Cheliderpeton

Cheliderpeton (often misspelled Chelyderpeton) is an extinct genus of temnospondyl amphibian. It lived during the Early Permian in what is now Europe and Germany. Fossils have been found from the Ruprechtice horizon of the Intrasudetic Basin of Bohemia in the Czech Republic, as well as the Saar-Nahe Basin of southwestern Germany. Cheliderpeton had a 16 cm skull, and reached about 65 cm in length.[1]

Life restoration of Cheliderpeton vranyi

Cheliderpeton
Temporal range: Early Permian
Fossil skull of Cheliderpeton varyi from Broumov (Czech Republic) at the National Museum (Prague)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Order: †Temnospondyli
Superfamily: †Archegosauroidea (?)
Genus: †Cheliderpeton
Fritsch, 1877
Type species
†Cheliderpeton vrayni
Fritsch, 1877
Skull of Cheliderpeton

The type species of Cheliderpeton is C. vranyi, named in 1877 by the Czech paleontologist Antonin Fritsch and found from the Czech Republic.[2] A second species named C. latirostre was described in 1993 by J. A. Boy from Germany, after having been assigned to Archegosaurus.[3] It differs from the type in having a less extensive preorbital region of the skull, a wider cheek region, and a snout tip that is less round and has lateral projections. Currently the two species are the only ones known to belong to Cheliderpeton, although there have been more assigned in the past that are now considered to be synonymous with the known species. "Actinodon germanicus", described by O. Kuhn in 1939 on the basis of a specimen found from the Czech Republic,[4][5] It is now known to be an unusually large individual of C. vranyi, probably in a late stage of ontogenic development.[6]

It has been suggested that C. latirostre may belong to a separate genus, either to Archegosaurus or its own distinct taxon, due to several differences between it and the type species. These differences include the concave outline of the skull roof, the expanded snout seen in adult individuals (similar to Archegosaurus decheni), and a nasal-maxilla contact in the adult stage.[6] In 2009, C. latirostre was placed in its own genus, Glanochthon.[7]

Cheliderpeton is closely related to the genus Intasuchus from the northern pre-Ural region of Russia. Both share similarities in skull shape, as well as a short and expanded dorsal branch of the ilium and a premaxilla-maxilla suture that is anterior to the choana.[3] Fossils of temnospondyls similar in appearance to Cheliderpeton have been found from the Autun Basin and the basin of Bourbon l'Archambault in France.[8][9]

Fossil

Cheliderpeton may be a member of the family Archegosauridae or the family Intasuchidae. The family Archegosauridae belongs to the superfamily Archegosauroidea, but it is unsure whether Intasuchus, the type species of Intasuchidae, belongs to Archegosauroidea or the family Eryopidae, within the superfamily Eryopoidea. Thus it is unsure whether Cheliderpeton is an eryopoid (and perhaps more specifically an eryopid) or an archegosauroid. Cheliderpeton had previously been placed in the family Actinodontidae along with Actinodon, Syndyodosuchus and Sclerocephalus,[10] but because Actinodon, the type genus of the family, is a junior synonym of the eryopid Onchiodon, the family is now considered polyphyletic.[6]

Notes and references
  1. http://www.angellis.net/Web/PDfiles/amphs.pdf
  2. Fritsch, A. (1877). "Ueber einen neuen Saurier aus den Kalksteinen der Permformation (U. Dyas) aus Braunau in Böhmen". Verlag der königlich-böhmischen Gesellschaft der Wissenschaften. 1877: 1–3.
  3. Boy, J. A. (1993). "Über einige Vertreter der Eryopidea (Amphibia: Temnospondyli) aus dem europäschen Rotliegend (?höchstes Karbon-Perm). 4. Cheliderpeton latirostre". Paläontologische Zeitschrift. 67 (1/2): 123–143. doi:10.1007/bf02985874.
  4. "A. germanicus", was named during the time when Germany occupied Bohemia, hence the specific name.
  5. Kuhn, O. (1939). "Ein neuer rhachitomer Stegocephale aus dem Unteren Rotliegenden". Paläontologische Zeitschrift. 21: 126–131. doi:10.1007/bf03043250.
  6. Werneburg, R.; Steyer, J. S. (2002). "Revision of Cheliderpeton vranyi Fritsch, 1877 (Amphibia, Temnospondyli) from the Lower Permian of Bohemia (Czech Republic)". Paläontologische Zeitschrift. 76 (1): 149–162. doi:10.1007/BF02988193.
  7. Schoch, R.R.; Witzmann, F. (2009). "The temnospondyl Glanochthon from the Permian Meisenheim Formation of Germany". Special Papers in Palaeontology. 81: 121–136.
  8. Steyer, J. S. (1996). "Une nouvelle espèce de Cheliderpeton (Amphibia, Temnospondyli) du Permien inférieur de Buxières-les-Mines (Allier, France). Position phylétique et relations ontogénie-phylogénie des eryopoides". DEA Report: 31p.
  9. Werenburg, R. (1997). "Der eryopide Onchiodon (Amphibia) aus dem Rotliegend des Beckens yon Autun (Frankreich)". Freiberger Forschungshefte C 466, Paläontologie, Stratigraphie, Fazies - Heft 4: 167–181.
  10. Schoch, R. R. (1997). "Cranial anatomy of the Permian temnospondyl amphibian Zatrachys serratus Cope 1878, and the phylogenetic position of the Zatrachydidae". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 206 (2): 223–248.
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