Holoaden

Holoaden (common name: highland frogs) is a small genus of craugastorid frogs endemic to southeastern Brazil.[1]

Holoaden
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Amphibia
Order: Anura
Family: Craugastoridae
Subfamily: Holoadeninae
Genus: Holoaden
Miranda-Ribeiro, 1920
Diversity
4 species (see text)

Taxonomy

Holoaden is the type genus of subfamily Holoadeninae that was erected in 2008 and placed in family Strabomantidae, erected at the same time. Before that, it was in Brachycephalidae.[2] However, later research has led the whole Strabomantidae, Holoadeninae included, to become included in Craugastoridae.[3][4] This arrangement is not followed by all, and the AmphibiaWeb still keeps Holoadeninae, and consequently Holoaden, in Strabomantidae.[5]

Description

Holoaden grow to 48 mm (1.9 in) snout–vent length at most. Head is narrower than the body. Differentiated tympanic membrane and tympanic annulus are absent. Dorsum is highly glandular. Venter is areolate.[2]

Species

There are four recognized species in this genus:[1][5]

  • Holoaden bradei Lutz, 1958
  • Holoaden luederwaldti Miranda-Ribeiro, 1920
  • Holoaden pholeter Pombal, Siqueira, Dorigo, Vrcibradic, and Rocha, 2008
  • Holoaden suarezi Martins and Zaher, 2013
gollark: There's not necessarily something in every slot.
gollark: no.
gollark: Who doesn't want to live in a glass/concrete cube?
gollark: You can totally build houses without wood.
gollark: It's still *vaguely* useful even without any actual numbers to be aware of some physics things.

References
  1. Frost, Darrel R. (2015). "Holoaden Miranda-Ribeiro, 1920". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 11 June 2015.
  2. Hedges, S. B., Duellman, W. E., and Heinicke, M. P (2008). "New World direct-developing frogs (Anura: Terrarana): Molecular phylogeny, classification, biogeography, and conservation" (PDF). Zootaxa. 1737: 1–182.CS1 maint: multiple names: authors list (link)
  3. Frost, Darrel R. (2015). "Craugastoridae Hedges, Duellman, and Heinicke, 2008". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 11 June 2015.
  4. Padial, J. M.; Grant, T.; Frost, D. R. (2014). "Molecular systematics of terraranas (Anura: Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria". Zootaxa. 3825: 1–132. doi:10.11646/zootaxa.3825.1.1.
  5. "Strabomantidae". AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. 2015. Retrieved 11 June 2015.
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