & S. L. Jewett, 2016 (Wedge dwarfgoby) [25]
• Eviota melasma Lachner & Karnella, 1980 (Head-spot dwarfgoby)
• Eviota mikiae G. R. Allen, 2001 (White-line dwarfgoby)
• Eviota mimica D. W. Greenfield & J. E. Randall, 2016 (Mimic dwarfgoby) [16]
• Eviota minuta D. W. Greenfield & S. L. Jewett, 2014 (Minute dwarfgoby) [26]
• Eviota monostigma Fourmanoir, 1971 (Single-spot dwarfgoby)
• Eviota natalis G. R. Allen, 2007 (Christmas dwarfgoby)
• Eviota nebulosa J. L. B. Smith, 1958 (Nebulous dwarfgoby)
• Eviota nigramembrana D. W. Greenfield & T. Suzuki, 2013 (Black-bar dwarfgoby) [27]
• Eviota nigripinna Lachner & Karnella, 1980
• Eviota nigrispina D. W. Greenfield & T. Suzuki, 2010 (Black-spine dwarfgoby) [28]
• Eviota nigriventris Giltay, 1933 (Red-belly dwarfgoby) [13]
• Eviota notata D. W. Greenfield & S. L. Jewett, 2012 (Bar-head dwarfgoby) [29]
• Eviota occasa D. W. Greenfield, R. Winterbottom & T. Suzuki, 2014 (Sunset dwarfgoby) [30]
• Eviota ocellifer Shibukawa & T. Suzuki, 2005
• Eviota oculopiperita D. W. Greenfield & Bogorodsky, 2014 (Peppermint-eye dwarfgoby) [21]
• Eviota pamae G. R. Allen, Brooks & Erdmann, 2013 [31]
• Eviota pardalota Lachner & Karnella, 1978 (Leopard dwarfgoby)
• Eviota partimacula J. E. Randall, 2008
• Eviota pellucida Larson, 1976 (Neon dwarfgoby)
• Eviota pinocchioi D. W. Greenfield & R. Winterbottom, 2012 (Pinocchio dwarfgoby) [22]
• Eviota piperata D. W. Greenfield & R. Winterbottom, 2014 (Peppered dwarfgoby) [32]
• Eviota prasina (Klunzinger, 1871) (Green dwarfgoby)
• Eviota prasites D. S. Jordan & Seale, 1906 (Hair-fin dwarfgoby)
• Eviota pseudostigma Lachner & Karnella, 1980
• Eviota punctulata S. L. Jewett & Lachner, 1983 (Dotted dwarfgoby)
• Eviota punyit Tornabene, Valdez & Erdmann, 2016 [33]
• Eviota queenslandica Whitley, 1932 (Queensland dwarfgoby)
• Eviota raja G. R. Allen, 2001 (King dwarfgoby)
• Eviota randalli D. W. Greenfield, 2009 (Randall's dwarfgoby)
• Eviota readerae A. C. Gill & S. L. Jewett, 2004 (Sally's dwarfgoby)
• Eviota richardi D. W. Greenfield & J. E. Randall, 2016 (Rick's dwarfgoby) [16]
• Eviota rubra D. W. Greenfield & J. E. Randall, 1999
• Eviota rubriceps D. W. Greenfield & S. L. Jewett, 2011 (Red-headed dwarfgoby) [34]
• Eviota rubriguttata D. W. Greenfield & T. Suzuki, 2011 [8]
• Eviota rubrimaculata T. Suzuki, D. W. Greenfield & Motomura, 2015 (Red-spot dwarfgoby) [19]
• Eviota rubrisparsa D. W. Greenfield & J. E. Randall, 2010 (Red-speckled dwarfgoby) [6]
• Eviota saipanensis Fowler, 1945
• Eviota santanai D. W. Greenfield & Erdmann, 2013 (Santana's dwarfgoby) [35]
• Eviota sebreei D. S. Jordan & Seale, 1906 (Striped dwarfgoby)
• Eviota shibukawai T. Suzuki & D. W. Greenfield, 2014 (Shibukawa's dwarfgoby) [18]
• Eviota shimadai D. W. Greenfield & J. E. Randall, 2010 (Shimada's dwarfgoby) [6]
• Eviota sigillata S. L. Jewett & Lachner, 1983 (Adorned dwarfgoby)
• Eviota singula D. W. Greenfield & R. Winterbottom, 2016 (One-spot dwarfgoby) [24]
• Eviota smaragdus D. S. Jordan & Seale, 1906 (Emerald dwarfgoby)
• Eviota sodwanaensis D. W. Greenfield & R. Winterbottom, 2016 (Sodwana dwarfgoby) [36]
• Eviota sparsa S. L. Jewett & Lachner, 1983 (Speckled dwarfgoby)
• Eviota specca D. W. Greenfield, T. Suzuki & Shibukawa, 2014 [20]
• Eviota spilota Lachner & Karnella, 1980 (Spotted-fin dwarfgoby)
• Eviota springeri D. W. Greenfield & S. L. Jewett, 2012 (Springer's dwarfgoby) [29]
• Eviota storthynx (Rofen, 1959) (Storthynx dwarfgoby)
• Eviota susanae D. W. Greenfield & J. E. Randall, 1999
• Eviota teresae D. W. Greenfield & J. E. Randall, 2016 (Terry's dwarfgoby) [16]
• Eviota tetha D. W. Greenfield & Erdmann, 2014 (Tetha's dwarfgoby) [37]
• Eviota thamani D. W. Greenfield & J. E. Randall, 2016 (Thaman's dwarfgoby) [16]
• Eviota tigrina D. W. Greenfield & J. E. Randall, 2008
• Eviota toshiyuki D. W. Greenfield & J. E. Randall, 2010 [6]
• Eviota variola Lachner & Karnella, 1980 (Fin-spot dwarfgoby)
• Eviota winterbottomi D. W. Greenfield & J. E. Randall, 2010 (Dusky-bar dwarfgoby) [6]
• Eviota zebrina Lachner & Karnella, 1978 (Zebra dwarfgoby)
• Eviota zonura D. S. Jordan & Seale, 1906 (Zoned dwarfgoby)

References

1. Tornabene, L., Ahmadia, G.N. & Williams, J.T. (2013): Four new species of dwarfgobies (Teleostei: Gobiidae: Eviota) from the Austral, Gambier, Marquesas and Society Archipelagos, French Polynesia. Systematics and Biodiversity, 11 (3): 363–380.
2. Hoese, D.F. & Stewart, A.L. (2012): A new species of the gobiid genus Eviota (Teleostei: Gobioidei) from the Kermadec Islands, New Zealand. Zootaxa, 3418: 61–67.
3. Bos, A.R. (2012): Fishes (Gobiidae and Labridae) associated with the mushroom coral Heliofungia actiniformis (Scleractinia: Fungiidae) in the Philippines. Coral Reefs, 31 (1): 133–133.
4. Bos, A.R. & Hoeksema, B.W. (2014): Cryptobenthic fishes and co-inhabiting shrimps associated with the mushroom coral Heliofungia actiniformis (Fungiidae) in the Davao Gulf, Philippines. Environmental Biology of Fishes, 98 (6): 1479–1489.
5. Froese, Rainer and Pauly, Daniel, eds. (2018). Species of Eviota in FishBase. June 2018 version.
6. Greenfield, D.W. & Randall, J.E. (2010): Four new gobiid fishes of the genus Eviota from the western Pacific, with clarification of Eviota guttata and Eviota albolineata (Teleostei: Gobiidae). Proceedings of the California Academy of Sciences, 61 (3): 269-289.
7. Greenfield, D.W. & Erdmann, M.V. (2014): Eviota algida, a new dwarfgoby species from the upwelling waters off Nusa Penida, Indonesia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 13: 27–34.
8. Greenfield, D.W. & Suzuki, T. (2011): Two new goby species of the genus Eviota from the Ryukyu Islands, Japan (Teleostei: Gobiidae). Zootaxa, 2812: 63–68.
9. Greenfield, D.W. & Jewett, S.L. (2014): Two New Dwarfgobies from the Western Pacific (Teleostei: Gobiidae: Eviota). Copeia, 2014 (1): 56–62.
10. Greenfield, D.W. & Jewett, S.L. (2016): Two new dwarfgobies from the Indian and Western Pacific Oceans (Teleostei: Gobiidae: Eviota). Zootaxa, 4121 (5): 589–599.
11. Greenfield, D.W. & Suzuki, T. (2012): Eviota atriventris, a New Goby Previously Misidentified as Eviota pellucida Larson (Teleostei: Gobiidae). Zootaxa, 3197: 55–62.
12. Greenfield, D.W. & Suzuki, T. (2016): Eviota bilunula, a new dwarfgoby species from Fiji, with a redescription of Eviota flebilis (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 24: 1–9.
13. Greenfield, D.W. & Tornabene, L. (2014): Eviota brahmi n. sp. from Papua New Guinea, with a redescription of Eviota nigriventris (Teleostei: Gobiidae). Zootaxa, 3793 (1): 133–146.
14. Greenfield, D.W. & Randall, J.E. (2011): Two new Indo-Pacific species in the Eviota nigriventris complex (Teleostei: Gobiidae). Zootaxa, 2997: 54–66.
15. Tornabene, L. & Greenfield, D.W. (2016): Eviota erdmanni (Teleostei: Gobiidae), a new dwarfgoby from the Savu Sea, Flores, Indonesia. Journal of the Ocean Science Foundation, 22: 1-9.
16. Greenfield, D.W. & Randall, J.E. (2016): A review of the dwarfgobies of Fiji, including descriptions of five new species (Teleostei: Gobiidae: Eviota). Journal of the Ocean Science Foundation, 20: 25–75.
17. Greenfield, D.W. & Allen, G.R. (2012): Eviota fallax, a New Dwarfgoby from the Western Pacific (Teleostei: Gobiidae). Zootaxa, 3522: 42–48.
18. Suzuki, T. & Greenfield, D.W. (2014): Two new dwarfgobies from the Ryukyu Islands, Japan: Eviota shibukawai and Eviota filamentosa (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 11: 32–39.
19. Suzuki, T., Greenfield, D.W. & Motomura, H. (2015): Two new dwarfgobies (Teleostei: Gobiidae) from the Ryukyu Islands, Japan: Eviota flavipinnata and Eviota rubrimaculata. Zootaxa, 4007 (3): 399–408.
20. Greenfield, D.W., Suzuki, T. & Shibukawa, K. (2014): Two new dwarfgobies of the genus Eviota from the Ryukyu Islands, Japan (Teleostei: Gobiidae). Zootaxa, 3774 (5): 481–488.
21. Greenfield, D.W., Bogorodsky, S.V. & Mal, A.O. (2014): Two new Red Sea dwarfgobies (Teleostei, Gobiidae, Eviota). Journal of the Ocean Science Foundation, 10: 1–10.
22. Greenfield, D.W. & Winterbottom, R. (2012): Two new dwarfgobies from the Southwestern Pacific Ocean (Teleostei: Gobiidae: Eviota). Zootaxa, 3572: 33–42.
23. Greenfield, D.W. & Randall, J.E. (2010): Eviota karaspila, a new gobiid fish from Fiji (Teleostei: Gobiidae). Zootaxa, 2672: 61–68.
24. Greenfield, D.W. & Winterbottom, R. (2016): Three new dwarfgobies from the western Pacific Ocean (Teleostei: Gobiidae: Eviota). Journal of the Ocean Science Foundation, 22: 28-40.
25. Greenfield, D.W. & Jewett, S.L. (2016): Eviota melanosphena, a new dwarfgoby from Australia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 21: 71-77.
26. Greenfield, D.W. & Jewett, S.L. (2014): Eviota minuta, a new dwarfgoby from the Philippine Islands (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 12: 12–17.
27. Greenfield, D.W. & Suzuki, T. (2013): Eviota nigramembrana, a new dwarfgoby from the Western Pacific (Teleostei: Gobiidae). Zootaxa, 3637 (2): 169–175.
28. Greenfield, D.W. & Suzuki, T. (2010): Eviota nigrispina, a New Goby from the Ryukyu Islands, Japan (Teleostei: Gobiidae). Zootaxa, 2655: 57–62.
29. Greenfield, D.W. & Jewett, S.L. (2012): Two new Gobiid fishes of the genus Eviota from the Indian Ocean (Teleostei: Gobiidae). Zootaxa, 3515: 67–74.
30. Greenfield, D.W., Winterbottom, R. & Suzuki, T. (2014): Eviota occasa, a new species of dwarfgoby from Palau and the Ryukyu Islands, Japan (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 10: 11–19.
31. Allen, G.R., Brooks, W.M. & Erdmann, M.V. (2013): Eviota pamae, a new species of coral reef goby (Gobiidae) from Indonesian seas. aqua, International Journal of Ichthyology, 19 (2): 79–84.
32. Greenfield, D.W. & Winterbottom, R. (2014): Eviota piperata, a new gobiid species from Palau (Teleostei: Gobiidae). Zootaxa, 3755 (3): 295–300.
33. Tornabene, L., Valdez, S., Erdmann, M.V. & Pezold, F.L. (2016): Multi-locus sequence data reveal a new species of coral reef goby (Teleostei: Gobiidae: Eviota), and evidence of Pliocene vicariance across the Coral Triangle. Journal of Fish Biology, 88 (5): 1811–1834.
34. Greenfield, D.W. & Jewett, S.L. (2011): Eviota rubriceps, a new goby from the Southwestern Pacific Ocean, with comments on E. mikiae and E. raja (Teleostei: Gobiidae). Zootaxa, 3134: 53–62.
35. Greenfield, D.W. & Erdmann, M.V. (2013): Eviota santanai, a new Dwarfgoby from Timor-Leste (Teleostei: Gobiidae). Zootaxa, 3741 (4): 593–600.
36. Greenfield, D.W. & Winterbottom, R. (2016): Eviota sodwanaensis, a new dwarfgoby from South Africa (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 22: 53–57.
37. Greenfield, D.W. & Erdmann, M.V. (2014): Eviota tetha, a new species of dwarfgoby from Cenderawasih Bay, West Papua, Indonesia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 11: 23–31.