Nycteola revayana

Nycteola revayana, the oak nycteoline, is a moth of the family Nolidae. The species was first described by Giovanni Antonio Scopoli in 1772. It is found from Europe and east across the Palearctic to Japan and India.

Illustration from John Curtis's British Entomology Volume 1

Nycteola revayana
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Family:	Nolidae
Genus:	Nycteola
Species:	N. revayana
Binomial name
Nycteola revayana (Scopoli, 1772)
Synonyms
Phalaena revayana Scopoli, 1772 Pyralis duplana Fabricius, 1777 Pyralis ilicana Fabricius, 1781 Phalaena lathamiana Swederus, 1787 Phalaena afzeliana Swederus, 1787 Pyralis riuagana Fabricius, 1787 Tortrix punctana Hübner, [1799] Tortrix undulana Hübner, [1799] (preocc.) Tortrix ramosana Hübner, [1799] Phalaena bifasciana Donovan, 1801 Sarrothripus stonius Curtis, 1829 Sarrothripa russiana Duponchel, [1845] Subrita bilineatella Walker, 1866 Sarrothripa fusculana Schmid, 1886 Plotheia diplographa Hampson, 1905

It is a small species and somewhat resembles the Tortricidae. It used to be thought to be related to that family, and was known as the large marbled tortrix.

Technical description and variation

For a key to the terms used, see Glossary of entomology terms.

S. revayana Scop. Forewing ashy grey, dusted with darker, especially in costal half of median area; lines darker, double, but indistinct;the inner outcurved above and below middle, and angled outwards below vein 1; the outer irregularly waved and strongly indented on submedian fold ; some faint dark spots indicate the submarginal line ; hindwing pale greyish fuscous; — in the ab. fusculana Schmid the ground colour is dull dark brownish fuscous, with the black lines visible and a ferruginous spot representing the reniform stigma; from the middle of base an oblique black streak runs to lower end of the inner line ; the submarginal spots are visible, and a row of small black marginal lunules; — ab. dilutana Hbn. has the ground colour dull grey or whitish grey, the median area brownish grey, more purely brown towards costa; the reniform diffuse, rufous; the pale basal area with grey spots between the veins between subbasal and inner lines ; the submarginal spots darker grey and the pale submarginal line beyond them more clearly marked; — ab. degenerana Hbn. is bright greenish white; the median area purplish grey and brown, blacker towards costa, the reniform purple red; the lines filled up with white and the cloudy spots in basal and marginal areas black; a black spot at apex ; fringe whitish ; intermediate forms occur between this and dilutana ; the ab. obscura ab. nov. has the whole forewing olive brown, the markings as in degenerana; — ab. glaucana Lampa has the forewing obscurely glaucescent, with a triangular dark fuscous blotch; — ab. ilicana F. (= punctana Hbn.) is pale fawn grey, sprinkled with whitish scales; the lines much finer; the cell spot, a spot on each side of the median vein before inner line, and the two lowest spots of the submarginal line black; — in ab. ramosana Hbn. (= ramulanus Curt.) the ground colour is brownish grey, with fuscous shading, and a deep black streak from below base of cell to below reniform stigma, shortly branched towards inner margin beyond subbasal and inner lines; the submarghial spots darker; — ab. russiana Dup. has the forewing grey green, with a large dark median blotch. Larva green with the segmental incisions yellowish, the tubercles with long white.[1]The wingspan is 20–25 millimetres (0.79–0.98 in).

Biology

Adults are on wing adults in late autumn, overwintering and appearing again in early spring.

The larvae mainly feed on Quercus species, including Quercus robur, but have also been recorded on Populus and Salix species.[2] They resting curled up between the leaves pupating in a yellow boat-shaped cocoon.

References

1. Warren. W. in Seitz, A. Ed., 1914 Die Großschmetterlinge der Erde, Verlag Alfred Kernen, Stuttgart Band 3: Abt. 1, Die Großschmetterlinge des palaearktischen Faunengebietes, Die palaearktischen eulenartigen Nachtfalter, 1914 This article incorporates text from this source, which is in the public domain.
2. Robinson, Gaden S.; Ackery, Phillip R.; Kitching, Ian J.; Beccaloni, George W.; Hernández, Luis M. (2010). "Search the database - introduction and help". HOSTS - A Database of the World's Lepidopteran Hostplants. Natural History Museum, London.

External links

• Kimber, Ian. "74.009 BF2423 Oak Nycteoline Nycteola revayana (Scopoli, 1772)". UKMoths. Retrieved 4 July 2019.
• Fauna Europaea
• Lepiforum e.V.