Acuticryphops

Acuticryphops; Temporal range: Frasnian PreꞒ Ꞓ O S D C P T J K Pg N ↓
	Acuticryphops
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Trilobita
Superorder:	Phacopina
Order:	Phacopida
Superfamily:	Phacopoidea
Family:	Phacopidae
Genus:	Acuticryphops; Cronier and Feist, 2000
species
	A. acuticeps (Kayser, 1889) (type) = Phacops acuticeps, Cryphops acuticeps ;

Acuticryphops is a genus of trilobite that lived during the late Frasnian, particularly between the Lower and Upper Kellwasser horizons. It is known from mid-European Avalonia (Harz Mountains and Rhenish Slate Mountains) and Armorica (Thuringia: Schleiz) and the North Gondwana margin (Coumiac, France and central Morocco).[1][2]

Taxonomy

Acuticryphops was probably ancestral to the species of the genus Trimerocephalus, and mainly differs from its supposed eyeless descendants by retaining small eyes.

Evolution

Acuticryphops acuticeps is an example of rapid evolution. Subsequent populations exhibit a declining average and an increasing variation in the number of lenses per eye from about 10 to between 7 and 1. This rapid trend does not occur in other characters of the exoskeleton. This suggests the absence rather than the presence of selective pressure on the number of lenses per eye. This is presumably associated with the eustatic deepening that occurred.[1]

Description

In dorsal view, Acuticryphops acuticeps has a rounded triangular headshield (or cephalon). The central raised area of the cephalon (or glabella) is roughly pentangular, and it substantially overhangs the frontal cephalic border. The cheeks are roughly triangular, with the eyes (or palpabral lobes) occupying the frontal quarter of its length. The eye has between 1 and 11 or more individual lenses. The pygidium is more than 3 times wider than long and is almost rectangular because the posterior thorax segments angle backwards when approaching the side of the exoskeleton.

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References

Crônier, Catherine; Feist, Raimund; Auffray, Jean-Christophe (2004). "Variation in the eye of Acuticryphops (Phacopina, Trilobita) and its evolutionary significance: a biometric and morphometric approach". Paleobiology. 30 (3): 471–481. doi:10.1666/0094-8373(2004)030<0471:viteoa>2.0.co;2. Retrieved 23 December 2013.
Feist, R. (2002). "Trilobites from the latest Frasnian Kellwasser Crisis in North Africa (Mrirt, central Moroccan Meseta)". Acta Palaeontologica Polonica. 47 (2): 203–210. cited in "Trilobites between Lower and Upper Kellwasser events, Mrirt area, Morocco (Devonian of Morocco)". Fossilworks. Retrieved 23 December 2013.

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[CFA-1] Crônier, Catherine; Feist, Raimund; Auffray, Jean-Christophe (2004). "Variation in the eye of Acuticryphops (Phacopina, Trilobita) and its evolutionary significance: a biometric and morphometric approach". Paleobiology. 30 (3): 471–481. doi:10.1666/0094-8373(2004)030<0471:viteoa>2.0.co;2. Retrieved 23 December 2013.

[2] Feist, R. (2002). "Trilobites from the latest Frasnian Kellwasser Crisis in North Africa (Mrirt, central Moroccan Meseta)". Acta Palaeontologica Polonica. 47 (2): 203–210. cited in "Trilobites between Lower and Upper Kellwasser events, Mrirt area, Morocco (Devonian of Morocco)". Fossilworks. Retrieved 23 December 2013.