Melanella

Melanella
	Melanella martinii
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Clade:	Caenogastropoda
Clade:	Hypsogastropoda
Order:	Littorinimorpha
Family:	Eulimidae
Genus:	Melanella; Bowdich, 1822
Synonyms
	Balcis Leach, 1847; Eulima (Balcis); Eulimaustra Laseron, 1955; Melanella (Balcis); Melanella (Melanella) Bowdich, 1822;

Melanella is a genus of very small ectoparasitic sea snails, marine gastropod mollusks or micromollusks in the family Eulimidae.[1]

Species

According to the World Register of Marine Species (WoRMS) the following species with accepted names are included within the genus Melanella[1]

Melanella abida Dall, 1927
Melanella abreojosensis Bartsch, 1917
Melanella acicula (Gould, 1849)
Melanella aciculata (Pease, 1861)
Melanella acuformis (G. Nevill & H. Nevill, 1875)
Melanella adamantina (de Folin, 1867)
Melanella aequatorialis (Thiele, 1925)
Melanella aethiopica (Thiele, 1925)
Melanella aethria (Melvill, 1918)
Melanella alayoi Espinosa, Ortea & Magana, 2001
Melanella alba (da Costa, 1778)
Melanella alertae (Dell, 1956)
Melanella alfredensis Bartsch, 1915
Melanella algoensis E. A. Smith, 1901
Melanella amabilis (Brazier, 1877)
Melanella ameliae Engl, 2007
Melanella anachorea Dall, 1927
Melanella angulata (A. Adams, 1861)
Melanella antarctica (Strebel, 1908)
Melanella archeyi (Finlay, 1928)
Melanella arleyi Espinosa, Ortea & Magana, 2001
Melanella atlantica (E. A. Smith, 1890)
Melanella attenuata (Sowerby II, 1866)
Melanella aucklandica (Suter, 1909)
Melanella augur (Angas, 1865)
Melanella aurata Hirase, 1920
Melanella baldra Bartsch, 1917
Melanella becki (Turton, 1932)
Melanella bermudezi Pilsbry & Aguayo, 1933
Melanella borealis (Bartsch, 1915)
Melanella boscheineni Engl, 2004
Melanella boscii (Payraudeau, 1826)
Melanella bovicornu (Pilsbry, 1905)
Melanella brevis (Requien, 1848)
Melanella brunnimaculata (Kay, 1979)
Melanella bryani Pilsbry, 1918
Melanella callistemma Dall, 1927
Melanella catalinensis Bartsch, 1917
Melanella chariessa (Verrill, 1884)
Melanella charissa (Verrill, 1884) : misspelling of Melanella chariessa (Verrill, 1884)
Melanella chrysallida (A. Adams, 1861)
Melanella chydaea (Watson, 1883)
Melanella cifara Bartsch, 1915
Melanella cinca Dall, 1927
Melanella clavula (A. Adams, 1861)
Melanella compactilis (Monterosato, 1884)
Melanella conoidea (Kurtz & Stimpson, 1851)
Melanella convexa (Smith, 1907)
Melanella corintonis (Hertlein & Strong, 1951)
Melanella cosmia Bartsch, 1917
Melanella costellata (Dautzenberg& Fischer H., 1897)
Melanella cumingi (A. Adams, 1854)
Melanella cunaeformis (May, 1915)
Melanella cuspidata (A. Adams, 1851)
Melanella debilis (A. Adams, 1861)
Melanella decagyra (Melvill & Standen, 1903)
Melanella decipiens (Hedley, 1899)
Melanella densicostata Bouchet & Warén, 1986
Melanella doederleini (Brusina, 1886)
Melanella draconis Bartsch, 1917
Melanella dufresnii Bowdich, 1822
Melanella eburnea (Mühlfeld, 1824)
Melanella elenensis (Bartsch, 1924)
Melanella elsa Thiele, 1930
Melanella eulimoides (C. B. Adams, 1850)
Melanella exasperata (Preston, 1905)
Melanella exilis (Pease, 1863)
Melanella exulata (Smith, 1915)
Melanella falcata (Carpenter, 1865)
Melanella fasciata (Watson, 1883)
Melanella frielei (Jordan, 1895)
Melanella gagei Bouchet & Warén, 1986
Melanella glypta Bouchet & Warén, 1986
Melanella gracilis (C. B. Adams, 1850)
Melanella gradata (Cotton & Godfrey, 1932)
Melanella grippi (Bartsch, 1917)
Melanella guenteri Engl, 2004
Melanella guildingii (A. Adams, 1854)
Melanella hadalis Knudsen, 1964
Melanella halia Bartsch, 1917
Melanella haloraphe (Dautzenberg & Fischer H., 1896)
Melanella hastata (Sowerby I, 1834)
Melanella hebes (Watson, 1883)
Melanella hemphilli Bartsch, 1917
Melanella hypsela (Verrill & Bush, 1900)
Melanella ima Pilsbry, 1918
Melanella indica'' (Preston, 1905)
Melanella insculpta Bouchet & Warén, 1986
Melanella iotoides Bartsch, 1916
Melanella ira Dall, 1927
Melanella jeffreysii (Tryon, 1886)
Melanella junieri Espinosa, Ortea & Magana, 2006
Melanella kahoolawensis Pilsbry, 1917
Melanella kanaka (Pilsbry, 1917)
Melanella karolinae Engl, Winfried, 2006
Melanella kawamurai (Kuroda & Habe, 1950)
Melanella kermadecensis W. R. B. Oliver, 1915
Melanella lastra Bartsch, 1917
Melanella laurae (Friele, 1886)
Melanella letsonae (Pilsbry, 1917)
Melanella levantina (Oliverio, Buzzurro & Villa, 1994)
Melanella lubrica (Monterosato, 1890)
Melanella lucida (Verrill, 1884)
Melanella luminosa Marshall, 1997
Melanella lunata Pilsbry, 1917
Melanella lutea Rodrigez Garcia del Castillo, 2000
Melanella magnifica (Bartsch, 1915)
Melanella maoria (Powell, 1940)
Melanella maracuya Espinosa, Ortea & Magana, 2001
Melanella martinii (A. Adams in Sowerby, 1854)
Melanella martynjordani (Jordan, 1895)
Melanella marviva Espinosa, Ortea, & Magaña, 2006
Melanella maui (Dell, 1952)
Melanella medipacifica (Pilsbry, 1917)
Melanella micans (Carpenter, 1865)
Melanella microsculpta Bouchet & Warén, 1986
Melanella mimus Pilsbry, 1918
Melanella minor (Pallary, 1904)
Melanella modesta Thiele, 1930
Melanella montereyensis Bartsch, 1917
Melanella muelleriae (Sturany, 1904)
Melanella mundula (A. Adams, 1861)
Melanella myriotrochi Bouchet & Warén, 1986
Melanella nilae Engl, Winfried, 2006
Melanella nisonida (Melvill, 1906)
Melanella nutans (Mühlfeld, 1824)
Melanella obtusa (Laseron, 1955)
Melanella obtusoapicata Bouchet & Warén, 1986
Melanella ochsneri (Bartsch, 1917)
Melanella odontoidea (A. Adams, 1861)
Melanella ogasawarana (Pilsbry, 1905)
Melanella oleacea (Kurtz & Stimpson, 1851)
Melanella olssoni (Bartsch, 1924)
Melanella opaca (Sowerby II, 1865)
Melanella opalina (Monterosato MS, Marshall, 1901)
Melanella ophiodon Dall, 1927
Melanella orphanensis Clarke A.H., 1974
Melanella orthophyes (Sturany, 1903)
Melanella otagoensis (Powell, 1927)
Melanella oxyacme (Suter, 1908)
Melanella oxytata (Watson, 1883)
Melanella pandata (A. Adams, 1861)
Melanella parallela Dall, 1927
Melanella paxillus (Hedley, 1904)
Melanella peninsularis (Bartsch, 1917)
Melanella penna Dall, 1927
Melanella pernula (Monterosato, 1890)
Melanella perplexa Oliver, 1915
Melanella persimilis (Kuroda & Habe in Kuroda et al., 1971)
Melanella perversa (Bush, 1909)
Melanella petitiana (Brusina, 1869)
Melanella pisinna Rehder, 1980
Melanella planisutis Bouchet & Warén, 1986
Melanella polita (Linnaeus, 1758)
Melanella praecurta (Pallary, 1904)
Melanella producta (Carpenter, 1863)
Melanella pseudoglabra (Dautzenberg & Fischer H., 1912)
Melanella puhana (Dell, 1956)
Melanella pyramidalis (A. Adams, 1854 in H. & A. Adams 1853–58)
Melanella retexta (Carpenter, 1863)
Melanella reticulata Bouchet & Warén, 1986
Melanella retrorsa (Sowerby, 1865)
Melanella richardi (Dautzenberg & Fischer H., 1896)
Melanella robusta (A. Adams, 1861)
Melanella roegerae (Cotton & Godfrey, 1932)
Melanella rubrotincta (Jeffreys, 1867)
Melanella salvadori Castellanos, Rolán & Bartolotta, 1987
Melanella sankurieae Engl, 2004
Melanella sansibarica (Thiele, 1925)
Melanella sarissa (Watson, 1883)
Melanella sarsi (Bush, 1909)
Melanella scarifata Gofas & Rueda, 2014
Melanella seguenzai Di Geronimo & La Perna, 1999
Melanella shibana (Yokoyama, 1927)
Melanella siberutensis (Thiele, 1925)
Melanella similior Bouchet & Warén, 1986
Melanella sinuosa (Scacchi, 1836)
Melanella solida (Sowerby II, 1865)
Melanella solidula (Adams & Reeve, 1850)
Melanella soliduloides (Habe, 1951)
Melanella solitaria (Smith, 1915)
Melanella spina (Boettger, 1893)
Melanella spinosa Oliver, 1915
Melanella spiridioni (Dautzenberg & Fischer H., 1896)
Melanella stalioi (Brusina, 1869)
Melanella stamina Dall, 1927
Melanella subantarctica (Strebel, 1908)
Melanella subula (A. Adams, 1861)
Melanella teinostoma (A. Adams, 1853)
Melanella teramachii (Habe, 1952)
Melanella thalia Bartsch, 1915
Melanella thersites (Carpenter, 1864)
Melanella tia (Bartsch, 1926)
Melanella tortuosa (Adams & Reeve, 1850)
Melanella townsendi Bartsch, 1917
Melanella translucens (Monterosato, 1890)
Melanella trunca (Watson, 1897)
Melanella truncata (Suter, 1908)
Melanella tumidula (Thiele, 1912)
Melanella turbonilloides Bouchet & Warén, 1986
Melanella turrita Bouchet & Warén, 1986
Melanella vegrandis (Murdoch & Suter, 1906)
Melanella versa Dall, 1927
Melanella xiphidiopsis (Dautzenberg & Fischer, 1896)
Melanella yamazii (Habe, 1952)
Melanella zugnigae Espinosa, Ortea & Magana, 2001

Melanella martinii, abapertural view

Three shells of Melanella solidula

The Indo-Pacific Molluscan Database also includes the following species with names in current use[2]

Melanella australiensis (Thiele, 1930)
Melanella campyla (Watson, 1883)
Melanella carneola (Gould, 1861 in 1859–61)
Melanella cunaeformis (May 1915)
Melanella curvata (A. Adams, 1861-b)
Melanella dentaliopsis (A. Adams, 1861-b)
Melanella flexa (A. Adams, 1861-b)
Melanella flexuosa (A. Adams in H. & A. Adams, 1853–58)
Melanella grandis (A. Adams, 1854 in H. & A. Adams, 1853–58)
Melanella labiosa (Sowerby II, 1834-b)
Melanella lactea (A. Adams, 1854-i)
Melanella musta (Yokoyama, 1928)
Melanella nivea (A. Adams, 1861): species inquerenda
Melanella oblonga (Boettger)
Melanella peronellicola (Kuroda & Habe, 1950-d)
Melanella pinguicula (A. Adams, 1861)
Melanella reclinata (A. Adams, 1861)
Melanella scitula (A. Adams, 1861-b)
Melanella semitorta (A. Adams, 1861-b)
Melanella stenostoma (A. Adams, 1861-b)
Melanella stylata (A. Adams, 1861-b)
Melanella valida (A. Adams, 1861-b)
Melanella venusta (Pease, 1868-b)

Species brought into synonymy

Subgenus Melanella (Balcis) accepted as Melanella Bowdich, 1822
Subgenus Melanella (Melanella) Bowdich, 1822 accepted as Melanella Bowdich, 1822
Melanella acanthyllis (Watson, 1886): synonym of Hemiliostraca acanthyllis (Watson, 1883)
Melanella acerrima (Watson, 1883): synonym of Eulima acerrima Watson, 1883
Melanella aciculata [sic] (Gould in Sowerby, 1866) misspelling of Melanella acicula (Gould, 1849)
Melanella acuta (Sowerby I, 1834): synonym of Hypermastus acutus (G. B. Sowerby I, 1834)
Melanella amblytera (Verrill & Bush, 1900): synonym of Polygireulima amblytera (Verrill & Bush, 1900)
Melanella anomala (Laseron, 1955): synonym of Melanella aciculata (Pease, 1861)
Melanella atypha (Verrill & Bush, 1900): synonym of Eulima atypha Verrill & Bush, 1900
Melanella breviuscula (Dunker, 1875): synonym of Eulima breviuscula Dunker, 1875
Melanella bryani (Pilsbry, 1917): synonym of Melanella aurata Hirase, 1920
Melanella candida (Marrat, 1880): synonym of Melanella martinii (A. Adams in Sowerby, 1854)
Melanella chascanon (Watson, 1883): synonym of Eulima chascanon Watson, 1883
Melanella chaunax (Watson, 1883): synonym of Eulima chaunax Watson, 1883
Melanella clypeastericola (Habe, 1976): synonym of Clypeastericola clypeastericola (Habe, 1976)
Melanella columbiana (Bartsch, 1917): synonym of Vitreolina columbiana (Bartsch, 1917)
Melanella compsa (Verrill & Bush, 1900): synonym of Eulima compsa Verrill & Bush, 1900
Melanella conoidalis (Sowerby, 1865): synonym of Melanella cumingi (A. Adams, 1851)
Melanella corrida Dall, 1927: synonym of Melanella abida Dall, 1927
Melanella crosseana (Brusina, 1886): synonym of Eulima crossei (Brusina, 1886)
Melanella cylindrata (Watson, 1883): synonym of Eulima cylindrata Watson, 1883
Melanella dilecta E. A. Smith, 1899: synonym of Parvioris dilecta (E. A. Smith, 1899)
Melanella engonia (Verrill & Bush, 1900): synonym of Melanella hypsela Verrill & Bush, 1900
Melanella fuscopunctata (E. A. Smith, 1890): synonym of Sticteulima fuscopunctata (E. A. Smith, 1890)
Melanella gomphus (Watson, 1883): synonym of Eulima gomphus Watson, 1883
Melanella helena Thiele, 1930: synonym of Melanella aciculata (Pease, 1861)
Melanella hians (Watson, 1883): synonym of Eulima hians Watson, 1883
Melanella hyalina (Watson, 1883): synonym of Eulima hyalina Watson, 1883
Melanella inconspicua (Watson, 1897): synonym of Eulima inconspicua Watson, 1897
Melanella indeflexa (A. Adams, 1861): synonym of Melanella dufresnei (Bodwich, 1822)
Melanella inflata (Tate & May 1901): synonym of Apicalia inflata (Tate & May 1901)
Melanella inflexa (Blainville, 1825): synonym of Melanella dufresnii Bowdich, 1822
Melanella intermedia (Cantraine, 1835): synonym of Melanella polita (Linnaeus, 1758)
Melanella jamaicensis (C. B. Adams, 1845): synonym of Melanella eburnea (Mühlfeld, 1824)
Melanella jeffreysi (Tryon, 1886): synonym of Melanella platyacme (Sykes, 1903)
Melanella kuronamako (Habe, 1952): synonym of Melanella aciculata (Pease, 1861)
Melanella kyurokusimensis Nomura & Hatai, 1960: synonym of Melanella odontoidea (A. Adams, 1861)
Melanella lampra Melvill, 1918: synonym of Oceanida lampra (Melvill, 1918)
Melanella laseroni Hedley, 1916: synonym of Melanella solitaria (E. A. Smith, 1915)
Melanella latipes (Watson, 1883): synonym of Eulima latipes Watson, 1883
Melanella legrandi (Beddome, 1883): synonym of Eulima legrandi Beddome, 1883
Melanella lineata (Monterosato, 1869): synonym of Fusceulima lineata (Monterosato, 1869)
Melanella luchuana (Pilsbry, 1901): synonym of Melanella cuspidata (A. Adams, 1851)
Melanella macra Bartsch, 1917: synonym of Vitreolina macra (Bartsch, 1917)
Melanella mabutii Nomura & Hatai, 1935: synonym of Parvioris fulvescens (A. Adams, 1866)
Melanella mamilla (May 1915): synonym of Eulima mamilla May, 1915
Melanella mayi (Tate, 1900): synonym of Melanella augur (Angas, 1865)
Melanella modicella (Adams, 1854): synonym of Eulima modicella Adams, 1854
Melanella montageuana (Iredale, 1914): synonym of Eulima montageuana Iredale, T. 1914
Melanella montebelloensis (Iredale, 1914): synonym of Subularia montebelloensis Iredale, T. 1914, synonym of Melanella attenuata (G. B. Sowerby II, 1866)
Melanella monterosatoi (Monterosato, 1890): synonym of Melanella pyramidalis (Sowerby II, 1866)
Melanella murrayae (Cotton & Godfrey, 1932): synonym of Melanella augur (Angas, 1865)
Melanella nitens (Brazier, 1877): synonym of Eulima nitens Brazier, 1876: synonym of Parvioris nitens (Brazier, 1876)
Melanella orthopleura (Tate, 1898): synonym of Melanella augur (Angas, 1865)
Melanella patula auct. non. (Dall & Simpson, 1901): Dall (1927, in part.: 67): synonym of Eulimacrostoma microsculpturata Souza & Pimenta, 2019
Melanella planicincta (Cotton & Godfrey, 1932): synonym of Melanella augur (Angas, 1865)
Melanella platyacme (Sykes, 1903): synonym of Melanella jeffreysii (Tryon, 1886)
Melanella polygyra (Adams, 1851): synonym of Eulima politissima Newton, 1895
Melanella proxima (Sowerby, 1866): synonym of Melanella augur (Angas, 1865)
Melanella psila (Watson, 1883): synonym of Eulima psila Watson, 1883
Melanella schoutanica (May 1915): synonym of Eulima schoutanica May, 1915; synonym of Sabinella schoutanica (May, 1916)
Melanella solida (Sowerby, 1865): synonym of Melanella platyacme (Sykes, 1903)
Melanella solidula (Adams and Reeve, 1850): synonym of Melanella lubrica (Monterosato, 1890)
Melanella tenisoni (Tryon, 1886): synonym of Eulima tenisoni Tryon, 1886
Melanella thaanumi Pilsbry, 1917: synonym of Eulima major G.B. Sowerby, 1834
Melanella tryoni (Tate & May 1900): synonym of Eulima tryoni Tate & May, 1900, synonym of Apicalia tryoni (Tate & May, 1900)
Melanella vafra Pilsbry, 1917: synonym of Parvioris saccata (Boettger, 1893)
Melanella victoriae (Gatliff & Gabriel, 1914): synonym of Eulima victoriae Gatliff & Gabriel, 1914

References

Gofas, S. (2011). Melanella Bowdich, 1822. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=137978 on 20 May 2011
OBIS : Melanella

Bowdich, T.E. 1822. Elements of Conchology, including the fossil genera and animals. Part 1. Univalves. Paris : Smith pp. i–xi, 14–73, index 1–3, 19 pls, 2 tables.
Leach, 1847 [in Gray] Annals and Magazine of Natural History, 20: 271; Placed on the Official List by Opinion 1739, 1993, Bulletin of Zoological Nomenclature, 50(3): 242
Laseron, C. 1955. Revision of the New South Wales eulimoid shells. The Australian Zoologist 12(2): 83–107, pls 1–3, figs 1–78 [85–86, 101] [Euliamaustra is an alternative original spelling (p. 85); spelling Eulimaustra selected by First Reviser action by McMichael & Whitley, 1961, The Marine Zoologist, 1(8): 124]
Warén A. (1984) A generic revision of the family Eulimidae (Gastropoda, Prosobranchia). Journal of Molluscan Studies suppl. 13: 1–96. page(s): 54
Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180–213

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[WoRMS-1] Gofas, S. (2011). Melanella Bowdich, 1822. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=137978 on 20 May 2011

[2] OBIS : Melanella