Zakerana

Minervarya is a genus of frogs in the family Dicroglossidae from south Asia (Sri Lanka, the Indian subcontinent including Pakistan, Nepal, and Bangladesh), and Nepal and Bhutan. They are sometimes known as the Cricket frogs or Rice frogs.[1]

Zakerana
Minervarya keralensis
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Amphibia
Order:	Anura
Family:	Dicroglossidae
Subfamily:	Dicroglossinae
Genus:	Minervarya Dubois, A., A. Ohler, and S. D. Biju, 2001
Type species
Minervarya sahyadris Dubois, Ohler, and Biju, 2001
Diversity
35 species, see text

Taxonomy

The genus Minervarya was proposed to be recognized as a distinct genera from Fejervarya in 2018.[1] This was the culmination of recognizing that Fejervarya (as known before) was paraphyletic and consisted of two different groups; this recognition had already developing some years before and the genus name Zakerana was proposed.[2]

Species

A male of the species kudremukhensis

There are 20 species in this genus:[2]

• Zakerana asmati (Howlader, 2011)
• Zakerana brevipalmata (Peters, 1871)
• Zakerana caperata (Kuramoto, Joshy, Kurabayashi, and Sumida, 2008)
• Zakerana dhaka (Howlader, 2016)
• Zakerana granosa (Kuramoto, Joshy, Kurabayashi, and Sumida, 2008)
• Zakerana greenii (Boulenger, 1905)
• Zakerana keralensis (Dubois, 1981)
• Zakerana kirtisinghei (Manamendra-Arachchi and Gabadage, 1996)
• Zakerana kudremukhensis (Kuramoto, Joshy, Kurabayashi, and Sumida, 2008)
• Zakerana mudduraja (Kuramoto, Joshy, Kurabayashi, and Sumida, 2008)
• Zakerana murthii (Pillai, 1979)
• Zakerana mysorensis (Rao, 1922)
• Zakerana nepalensis (Dubois, 1975)
• Zakerana nilagirica (Jerdon, 1854)
• Zakerana parambikulamana (Rao, 1937)
• Zakerana pierrei (Dubois, 1975)
• Zakerana rufescens (Jerdon, 1854)
• Zakerana sauriceps (Rao, 1937)
• Zakerana sengupti (Purkayastha and Matsui, 2012)
• Zakerana syhadrensis (Annandale, 1919)
• Zakerana teraiensis (Dubois, 1984)

Phylogeny

The following phylogeny of Zakerana is from Pyron & Wiens (2011).[3] 9 species are included.

Zakerana	Zakerana mudduraja         Zakerana pierrei       Zakerana greenii     Zakerana kirtisinghei             Zakerana kudremukhensis     Zakerana rufescens         Zakerana caperata       Zakerana syhadrensis     Zakerana granosa

Howlader, et al. (2016) gives a different phylogeny.[4] 11 species are included.

Zakerana	Zakerana mudduraja       Zakerana rufescens           Zakerana syhandrensis     Zakerana kudremukhensis         Zakerana caperata       Zakerana greenei     Zakerana kirtisinghei               Zakerana granosa     Zakerana pierrei         Zakerana asmati     Zakerana dhaka

Both phylogenetic trees place Zakerana mudduraja as the most basal species.

Suwannapoom, et al. (2016) place Fejervarya chiangmaiensis as a sister taxon to a clade containing Zakerana syhandrensis, Zakerana granosa, and Zakerana pierrei.[5]

References

1. Kurabayashi, Atsushi; Sanchez, Eugenia; Kurabayashi, Atsushi; Biju, S.D.; Islam, Mohammed M.; Hasan, Mahmudul (2018). "Phylogeny and classification of fejervaryan frogs (Anura: Dicroglossidae)". Salamandra. 54 (1): 109–116.
2. Frost, Darrel R. (2014). "Zakerana Howlader, 2011". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 6 February 2014.
3. R. Alexander Pyron; John J. Wiens (2011). "A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians". Molecular Phylogenetics and Evolution. 61 (2): 543–583. doi:10.1016/j.ympev.2011.06.012. PMID 21723399.
4. Howlader MSA, Nair A, Merilä J (2016) A New Species of Frog (Anura: Dicroglossidae) Discovered from the Mega City of Dhaka. PLoS ONE 11(3): e0149597. doi:10.1371/journal.pone.0149597
5. Suwannapoom, C,, Z.-y. Yuan, N. A. Poyarkov, Jr., F. Yan, S. Kamtaeja, R. W. Murphy, and J. Che. 2016. A new species of genus Fejervarya (Anura: Dicroglossidae) from northern Thailand. Zoological Research/Dōngwùxué yánjiū. Kunming 37(6): 1–11.