Gymnogeophagus

Gymnogeophagus
	Gymnogeophagus balzanii
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cichliformes
Family:	Cichlidae
Subfamily:	Cichlinae
Tribe:	Geophagini
Subtribe:	Geophagina
Genus:	Gymnogeophagus; A. Miranda-Ribeiro, 1918
Type species
	Gymnogeophagus cyanopterus; A. Miranda Ribeiro, 1918
	Geographic range of genus Gymnogeophagus

Gymnogeophagus is a genus of cichlid fishes from South America, where they are known from various river basins (notably Rio de la Plata and Lagoa dos Patos–Mirim) in southern Brazil, Paraguay, Uruguay and northern Argentina.[1] They are part of a group popularly known as eartheaters.

Species

There are currently 18 recognized species in this genus:[2]

Gymnogeophagus australis (C. H. Eigenmann, 1907) (Uruguayan eartheater)
Gymnogeophagus balzanii (Perugia, 1891) (Argentine eartheater)
Gymnogeophagus caaguazuensis Staeck, 2006
Gymnogeophagus che Casciotta, S. E. Gómez & Toresanni, 2000
Gymnogeophagus constellatus L. R. Malabarba, M. C. Malabarba & R. E. dos Reis, 2015[3]
Gymnogeophagus gymnogenys (R. F. Hensel, 1870) (Smooth-cheek eartheater)
Gymnogeophagus jaryi Alonso et al., 2019[4]
Gymnogeophagus labiatus (R. F. Hensel, 1870) (Earth eater)
Gymnogeophagus lacustris R. E. dos Reis & L. R. Malabarba, 1988
Gymnogeophagus lipokarenos L. R. Malabarba, M. C. Malabarba & R. E. dos Reis, 2015[3]
Gymnogeophagus mekinos L. R. Malabarba, M. C. Malabarba & R. E. dos Reis, 2015[3]
Gymnogeophagus meridionalis R. E. dos Reis & L. R. Malabarba, 1988
Gymnogeophagus missioneiro L. R. Malabarba, M. C. Malabarba & R. E. dos Reis, 2015[3]
Gymnogeophagus pseudolabiatus L. R. Malabarba, M. C. Malabarba & R. E. dos Reis, 2015[3]
Gymnogeophagus rhabdotus (R. F. Hensel, 1870) (Stripe-fin eartheater)
Gymnogeophagus setequedas R. E. dos Reis, L. R. Malabarba & Pavanelli, 1992
Gymnogeophagus terrapurpura Loureiro, Zarucki, L. R. Malabarba & González-Bergonzoni, 2016[5]
Gymnogeophagus tiraparae González-Bergonzoni, Loureiro & Oviedo, 2009[6]

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References

Froese, Rainer and Pauly, Daniel, eds. (2016). Species of Gymnogeophagus in FishBase. January 2016 version.
Froese, Rainer and Pauly, Daniel, eds. (2018). Species of Gymnogeophagus in FishBase. February 2018 version.
Malabarba, L.R., Malabarba, M.C. & Reis, R.E. (2015). "Descriptions of five new species of the Neotropical cichlid genus Gymnogeophagus Miranda Ribeiro, 1918 (Teleostei: Cichliformes) from the rio Uruguay drainage" (PDF). Neotropical Ichthyology. 13 (4): 637–662. doi:10.1590/1982-0224-20140188.CS1 maint: uses authors parameter (link)
Alonso, F., G.E. Terán, G. Aguilera, O. Říčan, J. Casciotta, W.S. Serra, A. Almirón, M.F. Benítez, I. García and J.M. Mirande (2019). Description of A New Species of the Neotropical Cichlid Genus Gymnogeophagus Miranda Ribeiro, 1918 (Teleostei: Cichliformes) from the Middle Paraná Basin, Misiones, Argentina. PLoS ONE. 14(2): e0210166. doi:10.1371/journal.pone.0210166
Loureiro, M., Zarucki, M., Malabarba, L.R. & González-Bergonzoni, I. (2016): A new species of Gymnogeophagus Miranda Ribeiro from Uruguay (Teleostei: Cichliformes). Neotropical Ichthyology, 14 (1): e150082.
González-Bergonzoni, I., Loureiro, M. & Oviedo, S. (2009). "A new species of Gymnogeophagus from the río Negro and río Tacuarí basins, Uruguay (Teleostei: Perciformes)". Neotropical Ichthyology. 7 (1): 19–24. doi:10.1590/s1679-62252009000100003.CS1 maint: uses authors parameter (link)

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[FB-1] Froese, Rainer and Pauly, Daniel, eds. (2016). Species of Gymnogeophagus in FishBase. January 2016 version.

[fishbase-2] Froese, Rainer and Pauly, Daniel, eds. (2018). Species of Gymnogeophagus in FishBase. February 2018 version.

[Malabarba2015-3] Malabarba, L.R., Malabarba, M.C. & Reis, R.E. (2015). "Descriptions of five new species of the Neotropical cichlid genus Gymnogeophagus Miranda Ribeiro, 1918 (Teleostei: Cichliformes) from the rio Uruguay drainage" (PDF). Neotropical Ichthyology. 13 (4): 637–662. doi:10.1590/1982-0224-20140188.CS1 maint: uses authors parameter (link)

[4] Alonso, F., G.E. Terán, G. Aguilera, O. Říčan, J. Casciotta, W.S. Serra, A. Almirón, M.F. Benítez, I. García and J.M. Mirande (2019). Description of A New Species of the Neotropical Cichlid Genus Gymnogeophagus Miranda Ribeiro, 1918 (Teleostei: Cichliformes) from the Middle Paraná Basin, Misiones, Argentina. PLoS ONE. 14(2): e0210166. doi:10.1371/journal.pone.0210166

[Loureiro2016-5] Loureiro, M., Zarucki, M., Malabarba, L.R. & González-Bergonzoni, I. (2016): A new species of Gymnogeophagus Miranda Ribeiro from Uruguay (Teleostei: Cichliformes). Neotropical Ichthyology, 14 (1): e150082.

[Gonzalez2009-6] González-Bergonzoni, I., Loureiro, M. & Oviedo, S. (2009). "A new species of Gymnogeophagus from the río Negro and río Tacuarí basins, Uruguay (Teleostei: Perciformes)". Neotropical Ichthyology. 7 (1): 19–24. doi:10.1590/s1679-62252009000100003.CS1 maint: uses authors parameter (link)