Helochares

Helochares
	Helochares lividus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Hydrophilidae
Subfamily:	Hydrophilinae
Tribe:	Acidocerini
Genus:	Helochares; Mulsant, 1844
Synonyms
	Hydrocombus Sharp, 1882

Helochares is a genus of hydrophilid beetles with over 180 species in five subgenera worldwide.[1][2][3][4][5][6][7][8][9][10]

Subgenera

Batochares Hansen, 1991
Helochares Mulsant, 1844
Helocharimorphus Kuwert, 1890
Hydrobaticus MacLeay, 1871
Sindolus Sharp, 1882

Species

Helochares abbreviatus (Fabricius, 1801)
Helochares anchoralis Sharp, 1890
Helochares anthonyae Watts, 1995[9]
Helochares atatus Bruch, 1915
Helochares atlanticus Clarkson & Ferreira, 2014[8]
Helochares atropiceus Regimbart, 1903
Helochares chaquensis Fernández, 1982[11]
Helochares ciniensis Hebauer, Hendrich & Balke, 1999[5]
Helochares clypeatus (Blackburn, 1890)
Helochares crenatus (Regimbart, 1903)
Helochares dalhuntyi Watts, 1995[9]
Helochares densus Sharp, 1890
Helochares discus Hebauer, Hendrich & Balke, 1999[5]
Helochares femoratus (Brullé, 1838)
Helochares foveicollis (Montrouzier, 1860)
Helochares fuliginosus Hendrich & Yang, 1999[12]
Helochares inornatus Orchymont, 1926
Helochares lacustris Hebauer, 1999
Helochares lentus Sharp, 1890
Helochares lividoides Hansen & Hebauer, 1988
Helochares lividus (Forster, 1771)
Helochares livornicus Kuwert, 1890
Helochares longipalpis (Murray, 1859)
Helochares loweryae Watts, 1995[9]
Helochares luridus (MacLeay, 1871)
Helochares maculicollis Mulsant, 1844
Helochares marreensis Watts, 1995[9]
Helochares melanophthalmus (Mulsant, 1844)
Helochares mentinotus Kuwert, 1888
Helochares mesostitialis Fernández, 1981
Helochares minutissimus Kuwert, 1889
Helochares minwissimus Kuwert
Helochares mundus (Sharp, 1882)
Helochares nigripalpis Hebauer & Hendrich, 1999[6]
Helochares nigritulus Kuwert, 1889
Helochares normatus (LeConte, 1861)
Helochares obliquus Mart, Incekara, & Karaca, 2010[3]
Helochares obscurus (O.F. Müller, 1776)
Helochares oculatus Sharp, 1882
Helochares ohkurai Sato, 1976
Helochares orbus Watanabe, 1987
Helochares pallens MacLeay, 1825
Helochares pallipes (Brullé, 1841)
Helochares percyi Watts, 1995[9]
Helochares punctatus Sharp, 1869
Helochares sallaei Sharp, 1882
Helochares sharpi (Kuwert, 1890)
Helochares simulator Knisch, 1922
Helochares talarum Fernández, 1983[13]
Helochares taprobanicus Sharp, 1890
Helochares tatei (Blackburn, 1896)
Helochares tectiformis Fernández, 1982[11]
Helochares tenuistriatus Regimbart, 1908
Helochares tristis (MacLeay, 1871)
Helochares ventricosus Bruch, 1915
Helochares wattsi Hebauer & Hendrich, 1999[6]
Helochares yangae Hebauer, Hendrich & Balke, 1999[5]

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References

"Helochares Mulsant, 1844". ITIS. Retrieved July 24, 2015.
"Helochares Mulsant, 1844". NBN Gateway. Retrieved July 24, 2015.
Mart, Abdullah; Incekara, Umit; Karaca, Hanifi (2010). "A new species and records of Hydrophiliae (Coleoptera) from Turkey" (PDF). Turkish Journal of Zoology. 34: 297–303. doi:10.3906/zoo-0905-29. Archived from the original (PDF) on July 24, 2015.
Hansen, Michael; Hebauer, Franz (1988). "A new species of Helochares from Israel, with a key to the European and some Near East species (Coleoptera:Hydrophilidae)". Insect Systematics and Evolution. 19 (1): 27–30. doi:10.1163/187631289X00023.
Hebauer, Franz; Hendrich, Lars; Balke, Michael (1999). "A contribution to the knowledge of the water beetle fauna (Col. Hydradephaga, Hydrophiloidea and Staphylinoidea) of a tropical freshwater lake: Tasek Cini, Pahang, West Malaysia" (PDF). The Raffles Bulletin of Zoology. 47 (2): 333–348. Archived from the original (PDF) on July 24, 2015.
Hebauer, Franz; Hendrich, Lars (1999). "Two new species of Helochares from Northern Australia (Coleoptera: Hydrophilidae)". Entomological Problems. 30 (1): 47–51.
Short, A.E.Z. "Hydrophilidae: Review of the subtribe Acidocerina of the Southwest Pacific islands" (PDF). Archived from the original (PDF) on July 24, 2015. Cite journal requires |journal= (help)
Clarkson, Bruno; Ferreira, Jr., Nelson (2014). "A new species and records of Helochares (Insecta: Coleoptera: Hydrophilidae) from Southeastern Brazil" (PDF). Zoologia. 31 (4): 400–404. doi:10.1590/s1984-46702014000400012. Archived from the original (PDF) on July 24, 2015.
Watts, C.H.S. (1995). "Revision of the Australasian genera Agraphydrus Rogimbart, Chasmogenus Sharp and Helochares Mulsant (Coleoptera: Hydrophilidae)". Records of the South Australian Museum. 28 (1): 113–130.
Short, Andrew E.Z.; Fikacek, Martin (2011). "World catalogue of the Hydrophiloidea (Coleoptera): additions and corrections II (2006-1010)" (PDF). Acta Entomologica Musei Nationalis Pragae. 51 (1): 83–122. Archived from the original (PDF) on July 26, 2015.
Fernández, Liliana.A. (1982). "Cinco especies nuevas del genero Helochares (Coleoptera: Hydrophilidae)". Physis. Secciones A, B y C (in Spanish). 40 (99): 85–90.
Hendrich, Lars; Yang, C.M. (1999). "A contribution to the knowledge of the water beetle fauna of Palau Tioman, peninsular Malaysia (Coleoptera: Noteridae, Dytiscidae, Hydrophilidae, Hydraenidae, Scirtidae, Limnichidae)" (PDF). The Raffles Bulletin of Zoology. 6: 253–262. Archived from the original (PDF) on July 24, 2015.
Fernández, Liliana.A. (1983). "Helochares (Sindolus) talarum sp. nov., y redscripción de Helochares (Helochares) pallipes (Brullé), con descripción de los estados preimaginales (Coleoptera: Hydrophilidae)". Limnobios (in Spanish). 2: 439–449.

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[1] "Helochares Mulsant, 1844". ITIS. Retrieved July 24, 2015.

[2] "Helochares Mulsant, 1844". NBN Gateway. Retrieved July 24, 2015.

[Martetal-3] Mart, Abdullah; Incekara, Umit; Karaca, Hanifi (2010). "A new species and records of Hydrophiliae (Coleoptera) from Turkey" (PDF). Turkish Journal of Zoology. 34: 297–303. doi:10.3906/zoo-0905-29. Archived from the original (PDF) on July 24, 2015.

[4] Hansen, Michael; Hebauer, Franz (1988). "A new species of Helochares from Israel, with a key to the European and some Near East species (Coleoptera:Hydrophilidae)". Insect Systematics and Evolution. 19 (1): 27–30. doi:10.1163/187631289X00023.

[Hebaueretal-5] Hebauer, Franz; Hendrich, Lars; Balke, Michael (1999). "A contribution to the knowledge of the water beetle fauna (Col. Hydradephaga, Hydrophiloidea and Staphylinoidea) of a tropical freshwater lake: Tasek Cini, Pahang, West Malaysia" (PDF). The Raffles Bulletin of Zoology. 47 (2): 333–348. Archived from the original (PDF) on July 24, 2015.

[Hebauer&Hendrich-6] Hebauer, Franz; Hendrich, Lars (1999). "Two new species of Helochares from Northern Australia (Coleoptera: Hydrophilidae)". Entomological Problems. 30 (1): 47–51.

[7] Short, A.E.Z. "Hydrophilidae: Review of the subtribe Acidocerina of the Southwest Pacific islands" (PDF). Archived from the original (PDF) on July 24, 2015. Cite journal requires |journal= (help)

[Clarkson&Ferreira-8] Clarkson, Bruno; Ferreira, Jr., Nelson (2014). "A new species and records of Helochares (Insecta: Coleoptera: Hydrophilidae) from Southeastern Brazil" (PDF). Zoologia. 31 (4): 400–404. doi:10.1590/s1984-46702014000400012. Archived from the original (PDF) on July 24, 2015.

[Watts-9] Watts, C.H.S. (1995). "Revision of the Australasian genera Agraphydrus Rogimbart, Chasmogenus Sharp and Helochares Mulsant (Coleoptera: Hydrophilidae)". Records of the South Australian Museum. 28 (1): 113–130.

[10] Short, Andrew E.Z.; Fikacek, Martin (2011). "World catalogue of the Hydrophiloidea (Coleoptera): additions and corrections II (2006-1010)" (PDF). Acta Entomologica Musei Nationalis Pragae. 51 (1): 83–122. Archived from the original (PDF) on July 26, 2015.

[Fernandez1982-11] Fernández, Liliana.A. (1982). "Cinco especies nuevas del genero Helochares (Coleoptera: Hydrophilidae)". Physis. Secciones A, B y C (in Spanish). 40 (99): 85–90.

[12] Hendrich, Lars; Yang, C.M. (1999). "A contribution to the knowledge of the water beetle fauna of Palau Tioman, peninsular Malaysia (Coleoptera: Noteridae, Dytiscidae, Hydrophilidae, Hydraenidae, Scirtidae, Limnichidae)" (PDF). The Raffles Bulletin of Zoology. 6: 253–262. Archived from the original (PDF) on July 24, 2015.

[13] Fernández, Liliana.A. (1983). "Helochares (Sindolus) talarum sp. nov., y redscripción de Helochares (Helochares) pallipes (Brullé), con descripción de los estados preimaginales (Coleoptera: Hydrophilidae)". Limnobios (in Spanish). 2: 439–449.